taxonID	type	description	language	source
7F18A5B148145124A6B0EA4154F90ABB.taxon	description	Figs 5 C, D, 9, 10, 11	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
7F18A5B148145124A6B0EA4154F90ABB.taxon	etymology	Etymology. The species epithet is an adjective formed from the type locality, Battambang Province, Cambodia.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
7F18A5B148145124A6B0EA4154F90ABB.taxon	diagnosis	Diagnosis. This new species is distinguished by its notably small body size, being the smallest known in the genus Orthomorpha, measuring 12.4 – 14.6 mm (♂) or 17.2 – 17.8 mm (♀) in length, and 1.6 – 1.8 mm (♂) or 2.1 – 2.3 mm (♀) in width at midbody. The gonopod solenophore is complex, best described as trifid, bearing a small median denticle between the long terminal lobe and a smaller subterminal lobule. Additionally, tarsal brushes are present until ♂ legs of ring 9.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
7F18A5B148145124A6B0EA4154F90ABB.taxon	description	Description. Length 12.4 – 14.6 mm (♂), 17.2 – 17.8 mm (♀), width of midbody pro- and metazona 1.0 – 1.1 and 1.6 – 1.8 mm (♂) or 1.5 – 1.7 and 2.1 – 2.3 mm (♀), respectively. Coloration of live animals dark castaneous brown (Fig. 5 C, D), with contrasting pale red or bright orange paraterga and epiproct; antennae dark brownish, venter and legs dark brown to brown; coloration of alcohol material after 10 months of preservation faded to uniformly reddish brown (Fig. 9 A – G) with contrasting light yellow paraterga and epiproct; antennae, venter and legs brown to yellowish brown. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long (Fig. 9 B), reaching to ring 3 (♂, ♀) when stretched dorsally. In width, collum <head <ring 3 <4 <2 <5 <6 <7 – 16 (Fig. 9 A, C, F), thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 4 + 4 anterior, 2 + 2 intermediate, and 3 + 3 posterior; a small incision laterally in posterior 1 / 3; caudal corner of paraterga pointed and dentiform; paraterga declined ventrad, not drawn past rear margin. Tegument generally smooth and shining (Fig. 9 A – F), metaterga only at places faintly rugulose, slightly more so near rear margin; prozona very finely shagreened, metazona below paraterga faintly rugulose (Fig. 9 B, D, E). Postcollum metaterga with two transverse rows of setae: 2 + 2, mostly abraded setae in anterior (pre-sulcus) row, 3 + 3 in posterior (post-sulcus) one, the latter setae borne on minute tubercles, gradually reduced in size thereafter. Tergal setae simple, rather long, about 1 / 3 metatergal length, mostly abraded. Axial line traceable both on pro- and metazona. Paraterga strongly developed (Fig. 9 A – F), lying level to or slightly above dorsum, only on rings 1 – 4, 18 and 19 either lying clearly below dorsum (♂) or all lying slightly below dorsum, set at about half of midbody height, subhorizontal (♀), in lateral view moderately enlarged in pore-bearing rings, thinner in poreless ones (Fig. 9 B, D, E); shoulders broadly rounded, narrowly bordered, fused to callus; caudal corner narrowly rounded to pointed, slightly drawn past rear tergal margin (Fig. 9 B, D, E). Calluses delimited by a sulcus only dorsally. Paraterga 2 broad, anterior edge convex, lateral edge with two acute denticles, one in anterior 1 / 3, the other at midway (Fig. 9 A). Each following poreless ring with two incisions, each pore-bearing one with one, often evident incision in front of ozopore. Posterior edge of paraterga oblique, especially clearly so in rings 17 – 19. Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 9 D, E). Transverse sulcus distinct (Fig. 9 A, C, F), slightly incomplete on ring 19, complete and clearly visible on metaterga 5 – 18, rather deep, reaching the bases of paraterga, line-shaped, beaded at bottom. Stricture between pro- and metazona rather wide, shallow, beaded at bottom down to base of paraterga (Fig. 9 A, C, F). Pleurosternal carinae well-developed, as complete, arcuate ridges with distinct caudal denticle on rings 2 – 8 (♂) (Fig. 9 B, D) or rings 2 – 4 (♀), as a sharp caudal tooth, the latter gradually reduced in size down to a small tooth until ring 17 (♂), or broken into an anterior bulge and a caudal tooth on rings 5 – 8, retained only as a small, caudal, mostly rounded tooth on rings 9 – 16 (♀), entirely absent thereafter. Epiproct (Fig. 9 E – G) conical, flattened dorsoventrally, with two evident apical papillae; tip subtruncate; pre-apical papillae small, but visible, lying rather close to tip (Fig. 9 F, G). Hypoproct nearly semi-circular, 1 + 1 setiferous knobs at caudal edge well-separated and evident (Fig. 9 G). Sterna sparsely setose, without modifications; cross-impressions shallow; with a paramedian pair of evident, anteroventrally directed, rounded prongs between ♂ coxae 4 (Fig. 9 H, I). A conspicuous ridge in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3 – 1.5 (♂) or 1.1 – 1.3 times (♀) as long as body height (Fig. 9 B, F), prefemora without modifications, ♂ tarsal brushes present until leg 4. Gonopods stout and suberect (Figs 10, 11). Coxite (cx) rather short, slightly curved caudally, rather densely setose distodorsally (Figs 10 A, B, 11 A, B). Prefemoral part (pfe) densely setose, less than half the length of acropodite (femoral + postfemoral parts) (Fig. 10 A, B). Femoral part (fe) rather stout, slightly curved and faintly enlarged distad, with a postfemoral part demarcated by an oblique lateral sulcus (Figs 10 A, B, 11 A, B). Tip of solenophore (sph) faintly bifid (Figs 10 B, C, 11 A, C, D), with a subtruncate and long terminal lobe (Figs 10, 11 A – H) and a small subterminal lobule (Figs 10 B, C, 11 A, C, D); solenomere (sl) long and flagelliform (Figs 10, 11 A, B, D – H).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
219CCE656B4F51F481507E964467422A.taxon	description	Figs 5 A, B, 6, 7, 8	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
219CCE656B4F51F481507E964467422A.taxon	etymology	Etymology. The species name ‘ efefai’ is a phonetic spelling representation of the Fauna and Flora Cambodia (formerly known by the acronym FFI). This new species is named in honour of their passionate commitment to protecting Earth’s biodiversity and acknowledges FFI Cambodia for supporting our team to explore the rich biodiversity of the country. When pronounced, ‘ efefai’ is with the ‘ e’ and ‘ a’ being silent.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
219CCE656B4F51F481507E964467422A.taxon	diagnosis	Diagnosis. This new species seems to be particularly similar to Orthomorpha parasericata Likhitrakarn, Golovatch & Panha, 2010, a species known only from Surat Thani and Phang Nga provinces in southern Thailand, in having likewise broad, contrastingly lighter-colored paraterga, coupled with a stout gonopod telopodite and a bifid solenophore. However, it clearly differs by its smaller body size (28.4 – 33 mm long and 4.2 – 4.9 mm wide in males, 32.5 mm long and 4.6 mm wide in females, vs 32 – 37 mm long and 4.8 – 5.0 mm wide in males, 34 – 37 mm long and 4.8 – 5.3 mm wide in females of O. parasericata), as well as by the shape of the solenophore apex. Additionally, tarsal brushes are present until ♂ legs of ring 9 in the new species, whereas in O. parasericata, they are present only until ♂ legs of ring 5. Orthomorpha efefai sp. nov. can also be distinguished from the sympatric O. tergoaurantia sp. nov., with which it shares a superficially similar gonopod structure. The key differences lie in the somatic characters. The metaterga in O. efefai sp. nov. are distinctly rugulose-tuberculate, bearing two rows of setiferous cones, especially on the anterior body rings (Fig. 6 A, B). In contrast, the metaterga of O. tergoaurantia are mostly smooth and leathery, with setiferous knobs being far less pronounced (Fig. 2 A – F). The male tarsal brushes extend until the legs of ring 9 in O. efefai sp. nov., but are far more extensive in O. tergoaurantia sp. nov., persisting until the legs of ring 17. Additionally, the caudal tooth of the pleurosternal carinae is traceable until ring 16 in O. efefai sp. nov., but only until rings 7 or 8 in O. tergoaurantia sp. nov. (Fig. 2 B, D). Although broadly similar, the solenophore tip in O. efefai sp. nov. is distinctly bifid (Figs 7, 8), while in O. tergoaurantia sp. nov. it is trifid, with a small but clear middle prong (Figs 3, 4 A, C – F).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
219CCE656B4F51F481507E964467422A.taxon	description	Description. Length 28.4 – 31.8 mm (♂), 32.5 mm (♀), width of midbody pro- and metazona 2.6 – 2.7 and 4.2 – 4.3 mm (♂) or 3.3 and 4.6 mm (♀), respectively. Coloration of live animals blackish (Fig. 5 A, B), with contrasting reddish pink or bright pink paraterga and epiproct, posterior halves of metaterga on rings 16 – 1 and antennae blackish, legs brown; coloration of alcohol material after 10 months of preservation faded to uniformly dark brown (Fig. 6 A – G) with contrasting bright pink or pale pinkish paraterga and epiproct, legs brown to light grey-brown. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long (Fig. 5 B), extending past ring 3 (♂) or reaching to ring 3 (♀) when stretched dorsally. In width, head <collum <ring 3 <4 <2 <5 <6 <7 – 15 (Fig. 6 A), thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 3 + 3 anterior, 2 + 2 intermediate, and 3 + 3 posterior; a very faint incision laterally in posterior 1 / 3; caudal corner of paraterga pointed, dentiform, paraterga declined ventrad, not drawn past rear margin. Tegument of metaterga shining, rugulose-tuberculate, especially so on several front metaterga; prozona very finely shagreened, metazona below paraterga faintly rugulose (Fig. 6 A – F). Metaterga 2 – 5 with two rows of 2 + 2 anterior and 3 + 3 setiferous cones, usually slightly smaller cones in anterior (pre-sulcus) row and more evident ones laterally in posterior row (Fig. 6 A, B); thereafter same pattern, but traceable only as insertion points in anterior row and as minute knobs gradually increasingly obliterate to become nearly absent from ring 10 onward. Tergal setae short, simple, slender, often abraded, about 1 / 4 metatergal length. Axial line traceable, especially clear on collum and following few metaterga. Paraterga very strongly developed (Fig. 6 A – F), broad, all lying below dorsum (at about 1 / 4 body height), mostly subhorizontal, slightly upturned on rings 2 – 5, in lateral view moderately enlarged on pore-bearing rings, thinner on poreless ones (Fig. 6 B, D, E); anterior margin well-developed, mostly nearly straight and narrowly bordered, fused to callus; caudal corner of most of paraterga very narrowly rounded, increasingly drawn past tergal margin, slightly curved mesad on rings 15 – 19 (Fig. 6 B, D, E). Calluses on paraterga delimited by a sulcus only dorsally. Paraterga 2 broad, lateral edge with two small, but evident incisions, one in anterior 1 / 3, the other in posterior 1 / 3. Paraterga 3 and 4 each with an evident incision in anterior 1 / 3 at lateral margin (Fig. 6 A). Lateral margins of following paraterga with two small incisions, one at midway, the other in posterior 1 / 3, the other at midway, caudal incision being smaller on pore-bearing rings (Fig. 6 A, C, F). Posterior margin of paraterga concave, especially clearly so in rings 15 – 19. Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 6 B, D, E). Transverse sulcus distinct (Fig. 6 A – F), slightly incomplete on ring 2, complete and clearly visible on metaterga 3 – 18, shallow, reaching the bases of paraterga, arcuate, faintly beaded at bottom. Stricture between pro- and metazona rather wide, deep, beaded at bottom down to base of paraterga (Fig. 6 A, C, F). Pleurosternal carinae complete crests on rings 2 – 4, following rings 5 – 7 (8) each broken into an anterior bulge and a sharp caudal tooth, the latter gradually reduced in size to a small tooth until ring 16, thereafter missing (♂, ♀) (Fig. 6 B, C, E). Epiproct (Fig. 6 E – G) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed ventrocaudally (Fig. 6 E, G); pre-apical papillae small, but evident, lying close to tip. Hypoproct subtrapeziform (Fig. 6 G), 1 + 1 small setiferous knobs at caudal edge well-separated and evident. Sterna sparsely setose, without modifications; cross-impressions rather deep; a paramedian pair of evident, fully separated, setose cones between ♂ coxae 4 (Fig. 6 H, I). A conspicuous ridge in front of gonopod aperture. Legs moderately long and slender, midbody ones ca 1.0 – 1.2 (♂) (Fig. 5 B) or 0.9 – 1.1 times (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes present until leg 9. Gonopods simple and suberect (Figs 7, 8). Coxite (cx) long and slender, slightly curved caudally, rather densely setose distodorsally (Figs 7 B, C, 8 A, B). Prefemoral part (pfe) densely setose, as usual, less than half the length of acropodite (femoral + postfemoral parts) (Fig. 7 B, C). Femoral part (fe) slender, slightly curved and slightly enlarged distad, with a postfemoral part demarcated by an oblique lateral sulcus (Figs 7 A, B, 8 B, E). Solenophore (sph) with an evidently bifid tip, with a subterminal lobule (Figs 7, 8 A – E) and a spiniform apical lobule (Figs 7, 8 A – F); solenomere (sl) long and flagelliform (Figs 7 A, B, 8 A, C, E).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
DFFD7DF8871059E8973788FFE7523BEA.taxon	description	Figs 1, 2, 3, 4	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
DFFD7DF8871059E8973788FFE7523BEA.taxon	etymology	Etymology. The species name ‘ tergoaurantia’ is derived from the Latin words ‘ tergum’, meaning ‘ back’, and ‘ aurantium’, meaning ‘ orange’. This name emphasizes the distinctive dark or bright orange coloration of the paraterga, which is a prominent characteristic distinguishing it from other Cambodian congeners.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
DFFD7DF8871059E8973788FFE7523BEA.taxon	diagnosis	Diagnosis. This new species seems to be particularly similar to Orthomorpha cambodjana (Attems, 1953) in gonopod conformation, sharing a very slender and suberect gonopodal telopodite. However, it clearly differs by its larger body size (35.4 – 39.4 mm long and 4.1 – 4.9 mm wide in ♂, 35.4 – 39.4 mm long and 4.2 – 4.3 mm wide in ♀, vs 17 – 30 mm long and 2.0 – 3.1 mm wide in ♂, 19 – 29 mm long and 2.0 – 3.4 mm wide in ♀ of O. cambodjana) and by the shape of the solenophore apex. The caudal denticle on the pleurosternal carinae is traceable until body rings 7 or 8 in the new species (vs rings 16 or 17 in O. cambodjana). Additionally, tarsal brushes are present until ♂ legs of ring 17 in the new species, vs only until ♂ legs 7 of O. cambodjana.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
DFFD7DF8871059E8973788FFE7523BEA.taxon	description	Description. Length 31.2 – 37.3 mm (♂), 35.4 – 39.4 mm (♀), width of midbody pro- and metazona 2.5 – 3.2 and 4.1 – 4.9 mm (♂) or 3.2 – 3.9 and 4.7 – 5.7 mm (♀), respectively. Coloration of live animals dark brown to blackish (Fig. 1), with contrasting dark orange to bright orange or yellowish paraterga and epiproct; antennae black; venter and legs brown to blackish; coloration of alcohol material after six years of preservation faded to dark castaneous brown, paraterga, venter, epiproct, and several basal podomeres more flavous, pale pinkish, brownish or pale yellow (Fig. 2 A – G). Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long (Fig. 2 B), reaching or extending past ring 3 (♂) or reaching to ring 3 (♀) when stretched dorsally. In width, head <collum <ring 3 <4 <2 <5 <6 <7 – 16 (Fig. 2 A), thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 4 + 4 anterior, 2 + 2 intermediate, and 4 + 4 posterior; a small incision laterally in posterior 1 / 3; caudal corner of paraterga pointed, dentiform, paraterga declined ventrad, not drawn past rear margin (Fig. 2 A, B). Tegument smooth and shining, prozona finely shagreened, metaterga smooth and leathery, posterior halves often rugulose, surface below paraterga microgranulate (Fig. 2 A – F). Postcollum metaterga with two transverse rows of rather long setae: 2 + 2 in anterior and 3 + 3 in posterior row (Fig. 2 A, C, F), the latter also borne on wrinkles and often abraded, but then readily traceable as insertion points. Tergal setae long, strong, slender, about 1 / 3 metatergal length (Fig. 2 A, D – F). Axial line visible both on pro- and metazona. Paraterga very strongly developed (Fig. 2 A – F), set high (at ca 1 / 4 metazonal height), upturned in ♂, lying below dorsum on rings 2 – 5 and 17 – 19, above dorsum on rings 6 – 16, in ♀ mostly below dorsum, rather thin in lateral view, a little thicker on pore-bearing rings (Fig. 2 B, D, E); anterior margin well-developed, mostly regularly rounded and narrowly bordered, fused to callus; caudal corner narrowly rounded, on postcollum rings extending increasingly past rear tergal margin, better so in ♂, nearly pointed to pointed, caudal tip on paraterga 16 – 19 evidently curved mesad (Fig. 2 E, F). Posterior margin of paraterga clearly concave, especially so in rings 15 – 19. Calluses on paraterga delimited by a sulcus only dorsally. Paraterga 2 broad, lateral margin with three small incisions, the one near caudal corner being particularly small (Fig. 2 A). Paraterga 3 and 4 with two small incisions at lateral margin, one at midway, the other at posterior 1 / 3; anterior incision particularly evident. Lateral margins of following paraterga often with a setigerous incision in anterior 1 / 3, being smaller on pore-bearing rings (Fig. 2 C). Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 metatergal length in front of posterior margin of metaterga (Fig. 2 B, D, E). Transverse sulcus usually distinct (Fig. 2 A, C, F), slightly incomplete on ring 4, complete and clearly visible on metaterga 5 – 18, narrow, rather deep, reaching the bases of paraterga, arcuate, beaded at bottom. Stricture between pro- and metazona narrow, deep, beaded at bottom down to base of paraterga (Fig. 2 A, C, F). Pleurosternal carinae complete crests on rings 2 and 3, a sharp caudal tooth on ring 4, the tooth gradually reduced into small, caudally roughly granulate crests until ring 7 (8), thereafter missing (♂, ♀) (Fig. 2 B, D). Epiproct (Fig. 2 E – G) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed ventrocaudally (Fig. 2 E, G); pre-apical papillae small, but evident, lying close to tip. Hypoproct subtriangular (Fig. 2 G), 1 + 1 setiferous knobs at caudal edge well-separated and evident. Sterna sparsely setose, without modifications; cross-impressions rather deep; a paramedian pair of evident, rounded, fully separated, setose cones between ♂ coxae 4 (Fig. 2 H, I). A conspicuous ridge in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3 – 1.5 (♂) (Fig. 2 B, F, G) or 1.1 – 1.3 times (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes present until legs of ring 17. Gonopods long, slender and suberect (Figs 3, 4). Coxite long and slender, slightly curved caudally, rather densely setose distodorsally (Figs 3 B, C, 4 A, B). Prefemoral part (pfe) densely setose, as usual, about 1 / 3 as long as acropodite (femoral + postfemoral parts) (Fig. 3 B, C). Femoral part (fe) long and slender, slightly curved and suberect distad, with a postfemoral part demarcated by an oblique lateral sulcus (Figs 3 B, 4 B). Solenophore (sph) trifid, its terminal lobule longest, middle prong spiniform and shorter than subterminal lobule (Figs 3, 4 A, C – F); solenomere (sl) long and flagelliform.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Thi, Sothearen, Sophea, Chhin, Lou, Vanny, Sinovas, Pablo, Jeratthitikul, Ekgachai, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Panha, Somsak, Sutcharit, Chirasak (2025): Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Cambodia, with new records and descriptions of three new species. ZooKeys 1251: 251-274, DOI: 10.3897/zookeys.1251.158776
