taxonID	type	description	language	source
3A69878C3476FFDD3126F8E3FC51BD94.taxon	description	Figure 1	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3475FFDF3126F880FD0BBD47.taxon	description	Figure 3 a – m	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3475FFDF3126F880FD0BBD47.taxon	materials_examined	Material. Eight incomplete frontoparietals (MHNT. PAL. 2023.0.31.81 – MHNT. PAL. 2023.0.31.87 and MHNT. PAL. 2023.0.31.103); 41 incomplete squamosals (MHNT. PAL. 2023.0.31.63 – MHNT. PAL. 2023.0.31.71, MHNT. PAL. 2023.0.31.88, MHNT. PAL. 2023.0.31.91, MHNT. PAL. 2023.0.31.92, MHNT. PAL. 2023.0.31.94 – MHNT. PAL. 2023.0.31.102, MHNT. PAL. 2023.0.31.104 – MHNT. PAL. 2023.0.31.109, MHNT. PAL. 2023.0.31.111, MHNT. PAL. 2023.0.31.112, and MHNT. PAL. 2023.0.31.117); one putative nasal (MHNT. PAL. 2023.0.31.93); 64 fragmentary maxillae (MHNT. PAL. 2023.0.31.1 – MHNT. PAL. 2023.0.31.62); and one fragment of an exoccipital (MHNT. PAL. 2023.0.31.72).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3472FFDB32C9F9C6FC49B810.taxon	description	Gekkota indet. Figure 6	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3472FFDB32C9F9C6FC49B810.taxon	materials_examined	Material. Eight dentaries (MHNT. PAL. 2023.0.31.118 a – MHNT. PAL. 2023.0.31.118 g, UM-BFI 3060, UM-BFI 3072, and UM-BFI 3090); one trunk vertebra (UM-BFI 3195); and one sacral vertebra (UM-BFI 3193).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3472FFDB32C9F9C6FC49B810.taxon	description	Description Dentaries: All dentaries are broken, with almost no teeth preserved (Fig. 6 a – f). Te largest dentary (MHNT. PAL. 2023.0.31.118 b) has around 23 tooth positions preserved (Fig. 6 b). When preserved the teeth are small, slender and monocuspid (Fig. 6 a). Te sulcus Meckeli is fully closed and fused, and the dentaries are “ tubeshaped ” (Fig. 6 a – c). Trunk vertebra: UM-BFI 3195 is small and lightly build (Fig. 6 g – k). Te centrum is procoelous, with a small cotyle and condyle flattened dorsoventrally (Fig. 6 g, h, j). Te condyle is not inclined in lateral view (Fig. 6 k). Te prezygapophyses are small and strongly oriented dorsomedially (Fig. 6 a, i). Te neural crest is broken but does not seem to extend past the postzygapophyses (Fig. 6 i, k). Te subcentral margins are barely visible, and a pair of subcentral foramina are present on the ventral surface of the centrum (Fig. 6 j). Sacral vertebra: UM-BFI 3193 is small and lightly build (Fig. 6 l – p). Te centrum is amphicoelous, with the notochordal canal opened (Fig. 6 l, m). Te vertebra is very damaged, but the ventral view shows the fusion between two vertebrae (Fig. 6 o). Te pleurapophyses are poorly preserved, but their orientation shows their meet distally (Fig. 6 o, p). Attribution and remarks. Te dentition and closure of the sulcus Meckeli are diagnostic for dentaries of Gekkota in Europe (Augé, 2005; Čerňanský et al., 2023 a; Daza et al., 2014). In the Quercy Phosphorites two genera have been recognized, i. e., Bauersaurus Čerňanský et al., 2023 a, known exclusively from the early Eocene (MP 10 – 11) locality of Cos, and the younger, relatively more widespread, Cadurcogekko (Augé, 2005; Čerňanský et al., 2023 a; Daza et al., 2014; Georgalis et al., 2021 a). Unfortunately, no differences between the two genera are known on their dentaries (the diagnosis of Bauersaurus is based on the maxilla; see Čerňanský et al., 2023 a). Besides, outside Quercy, five other genera of gekkotans have also been established from the Eocene of Europe: Dollogekko Čerňanský et al., 2022, from the early Eocene of Belgium (Čerňanský et al., 2022), Laonogekko Augé, 2003, from the early Eocene of the Paris Basin in France (Augé, 2003), the amber-embedded Yantarogekko Bauer, Böhme & Weitschat, 2005, from the early Eocene of the Baltic Sea region (Bauer et al., 2005), Geiseleptes Villa et al., 2022, from the early to early middle Eocene of Germany (Villa et al., 2022), and Rhodanogekko Hoffstetter, 1946, from the middle Eocene of France (Hoffstetter, 1946). Tus, we tentatively assign all our specimens to Gekkota indet. UM-BFI 3193 is assigned to Gekkota on the basis of an amphicoelous condition (known, among squamates, only in Gekkota in the Cenozoic of Europe; Augé, 2005; Čerňanský et al., 2018; Daza et al., 2014; Delfino et al., 2011; Estes, 1983). Amphicoelous vertebrae have in the past been assigned to Cadurcogekko (Augé, 2005; Rage, 1988) but this assignment is not fully confirmed, because no articulated specimens of this genus are known. UM-BFI 3195 is more puzzling. It does matches the general morphology of known Cenozoic gekkotans from Europe (Daza et al., 2014) but differs in being procoelous. Procoelous vertebrae are known in certain extant gekkotan taxa (Hoffstetter & Gasc, 1969) but have never been documented in the Cenozoic fossil record in Europe (Daza et al., 2014). Te presence of both amphicoelous and procoelous vertebral material assigned to Gekkota indicates that at least two species of gekkotans were present in La Bouffie. It also marks the first occurrence of a procoelous gekkotan in the Cenozoic fossil record of Europe.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347DFFD032C9F9E0FB96B9B0.taxon	description	Figure 10	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347DFFD032C9F9E0FB96B9B0.taxon	materials_examined	Material. One incomplete right dentary (UM-BFI 3079).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347DFFD032C9F9E0FB96B9B0.taxon	description	Description. UM-BFI 3079 is a fragment of a large dentary (Fig. 10 a). Eight teeth emplacements (including six teeth) are preserved (Fig. 10 a). Te teeth are high-crowned, with one-third of their height projecting above the parapet (Fig. 10 b). Te teeth are straight and slender, expect for the posteriormost one, the latter being wider (Fig. 10 a). One tooth is complete and shows a tricuspid condition, with a large central cusp and two small lateral ones (Fig. 10 a, b). Te subdental shelf is moderately developed, with a rounded lingual margin and lacks a sulcus dentalis (Fig. 10 a). Te ventral wall bears an inframeckelian lip parallel to the subdental shelf (Fig. 10 a). As such, the sulcus Meckeli narrowly opens lingually (Fig. 10 a). On the labial surface, two mental foramina are preserved (Fig. 10 b). Attribution and remarks. UM-BFI 3079 is reminiscent of Geiseltaliellus lamandini based on the following combination of characters (see Augé, 2005): (1) absence of a sulcus dentalis; (2) a sulcus Meckeli narrowing anteriorly; (3) slender and straight tricuspid teeth; (4) presence of inframeckelian lip parallel to the subdental shelf; and (5) enlarged posterior teeth. It is worth noting though that the first four characters of this combination are also present in other pleurodontans, including the early Eocene North American genus Suzanniwana Smith, 2009 a, which has anyway been suggested to bear much resemblance to Geiseltaliellus, though being more primitive (Smith, 2009 a). In any case, due to the incompleteness of the only specimen from La Bouffie, we treat this species level identification as tentative. Furthermore, it should be noted that the referral of G. lamandini (originally established as a new species of Lacerta Linnaeus, 1758, by Filhol [1877]; subsequently recombined into the genus Pseudolacerta by Hoffstetter [1942]; and subsequently recombined into the genus Geiseltaliellus by Augé [2005]) to Geiseltaliellus is uncertain, as its lacks the known autapomorphies of the genus (Smith, 2009 b). Pleurodonta incertae sedis	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347DFFD032C9F9E0FB96B9B0.taxon	materials_examined	Material. Two incomplete left dentaries (MHNT. PAL. 2023.0.31.119 a and MHNT. PAL. 2023.0.31.119 b).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347DFFD032C9F9E0FB96B9B0.taxon	description	Description. Te dentary preserves 13 tooth positions, with four teeth (Fig. 11 a, b). Te dentition is pleurodont and heterodont (Fig. 11 a, b). Teeth are straight and slen- der (Fig. 11 a, b). Te anteriormost tooth has a pointed apex, while the other three are tricuspid with their lateral cuspids small (Fig. 11 a). In labial view, a third of the teeth are visible above the parapet (Fig. 11 c, d). Te labial surface is pierced by two foramina (Fig. 11 c, d). In lingual view, the sulcus Meckeli is fully closed by the expansion of the ventral wall (Fig. 11 a, b). A thin groove / suture is visible medially between the subdental shelf and the ventral wall of the dentary (Fig. 11 a, b). Attribution and remarks. Tese small dentaries are referred to Cadurciguana hoffstetteri (the type and only known species of the genus Cadurciguana, originally typified from the MP 19 locality of Escamps C, Quercy; Augé, 1987, 2005) based on the following characters (see Augé, 2005): (1) heterodont dentition with anterior teeth monocuspid and posterior teeth tricuspid with small lateral cusps; (2) a sulcus Meckeli fully closed and fused ventromedially by the expansion of the ventral wall; and (3) presence of a small groove extending anteriorly from the posterior end of the closed and fused sulcus Meckeli. Nevertheless, it has to be emphasized that the particularly the former two characters are not really diagnostic, as they represent widespread features, shared also with numerous (perhaps even hundreds of) extant pleurodontan taxa. However, still, as far as it regards the European fossil record, this combination of characters appears to be unique to Cadurciguana.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	description	Figure 13 ZooBank registration. urn: lsid: zoobank. org: act: 8 C 92 DEA 5 - EE 9 E- 427 F- 9929 - D 21 F 88366 D 87.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	etymology	Etymology. Te species epithet originates from the Greek word “ peritechne ” (“ περίτεΧνη ”), meaning “ elaborate ”, “ crafted with beautiful art ”, referring to the beautiful, elaborate shape of the teeth of the new taxon.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	materials_examined	Holotype. One complete left dentary (UM-BFI 3057).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	diagnosis	Differential Diagnosis. Phosphoriguana peritechne gen. et sp. nov. differs from Cadurciguana in: (1) having an opened sulcus Meckeli (closed and fused in C. hoffstetteri) (see Augé, 2005); (2) having more developed accessory cusps; and (3) having a more arched ventrally margin of the dentary. Phosphoriguana peritechne gen. et sp. nov. differs from Geiseltaliellus in lacking the presence of an inframeckelian lip narrowing or closing the sulcus Meckeli (see Augé, 2005; Smith, 2009 b). Phosphoriguana peritechne gen. et sp. nov. differs from Geiseltaliellus lamandini in having an arched ventral margin of the dentary (see Augé, 2005). Phosphoriguana peritechne gen. et sp. nov. differs from Bifurcodentodon Čerňanský et al., 2023 b in lacking the dentition made of two main cuspids, with several accessory cusps (see Čerňanský et al., 2023 b). Phosphoriguana peritechne gen. et sp. nov. differs from Pseudolacerta De Stefano, 1903, in lacking the strongly heterodont dentition and lacking the enlarged anterior caniniform teeth present in the latter genus (see Augé, 2005).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	materials_examined	Type locality and age. La Bouffie, fissure fill located in the village of Vaylats (Lot, SW France); late Eocene (MP 17 A); the species is so far known exclusively from the type locality.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C347AFFD332C9FA66FB29BF50.taxon	description	Description of the holotype. UM-BFI 3057 is a complete dentary missing only a few teeth (Fig. 13). Te dentary is elongated anteroposteriorly and large (Fig. 13 a, b). 15 teeth are preserved, and at least six additional tooth positions can be identified, but the presence of sediment obscures most of the region, preventing a definitive total count (Fig. 13 c). A broken tooth shows the implantation to be pleurodont; the dentition is heterodont (Fig. 13 a, c). Te first three anterior teeth are monocuspid, with a posteriorly oriented triangular apex (Fig. 13 a, b). Posteriorly, the teeth become tricuspid, with a large central cusp and two smaller mesial and distal accessory cusps of equal size well separated from the central one (Fig. 13 d). Te posterior teeth are also larger and longer than the anterior ones (Fig. 13 a). No sulcus dentalis seems to be present (Fig. 13 c). Te subdental shelf is concave ventrally and expands only slightly anteriorly (Fig. 13 b). On the ventral surface of the subdental shelf, a well-developed facet for the splenial extends anteriorly up to the sixth tooth position (starting from the anteriormost; Fig. 13 a). Te ventral side of the sulcus Meckeli also bears a facet for the splenial, extending from the second posteriormost tooth up to the third tooth position anteriorly. Te sulcus Meckeli is widely opened ventrolingually posteriorly, and progressively narrows anteriorly (Fig. 13 a). At the level of the anteriormost five teeth, the subdental shelf covers lingually the sulcus Meckeli up to the symphysis (Fig. 13 a). On the labial surface, a row of eight labial foramina opens mid-length of the surface (Fig. 13 b). Both angular and coronoid processes are symmetrically developed (Fig. 13 b). Moreover, there is a distinct groove on the posterolabial surface of the dentary for an anterolateral process of the coronoid. Tis groove seems to be more complex than usual (for pleurodontan standards), with an anteroposterior ridge running from front to back. It does not broaden anteriorly but might be dorsoventrally short (Fig. 13 a). Attribution and remarks. Two synapomorphies have been proposed on the dentary for Pleurodonta (Gauthier et al., 2012): (1) the restriction of the sulcus Meckeli; and (2) the anteromedial process of the coronoid wrapping under the level of the teeth row. UM-BFI 3057 only possesses the second character and thus is tentatively referred to Pleurodonta. Among known European pleurodontans, Phosphoriguana gen. nov. can be differentiated from all known genera (see Differential Diagnosis above). Outside Europe, a large number of pleurodontans is known from the Eocene of North America, pertaining to a number of extinct genera (e. g., Estes, 1983; Gilmore, 1928; Scarpetta, 2020; Smith & Gauthier, 2013; Smith, 2006, 2009 a, 2011 a, 2011 b). Te shape and dentition of the holotype dentary UM-BFI 3057 does not also match with the corresponding morphology of the North American taxa. Tus, we establish a new genus and species in order to accommodate the dentary UM-BFI 3057. Even though its distinctive anatomical features, the new taxon Phosphoriguana peritechne gen. et sp. nov. cannot be confidently referred to Pleurodonta based solely on the above-described features. As a matter of fact, pleurodont tooth implantation, tricuspid dentition, and lack of a sulcus dentalis are indeed typical features of pleurodontans, however, these can be also observed also in some members of Teiidae (e. g., genus Kentropyx Spix, 1825) and Lacertidae (e. g., Arnold et al., 2007). Furthermore, our current knowledge about potential synapomorphies in the jaws of Pleurodonta is still limited, and as a matter of fact, only a few have now been pointed out. Two were proposed by Smith (2009 b): (1) in the maxilla, a separate foramen for the subnarial artery on the premaxillary process; and possibly (2) in the dentary, a wrapping of the anteromedial process of the coronoid underneath the tooth row. (that latter was suggested by Smith, 2009 b but was recovered as a synapomorphy by Gauthier et al., 2012) Additionally, Gauthier et al. (2012) identified a third synapomorphy: (3) a restriction (but not closure, much less fusion) of the Meckelian groove as a synapomorphy of Pleurodonta and part of its stem. For further discussion of these characters, see Smith (2020 a, 2020 b, 2020 c) definitions of Pan-Iguania, Pan-Acrodonta and Pan-Iguanidae. Te holotype dentary (UM-BFI 3057) of Phosphoriguana peritechne gen. et sp. nov. from La Bouffie seems to possess character (2) but not (3), so the evidence for its taxonomic attribution to Pleurodonta is somehow ambiguous. To the contrary, character (3) in UM-BFI 3057 really deviates from the typical pleurodontan pattern: practically, pleurodontans - with the exception of Hoplocercidae - conform to character (3) and do not normally show such a widely open Meckelian groove, as observed in the new taxon from La Bouffie. Outside Pleurodonta, an alternative, potential taxonomic allocation of Phosphoriguana peritechne gen. et sp. would be that this holotype dentary (UM-BFI 3057) pertains to the same taxon as the frontal from La Bouffie that was originally described as a tupinambine teiid by Augé and Brizuela (2020: fig. 2; specimen MNHN. F. BFI 1877). All these being said, Phosphoriguana peritechne gen. et sp. is only tentatively referred to Pleurodonta and we anticipate that future (ideally articulated) specimens may shed some more light on its precise taxonomic affinities within other lizards.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3467FFCC3126F899FC13BD9F.taxon	description	Figures 14, 15	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3462FFCB3126FE46FE82BBF0.taxon	description	Figure 18	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3462FFCB3126FE46FE82BBF0.taxon	materials_examined	Material. Seven trunk vertebrae (UM-BFI 3077, UM-BFI 3088, UM-BFI 3132, UM-BFI 3136, UM-BFI 3192, UM-BFI 3194, and UM-BFI 3196); and one caudal vertebra (UM-BFI 3172).	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
3A69878C3462FFCB3126FE46FE82BBF0.taxon	description	Description. UM-BFI 3077 is a trunk vertebra, missing the right prezygapophysis and the anterior portion of the neural arch (Fig. 18 a – e). Te centrum is procoelous, with a dorsoventrally compressed cotyle and condyle (i. e., wider than high; Fig. 18 a, b). Its ventral surface is flat, with no haemal keel visible (Fig. 18 d). Te centrum is elongated anteroposteriorly (Fig. 18 c). Te neural canal is arched and taller than the cotyle (Fig. 18 a). Te neural arch has a marked interzygapophyseal constriction (Fig. 18 c). Te prezygapophysis bears an ovoid articular facet extended anteriorly (Fig. 18 c). Te facet is oriented dorsomedially (Fig. 18 a). Te subcentral ridges of the centrum are concave (Fig. 18 d). A thin dorsal ridge extends on the neural arch on its entire length (Fig. 18 c). Posteriorly, it thickens into a short neural spine (Fig. 18 c, e). UM-BFI 3132 is identical to UM-BFI 3077 but is poorly preserved (Fig. 18 f – j). UM-BFI 3192 and UM-BFI 3196 are also similar to UM-BFI 3077. UM-BFI 3088 (Fig. 18 k – o) and UM-BFI 3136 (Fig. 18 p – t) are similar to UM-BFI 3077 and UM-BFI 3132, with the exception of the following characters: (1) articular facet of the prezygapophyses less extended anteriorly; (2) shorter and wider centrum; and (3) shorter neural spine (likely linked to a poorer preservation). UM-BFI 3194 is also similar to UM-BFI 3088 and UM-BFI 3136. UM-BFI 3172 (Fig. 18 u – w) is a caudal vertebra missing most of its neural arch. Te vertebra is similar to the trunk vertebrae on the following characters: (1) cotyle and condyle wider than high; (2) relatively smooth ventral surface of the centrum; and (3) neural arch not vaulted. Te para- and diapophyses are fused into a large pleurapophysis, extending laterally (Fig. 18 u – w). Attribution and remarks. All these specimens are referred to Anguinae based on the following characters: (1) cotyle and condyle depressed; (2) flattened ventral surface of the centrum; and (3) neural arch not very vaulted posteriorly (Augé, 2005; Meszoely, 1970; Rage & Augé, 2015). Note that the two former characters are also present in the North American related group Anniellinae (Bell et al., 1995; Meszoely, 1970; Smith, 2011 a). In any case, a taxonomic identification below the family level in pre-Miocene anguines is difficult when based on non-cranial elements, as several genera were present during the Paleogene of Europe and for certain of these vertebrae are so far unknown (Augé, 2005; Čerňanský et al., 2016 a; Georgalis et al., 2021 a, 2021 b; Klembara & Green, 2010; Vasilyan et al., 2022) and variation within the vertebral column can be high (Georgalis & Scheyer, 2021). It should be noted that UM-BFI 3077, UM-BFI 3132, UM-BFI 3192, and UM-BFI 3196 represent an Ophisaurus - like morphotype vertebra, with: (1) subcentral ridges concave; and (2) height of the neural canal taller than the cotyle (see Čerňanský et al., 2019). As UM-BFI 3088, UM-BFI 3136, and UM-BFI 3194 differ from UM-BFI 3077 on these features, they could potentially represent a second anguine taxon from La Bouffie, but it is impossible to assess this based on such limited amount of isolated vertebrae, while admittedly, we cannot exclude that these differences can be merely attributed to some (unknown degree of) intracolumnar variation. Te genus Ophisaurus Daudin, 1803, has been mentioned (on non-illustrated materials) from La Bouffie (Crochet et al., 1981; later mentioned only as Anguinae indet. in Augé, 2005). Tese being said, we consider these genus-level identifications of Crochet et al. (1981) to be unfounded, pending the discovery of cranial elements from La Bouffie.	en	Lemierre, Alfred, Georgalis, Georgios L. (2025): Diversity in a greenhouse world: herpetofauna from the late Eocene (MP 17 A) of La Bouffie, Quercy Phosphorites (Lot, SW France). Swiss Journal of Palaeontology 144 (1): 1-39, DOI: 10.1186/s13358-025-00370-9, URL: https://doi.org/10.1186/s13358-025-00370-9
