taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
632872F793A858A6A18B482B3EEB3773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434024	https://doi.org/10.3897/zookeys.1255.158380.figure1	Figure 1. Nyctimus bistriatus Thorell, 1877 from Borneo; habitus, dorsal view. A. Male (RMNH. ARA. 17873); B. Female (RMNH. ARA. 17875). Scale bars: 0.5 mm.	Figure 1. Nyctimus bistriatus Thorell, 1877 from Borneo; habitus, dorsal view. A. Male (RMNH. ARA. 17873); B. Female (RMNH. ARA. 17875). Scale bars: 0.5 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
632872F793A858A6A18B482B3EEB3773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434025	https://doi.org/10.3897/zookeys.1255.158380.figure2	Figure 2. Nyctimus bistriatus Thorell, 1877 female from Sumatra (2013 BF 2.1 AraThom 051 N 001). A. Habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	Figure 2. Nyctimus bistriatus Thorell, 1877 female from Sumatra (2013 BF 2.1 AraThom 051 N 001). A. Habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
632872F793A858A6A18B482B3EEB3773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434026	https://doi.org/10.3897/zookeys.1255.158380.figure3	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
632872F793A858A6A18B482B3EEB3773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10A23D924157560CB4FD69AF2B3C67A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434027	https://doi.org/10.3897/zookeys.1255.158380.figure4	Figure 4. Nyctimus falcatus Benjamin & Dhiya’ulhaq, sp. nov., male holotype (RMNH. ARA. 17861). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	Figure 4. Nyctimus falcatus Benjamin & Dhiya’ulhaq, sp. nov., male holotype (RMNH. ARA. 17861). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10A23D924157560CB4FD69AF2B3C67A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434039	https://doi.org/10.3897/zookeys.1255.158380.figure16	Figure 16. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	Figure 16. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10A23D924157560CB4FD69AF2B3C67A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434040	https://doi.org/10.3897/zookeys.1255.158380.figure17	Figure 17. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Left palp, ventral view; B. Left palp, retrolateral view. Abbreviations: DL = dorsal lobe; E = embolus; ML = medial lobe; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis; VL = ventral lobe. Scale bar: 0.2 mm.	Figure 17. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Left palp, ventral view; B. Left palp, retrolateral view. Abbreviations: DL = dorsal lobe; E = embolus; ML = medial lobe; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis; VL = ventral lobe. Scale bar: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10A23D924157560CB4FD69AF2B3C67A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
A719868B03C75999BD872DD47B99BD6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434028	https://doi.org/10.3897/zookeys.1255.158380.figure5	Figure 5. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq sp. nov., from Borneo. A. Male holotype (RMNH. ARA. 17862), prosoma, dorsal view; B. ibid., opisthosoma, dorsal view; C. Female (RMNH. ARA. 17859), habitus, dorsal view. Scale bars: 0.5 mm.	Figure 5. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq sp. nov., from Borneo. A. Male holotype (RMNH. ARA. 17862), prosoma, dorsal view; B. ibid., opisthosoma, dorsal view; C. Female (RMNH. ARA. 17859), habitus, dorsal view. Scale bars: 0.5 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
A719868B03C75999BD872DD47B99BD6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434029	https://doi.org/10.3897/zookeys.1255.158380.figure6	Figure 6. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Sumatra. A. Female (2013 _ BF 2.1 _ AraThom 004 N _ 001), habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	Figure 6. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Sumatra. A. Female (2013 _ BF 2.1 _ AraThom 004 N _ 001), habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
A719868B03C75999BD872DD47B99BD6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434030	https://doi.org/10.3897/zookeys.1255.158380.figure7	Figure 7. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Borneo. A. Male (RMNH. ARA. 17859), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17862), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 7. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Borneo. A. Male (RMNH. ARA. 17859), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17862), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
A719868B03C75999BD872DD47B99BD6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
5A830190ADB7526B8CE7B63CF6B773FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434031	https://doi.org/10.3897/zookeys.1255.158380.figure8	Figure 8. Nyctimus mutilloides Benjamin & Dhiya’ulhaq, sp. nov., male holotype (RMNH. ARA. 17882). A. Habitus, dorsal view; B. Left palp ventral view; C. Left palp retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	Figure 8. Nyctimus mutilloides Benjamin & Dhiya’ulhaq, sp. nov., male holotype (RMNH. ARA. 17882). A. Habitus, dorsal view; B. Left palp ventral view; C. Left palp retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
5A830190ADB7526B8CE7B63CF6B773FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434032	https://doi.org/10.3897/zookeys.1255.158380.figure9	Figure 9. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., male holotype (AraThom 001 N _ 2013 _ HF 4.2 _ 001). A. Habitus, dorsal view; B. Habitus ventral view; C. Right palp, mirrored, ventral view; D. Right palp, mirrored, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	Figure 9. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., male holotype (AraThom 001 N _ 2013 _ HF 4.2 _ 001). A. Habitus, dorsal view; B. Habitus ventral view; C. Right palp, mirrored, ventral view; D. Right palp, mirrored, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434033	https://doi.org/10.3897/zookeys.1255.158380.figure10	Figure 10. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001). A. Habitus, ventral view; B. Habitus, dorsal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	Figure 10. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001). A. Habitus, ventral view; B. Habitus, dorsal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434034	https://doi.org/10.3897/zookeys.1255.158380.figure11	Figure 11. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), right palp, mirrored, ventral view; B. ibid., RTA, ventro-retrolateral view; C. Female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VH = ventral hook. Scale bars: 0.2 mm.	Figure 11. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), right palp, mirrored, ventral view; B. ibid., RTA, ventro-retrolateral view; C. Female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VH = ventral hook. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434035	https://doi.org/10.3897/zookeys.1255.158380.figure12	Figure 12. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., non-type male specimen from Borneo (RMNH. ARA. 17857). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VH = ventral hook. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	Figure 12. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov., non-type male specimen from Borneo (RMNH. ARA. 17857). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VH = ventral hook. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434025	https://doi.org/10.3897/zookeys.1255.158380.figure2	Figure 2. Nyctimus bistriatus Thorell, 1877 female from Sumatra (2013 BF 2.1 AraThom 051 N 001). A. Habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	Figure 2. Nyctimus bistriatus Thorell, 1877 female from Sumatra (2013 BF 2.1 AraThom 051 N 001). A. Habitus, dorsal view; B. Prosoma, frontal view; C. Epigynum, ventral view; D. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434026	https://doi.org/10.3897/zookeys.1255.158380.figure3	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434037	https://doi.org/10.3897/zookeys.1255.158380.figure14	Figure 14. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 010 N 2013 BJ 6.1 003). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigynum, ventral view; D. Epigynum, dorsal view; E. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C – E).	Figure 14. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 010 N 2013 BJ 6.1 003). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigynum, ventral view; D. Epigynum, dorsal view; E. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C – E).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434038	https://doi.org/10.3897/zookeys.1255.158380.figure15	Figure 15. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 010 N 2013 BJ 6.1 001), left palp, ventral view; B. ibid., left palp, retrolateral view; C. Female paratype (AraThom 010 N _ 2013 _ BJ 6.1 _ 003), epigynum, dorsal view; D. ibid., epigynum, ventral view. Abbreviations: CO = copulatory opening; E = embolus; PE = posterior extension of spermatheca; RTA = retrolateral tibial apophysis; SP = spine; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 15. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 010 N 2013 BJ 6.1 001), left palp, ventral view; B. ibid., left palp, retrolateral view; C. Female paratype (AraThom 010 N _ 2013 _ BJ 6.1 _ 003), epigynum, dorsal view; D. ibid., epigynum, ventral view. Abbreviations: CO = copulatory opening; E = embolus; PE = posterior extension of spermatheca; RTA = retrolateral tibial apophysis; SP = spine; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
14EA4F9E30F15BC0907EE5D2595607ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
043239DA7E405B979B4129D6254CD462.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434036	https://doi.org/10.3897/zookeys.1255.158380.figure13	Figure 13. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., male holotype (AraThom 010 N _ 2013 _ BJ 6.1 _ 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	Figure 13. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., male holotype (AraThom 010 N _ 2013 _ BJ 6.1 _ 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
043239DA7E405B979B4129D6254CD462.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434037	https://doi.org/10.3897/zookeys.1255.158380.figure14	Figure 14. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 010 N 2013 BJ 6.1 003). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigynum, ventral view; D. Epigynum, dorsal view; E. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C – E).	Figure 14. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov., female paratype (AraThom 010 N 2013 BJ 6.1 003). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigynum, ventral view; D. Epigynum, dorsal view; E. Epigynum, dorsal view, cleared. Scale bars: 1 mm (A, B); 0.2 mm (C – E).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
043239DA7E405B979B4129D6254CD462.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434038	https://doi.org/10.3897/zookeys.1255.158380.figure15	Figure 15. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 010 N 2013 BJ 6.1 001), left palp, ventral view; B. ibid., left palp, retrolateral view; C. Female paratype (AraThom 010 N _ 2013 _ BJ 6.1 _ 003), epigynum, dorsal view; D. ibid., epigynum, ventral view. Abbreviations: CO = copulatory opening; E = embolus; PE = posterior extension of spermatheca; RTA = retrolateral tibial apophysis; SP = spine; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 15. Nyctimus rendang Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 010 N 2013 BJ 6.1 001), left palp, ventral view; B. ibid., left palp, retrolateral view; C. Female paratype (AraThom 010 N _ 2013 _ BJ 6.1 _ 003), epigynum, dorsal view; D. ibid., epigynum, ventral view. Abbreviations: CO = copulatory opening; E = embolus; PE = posterior extension of spermatheca; RTA = retrolateral tibial apophysis; SP = spine; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
043239DA7E405B979B4129D6254CD462.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434034	https://doi.org/10.3897/zookeys.1255.158380.figure11	Figure 11. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), right palp, mirrored, ventral view; B. ibid., RTA, ventro-retrolateral view; C. Female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VH = ventral hook. Scale bars: 0.2 mm.	Figure 11. Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin, sp. nov. A. Male holotype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), right palp, mirrored, ventral view; B. ibid., RTA, ventro-retrolateral view; C. Female paratype (AraThom 001 N _ 2013 _ HF 3.2 _ 001), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VH = ventral hook. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
043239DA7E405B979B4129D6254CD462.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
CD696F8764635550B8478967D3B46BE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434039	https://doi.org/10.3897/zookeys.1255.158380.figure16	Figure 16. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	Figure 16. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left palp, ventral view; D. Left palp, retrolateral view. Scale bars: 1 mm (A, B); 0.2 mm (C, D).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
CD696F8764635550B8478967D3B46BE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434040	https://doi.org/10.3897/zookeys.1255.158380.figure17	Figure 17. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Left palp, ventral view; B. Left palp, retrolateral view. Abbreviations: DL = dorsal lobe; E = embolus; ML = medial lobe; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis; VL = ventral lobe. Scale bar: 0.2 mm.	Figure 17. Nyctimus saksang Dhiya’ulhaq & Benjamin, sp. nov. male holotype (AraThom 036 N 2013 HF 2.1 001). A. Left palp, ventral view; B. Left palp, retrolateral view. Abbreviations: DL = dorsal lobe; E = embolus; ML = medial lobe; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis; VL = ventral lobe. Scale bar: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
CD696F8764635550B8478967D3B46BE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10DC5F151D155FB7AE1FEDF552C40744.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434024	https://doi.org/10.3897/zookeys.1255.158380.figure1	Figure 1. Nyctimus bistriatus Thorell, 1877 from Borneo; habitus, dorsal view. A. Male (RMNH. ARA. 17873); B. Female (RMNH. ARA. 17875). Scale bars: 0.5 mm.	Figure 1. Nyctimus bistriatus Thorell, 1877 from Borneo; habitus, dorsal view. A. Male (RMNH. ARA. 17873); B. Female (RMNH. ARA. 17875). Scale bars: 0.5 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10DC5F151D155FB7AE1FEDF552C40744.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434026	https://doi.org/10.3897/zookeys.1255.158380.figure3	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
10DC5F151D155FB7AE1FEDF552C40744.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434030	https://doi.org/10.3897/zookeys.1255.158380.figure7	Figure 7. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Borneo. A. Male (RMNH. ARA. 17859), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17862), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	Figure 7. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Borneo. A. Male (RMNH. ARA. 17859), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17862), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
6487E2EB928A57BE84E8F71297687052.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
68082F374A2F5D83A03EFE75127061E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434041	https://doi.org/10.3897/zookeys.1255.158380.figure18	Figure 18. Zametopina wanliae Lin & Li, 2023, male (MCZ 6652). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: DP = denticulate process; E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	Figure 18. Zametopina wanliae Lin & Li, 2023, male (MCZ 6652). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: DP = denticulate process; E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
83804B747CD85BBE9E96196B32D20079.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434041	https://doi.org/10.3897/zookeys.1255.158380.figure18	Figure 18. Zametopina wanliae Lin & Li, 2023, male (MCZ 6652). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: DP = denticulate process; E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	Figure 18. Zametopina wanliae Lin & Li, 2023, male (MCZ 6652). A. Habitus, dorsal view; B. Left palp, ventral view; C. Left palpal tibia, retrolateral view. Abbreviations: DP = denticulate process; E = embolus; RTA = retrolateral tibial apophysis; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.5 mm (A); 0.2 mm (B, C).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
83804B747CD85BBE9E96196B32D20079.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1434042	https://doi.org/10.3897/zookeys.1255.158380.figure19	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).	2025-10-09	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen		Zenodo	biologists	Benjamin, Suresh P.;Dhiya’ulhaq, Naufal Urfi;Deeleman-Reinhold, Christa;Buchori, Damayanti;Hidayat, Purnama;Scheu, Stefan;Drescher, Jochen			
