taxonID	type	description	language	source
632872F793A858A6A18B482B3EEB3773.taxon	description	Figs 1, 2, 3	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
632872F793A858A6A18B482B3EEB3773.taxon	diagnosis	Diagnosis. Males can be easily distinguished from all other congeners by the longer embolus, more than length of cymbium (in all congeners, the embolus is at most half the length of cymbium; Fig. 3 A, B) and the denticulate RTA. Females can be easily distinguished by the single CO; CD arched, resembling a rams’ horn, more than twice the length of spermathecae (in all congeners, the CO is paired, and the CD are not longer than spermathecae; Figs 2 C, D, 3 C, D).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
632872F793A858A6A18B482B3EEB3773.taxon	description	Description. Male (RMNH. ARA. 17873; Figs 1 A, 3 A, B). Total length 2.85. Prosoma length 1.42; width 1.26. Opisthosoma length 1.43; width 1.17. Diameter of eyes: AME 0.09; ALE 0.15; PLE 0.14 PME 0.07. Interdistances between eyes: AME – AME 0.50; AME – ALE 0.10; ALE – ALE 0.86; PME – PME 0.70; PME – PLE 0.20; ALE – PLE 0.18; AME – PME 0.13; PLE – PLE 1.09. Leg measurements: leg I 3.3 (1.0, 0.5, 1.0, 0.8, 0.5). Prosoma cuboid, almost as long as wide; dorsally blackish brown, surface granulated; laterally with two rows of long, curved macrosetae originating on small tubercles. All eyes are very far apart, especially between the median eyes. Chelicerae, labium and sternum blackish brown; sternum with small dents. Legs dark brown except femora III and IV, which are yellow; femora and tibiae I – IV with two long spines; patellae I – IV with one distal long spine. Opisthosoma dorsally greyish black with many long setae; five large sigillae present, arranged in a triangular formation; venter greyish black. Palp (Fig. 3 A, B): cymbium oval. Embolus long, whip-like, looping over the retrolateral side of bulb, half of its length covered by the cymbium. Tegular hood small, located close to the base of bulb. RTA large, roughly trapezoid, laterally protruding, distal half with denticles on all sides. VTA long, digitiform; apically flexed with tip ventrally curved. Female (RMNH. ARA. 17875; Figs 1 B, 3 C, D). Total length 3.32. Prosoma length 1.15; width 1.30. Opisthosoma length 2.17; width 1.95. Diameter of eyes: AME 0.08; ALE 0.13; PLE 0.16; PME 0.06. Interdistances between eyes: AME – AME 0.56; AME – ALE 0.10; ALE – ALE 0.86; PME – PME 0.64; PME – PLE 0.21; ALE – PLE 0.18; AME – PME 0.16; PLE – PLE 1.09. Leg measurements: leg I 5.3 (1.6, 0.6, 1.3, 1.0, 0.8). Habitus as in male, except opisthosoma dorsally lighter than male and the presence of white spots as shown in Fig. 1 B. Epigynum (Figs 2 C, D, 3 C, D): exterior plate roughly pentagonal. Atrium inconspicuous. CO oval, single, large, CD long; divided into a very wide, ramshorn-shaped initial part and a narrow, C-shaped later part. Spermatheca globular, approximately as wide as CO; anteriorly with a small bump; posteriorly extended.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
632872F793A858A6A18B482B3EEB3773.taxon	distribution	Distribution. Brunei, Malaysia (Borneo: Sabah), Indonesia (Sumatra: Jambi; Sulawesi: Southeast Sulawesi) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10A23D924157560CB4FD69AF2B3C67A8.taxon	description	Fig. 4	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10A23D924157560CB4FD69AF2B3C67A8.taxon	etymology	Etymology. The species epithet falcatus is derived from the Latin word falx / falcis, meaning “ sickle ” or “ scythe. ” It refers to the distinctly curved, sickle-shaped embolus of the male palp, a diagnostic feature of the species.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10A23D924157560CB4FD69AF2B3C67A8.taxon	diagnosis	Diagnosis. Males are very similar to N. saksang sp. nov., in having a similar sickle-shaped embolus, but they can be easily distinguished from it by the large, digitiform VTA, attached distally on the RTA (the corresponding part in N. saksang, sp. nov., is much smaller and attached medially on the ventral lobe of RTA; Fig. 4 B vs Figs 16 C, 17 A).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10A23D924157560CB4FD69AF2B3C67A8.taxon	description	Description. Male (holotype RMNH. ARA. 17861; Fig. 4). Total length: 2.36. Prosoma length 1.23; width: 1.05. Opisthosoma length 1.13; width 1.02. Diameter of eyes: AME 0.08; ALE 0.15; PLE 0.14; PME 0.05. Interdistances between eyes: AME – AME 0.27; AME – ALE 0.08; ALE – ALE 0.59; PME – PME 026; PME – PLE 0.27; ALE – PLE 0.18; AME – PME 0.14; PLE – PLE 0.79. Prosoma cuboid, almost as long as wide, dark reddish brown; surface weakly granulated; laterally adorned with long macrosetae, each arising from a small tubercle. Chelicerae and sternum coloured as prosoma. Front legs brown; back legs brown except for the white femora III and IV. Opisthosoma round, light brown; dorsally covered with scutum; five large sigillae present, arranged in a triangular formation. Palp (Fig. 4 B, C): cymbium oval. Embolus sickle-shaped. Tegulum round, with a small triangular projection next to the embolus; tegular hood small. RTA wide trapezoid; VTA large, digitiform, attached distally on the RTA. Female. Unknown.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10A23D924157560CB4FD69AF2B3C67A8.taxon	distribution	Distribution. Only known from the type locality (Thailand: Phang Nga) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
A719868B03C75999BD872DD47B99BD6E.taxon	description	Figs 5, 6, 7	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
A719868B03C75999BD872DD47B99BD6E.taxon	etymology	Etymology. The specific epithet is derived from the name of the type locality.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
A719868B03C75999BD872DD47B99BD6E.taxon	diagnosis	Diagnosis. Males can be easily distinguished from all other congeners by the embolic base abruptly bent 90 ° towards the retrolateral side of the cymbium and the tapered, bent embolus (Fig. 7 A, B). Females are distinguishable by the long, oval spermatheca (globular or short-oval in all other congeners; Figs 6 C, D, 7 C, D).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
A719868B03C75999BD872DD47B99BD6E.taxon	description	Description. Male (holotype RMNH. ARA. 17862; Figs 5 A, B, 7 A, B). Total length 3.24. Prosoma length 1.61; width 1.50. Opisthosoma length 1.63; width 1.31. Diameter of eyes: AME 0.11; ALE 0.20; PLE 0.17; PME 0.08. Interdistances between eyes: AME – AME 0.44; AME – ALE 0.11; ALE – ALE 0.81; PME – PME 0.45; PME – PLE 0.36; ALE – PLE 0.21; AME – PME 0.17; PLE – PLE 1.15. Leg measurements: leg I 5.0 (1.5, 0.6, 1.3, 1.0, 0.6). Prosoma cuboid, almost as long as wide, dark reddish brown; surface weakly granulated; laterally adorned with long macrosetae, each arising from a small tubercle. Chelicerae and sternum coloured as prosoma. Endites brown with white tip. Front legs brown; back legs brown except for the white femora. Opisthosoma oval, greyish brown with white spots (most dense in the anterior half) and paired white bars in the middle; anteriorly with a white border; five large sigillae present, arranged in a triangular formation. Palp (Fig. 7 A, B): cymbium oval. Embolus elongated, tapering, with a 150 ° bend in the middle. Tegular hood small. RTA bilobed; ventral lobed very large, claw-shaped; dorsal lobe much smaller than ventral lobed, roughly triangular with an obtuse tip and a small triangular projection; VTA hook-shaped, small, attached medially on the RTA. Female (RMNH. ARA. 17859; Figs 5 C, 7 C, D). Total length: 3.38. Prosoma length 1.48; width 1.27. Opisthosoma length 1.90; width 1.70. Diameter of eyes: AME 0.07; ALE 0.17; PLE 0.14; PME 0.06. Interdistances between eyes: AME – AME 0.38; AME – ALE 0.12; ALE – ALE 0.74; PME – PME 0.39; PME – PLE 0.33; ALE – PLE 0.22; AME – PME 0.17; PLE – PLE 0.98. Leg measurements: leg I 4.6 (1.3, 0.4, 1.1, 1.0, 0.8). Habitus as in male. Epigynum (Fig. 7 C, D): exterior plate large, roughly square with rounded sides. Atrium oval. CO diagonally oriented, anteriorly facing. CD arched. Spermatheca long oval, with a slight bump anteriorly.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
A719868B03C75999BD872DD47B99BD6E.taxon	distribution	Distribution. Brunei; Malaysia (Borneo: Sabah); Indonesia (Sumatra: Jambi) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
5A830190ADB7526B8CE7B63CF6B773FB.taxon	description	Fig. 8	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
5A830190ADB7526B8CE7B63CF6B773FB.taxon	etymology	Etymology. The specific epithet is derived from the latin name of the velvet ant family Mutillidae, and the Greek-derived suffix - oides, meaning “ resembling ” or “ like. ” Thus, mutilloides translates to “ mutillid-like, ” referring to the spider’s striking abdominal patterns resembling abdominal markings of certain trogaspidiine mutillids.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
5A830190ADB7526B8CE7B63CF6B773FB.taxon	diagnosis	Diagnosis. Nyctimus mutilloides, sp. nov., can be readily distinguished from its congeners by its vivid white markings on the opisthosoma, forming a pattern highly suggestive of mutillid wasps (spots on all congeners). Furthermore, the male palp has a claw-shaped embolus that juts out from the distal margin of the bulb (different in other congeners; Fig. 8 B, C).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
5A830190ADB7526B8CE7B63CF6B773FB.taxon	description	Description. Male (holotype RMNH. ARA. 17882; Fig. 8). Total length 2.80. Prosoma length 1.22; width 1.23. Opisthosoma length 1.57; width 1.22. Diameter of eyes: AME 0.07; ALE 0.12; PLE 0.10; PME 0.08. Interdistances between eyes: AME – AME 0.16; AME – ALE 0.11; ALE – ALE 0.39; PME – PME 0.2; PME – PLE 0.18; ALE – PLE 0.10; AME – PME 0.07; PLE – PLE 0.69. Prosoma cuboid, almost as long as wide, slightly flattened, weakly granulated, dark brown with darker margins, laterally adorned with long macrosetae; cephalic region elevated. Legs slender, pale yellow with indistinct annulations. No conspicuous spines observed. Opisthosoma oval, dorsoventrally flattened, with white spots and bands on black background. Palp (Fig. 8 B, C): cymbium slender, oval; tip rounded. Embolus claw-shaped, curved with a pointed tip, emerging distally from the tegulum. Tegulum oval; tegular hood wide, located retrolaterally near the base. RTA bilobed; both lobes roughly triangular; dorsal lobe with a serrated edge. VTA short, thumb-shaped, and slightly curved. Female. Unknown.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
5A830190ADB7526B8CE7B63CF6B773FB.taxon	distribution	Distribution. Known only from the type locality (Malaysia: Borneo: Sabah) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
14EA4F9E30F15BC0907EE5D2595607ED.taxon	description	Figs 9, 10, 11, 12	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
14EA4F9E30F15BC0907EE5D2595607ED.taxon	etymology	Etymology. The specific epithet is taken from Latin, referring to the four white spots on the opisthosoma.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
14EA4F9E30F15BC0907EE5D2595607ED.taxon	diagnosis	Diagnosis. Males are easily distinguished from other congeners by the anvil-shaped RTA ending in two acute lobes (Fig. 11 A, B). Females can be distinguished from N. bistriatus by the much thinner, shorter CD and paired CO (Figs 10 C, D, 11 C, D; vs single CO in N. bistriatus, Figs 2 C, 3 C), and from N. rendang, sp. nov., by the spermatheca lacking a prominent posterior extension (Figs 10 D, 11 D; vs Figs 14 D, 15 D).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
14EA4F9E30F15BC0907EE5D2595607ED.taxon	description	Description. Male (holotype 2013 _ HF 4.2 _ AraThom 001 N _ 001; Figs 9, 11 A, B). Total length 2.95. Prosoma length 1.35; width 1.23. Opisthosoma length 1.60; width 1.22. Diameter of eyes: AME 0.07; ALE 0.16; PLE 0.16; PME 0.06. Interdistances between eyes: AME – AME 0.38; AME – ALE 0.08; ALE – ALE 0.68; PME – PME 0.34; PME – PLE 0.32; ALE – PLE 0.19; AME – PME 0.18; PLE – PLE 0.98. Clypeus height 0.00. Leg measurements: leg I 4.25 (1.19, 0.49, 1.06, 0.90, 0.61); leg II 3.89 (1.11, 0.42, 0.90, 0.81, 0.65); leg III 2.63 (0.86, 0.35, 0.57, 0.49, 0.36); leg IV 2.74 (0.83, 0.27, 0.66, 0.52, 0.46). Prosoma cuboid, almost as long as wide, box-shaped, weakly granulated, highest at mid-point, light brown; lateral margins furnished with a row of long macrosetae. Lateral eyes large, situated on tubercles on the lateral edge of the prosoma. Chelicerae, endites, and labium light brown. Sternum light brown. Legs I light brown; others yellow-brown, with sparse spines and black setae. Opisthosoma oval, brown, with two pairs of large white spots. Spinnerets brown. Palp (Figs 9 C, D, 11 A, B): cymbium oval. Embolus sickle-shaped; base slightly wrinkled. Tegular hood very wide. RTA large, rectangular, anvil-shaped; dorsally bifurcated, ending into two long, acute lobes, hidden behind a brush of setae; VTA hook-shaped attached medially on RTA. Female (paratype 2013 _ HF 3.2 _ AraThom 001 N _ 002; Figs 10, 11 C, D). Total length 4.81. Prosoma length 1.95; width 1.77. Opisthosoma length 2.86; width 2.65. Diameter of eyes: AME 0.09; ALE 0.20; PLE 0.18; PME 0.07. Interdistances between eyes: AME – AME 0.53; AME – ALE 0.14; ALE – ALE 0.97; PME – PME 0.52; PME – PLE 0.39; ALE – PLE 0.23; AME – PME 0.24; PLE – PLE 1.37. Leg measurements: leg I 5.47 (1.66, 0.57, 1.34, 1.13, 0.77); leg II -; leg III 3.69 (1.22, 0.47, 0.87, 0.63, 0.50); leg IV 4.12 (1.36, 0.46, 0.96, 0.76, 0.58). Habitus as in males, except prosoma and leg I reddish brown. Epigynum (Figs 10 C, D, 11 C, D): exterior plate large, hexagonal-shaped with wide anterior side. Atrium conspicuous, inverted heart-shaped with a separation in the middle. CO diagonally oriented, posteriorly facing. CD arch-shaped. Spermatheca elongated, peanut-shaped, with a small bump anteriorly.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
14EA4F9E30F15BC0907EE5D2595607ED.taxon	distribution	Distribution. Malaysia (Borneo: Sabah); Indonesia (Sumatra: Jambi) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
043239DA7E405B979B4129D6254CD462.taxon	description	Figs 13, 14, 15	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
043239DA7E405B979B4129D6254CD462.taxon	etymology	Etymology. The specific epithet refers to a meat dish associated with the cuisine of the Minangkabau people from West Sumatra. The meat is slow-cooked in a plethora of spices, giving it a dark colouration. The dark colour of N. rendang, sp. nov., is reminiscent of the colour of this dish.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
043239DA7E405B979B4129D6254CD462.taxon	diagnosis	Diagnosis. Males are similar to N. quadripunctatus, sp. nov., in having a sickle-shaped embolus and in the general shape of the bulb, but the VTA is digitiform and free (Fig. 15 A, B; vs hook-shaped, attached to RTA in N. quadripunctatus, Fig. 11 A, B) and the RTA possesses a pillar-shaped ventral lobe and paired, thick spines (vs both absent). Females are distinct in having the spermathecae with a posterior extension and a short, barely noticeable CD (Fig. 11 D).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
043239DA7E405B979B4129D6254CD462.taxon	description	Description. Male (holotype 2013 _ BJ 6.1 _ AraThom 010 N _ 001; Figs 13, 15 A, B). Total length 2.41. Prosoma length 1.16; width 1.03. Opisthosoma length 1.25; width 1.21. Diameter of eyes: AME 0.07; ALE 0.15; PLE 0.14; PME 0.05. Interdistances between eyes: AME – AME 0.35; AME – ALE 0.08; ALE – ALE 0.62; PME – PME 0.32; PME – PLE 0.24; ALE – PLE 0.17; AME – PME 0.19; PLE – PLE 0.77. Leg measurements: leg I 3.36 (0.90, 0.29, 0.92, 0.75, 0.50); leg II 3.46 (1.04, 0.32, 0.84, 0.75, 0.51); leg III 2.11 (0.67, 0.20, 0.54, 0.38, 0.32); leg IV 2.25 (0.75, 0.18, 0.59, 0.42, 0.31). Prosoma cuboid, almost as long as wide, dark reddish brown; surface weakly granulated; laterally adorned with long macrosetae, each arising from a small tubercle. Chelicerae, and sternum coloured as prosoma. Endites brown with white tip. Front legs brown; back legs brown except for the white femora. Opisthosoma round, dark brown with paired white bars in the middle; dorsally covered with scutum; five large sigillae present, arranged in a triangular formation. Palp (Figs 13 C, D, 15 A, B): cymbium oval. Embolus sickle-shaped. Tegulum with a small triangular projection next to the embolus; tegular hood small. RTA bilobed; ventral lobe finger-shaped; dorsal lobe wide trapezoid, apically with a pair of short, thick spines. VTA long, finger-shaped, apically flexed. Female (paratype AraThom 010 N _ 2013 _ BJ 6.1 _ 003; Figs 14, 15 C, D). Total length 3.86. Prosoma length 1.67; width 1.43. Opisthosoma length 2.19; width 2.21. Diameter of eyes: AME 0.08; ALE 0.18; PLE 0.19; PME 0.07. Interdistances between eyes: AME – AME 0.55; AME – ALE 0.12; ALE – ALE 0.90; PME – PME 0.45; PME – PLE 0.34; ALE – PLE 0.23; AME – PME 0.23; PLE – PLE 1.11. Leg measurements: leg I 4.85 (0.71, 1.01, 1.18, 0.49, 1.46); leg II -; leg III 3.05 (1.00, 0.30, 0.75, 0.54, 0.46); leg IV 3.41 (1.14, 0.32, 0.83, 0.65, 0.47). Habitus as in male, except opisthosoma not covered in scutum. Epigynum (Fig. 14 C, D, 15 C, D): exterior plate with irregular border. Atrium inconspicuous. CO semicircular, posteriorly facing. CD very short. Spermatheca oval, anteriorly with a large bump, posteriorly extended.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
043239DA7E405B979B4129D6254CD462.taxon	distribution	Distribution. Indonesia (Sumatra: Jambi) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
CD696F8764635550B8478967D3B46BE2.taxon	description	Figs 16, 17	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
CD696F8764635550B8478967D3B46BE2.taxon	etymology	Etymology. The specific epithet refers to a meat-stew dish associated with the Batak people from North Sumatra. The meat is cooked in blood, giving it a dark colouration. The dark colour of Nyctimus saksang Benjamin & Dhiya’ulhaq, sp. nov., is reminiscent of the colour of this dish.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
CD696F8764635550B8478967D3B46BE2.taxon	diagnosis	Diagnosis. Males can be easily distinguished from all other congeners by the very wide, axe-shaped RTA which consists of three large lobes (Figs 16 C, D, 17).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
CD696F8764635550B8478967D3B46BE2.taxon	description	Description. Male (holotype 2013 _ HF 2.1 _ AraThom 036 N _ 001; Figs 16, 17). Total length 3.11. Prosoma length 1.50; width 1.25. Opisthosoma length 1.61; width 1.40. Diameter of eyes: AME 0.08; ALE 0.17; PLE 0.14; PME 0.05. Interdistances between eyes: AME – AME 0.40; AME – ALE 0.10; ALE – ALE 0.73; PME – PME 0.38; PME – PLE 0.33; ALE – PLE 0.21; AME – PME 0.22; PLE – PLE 0.95. Leg measurements: leg I 4.20 (1.20, 0.44, 1.05, 0.89, 0.62); leg II 3.74 (1.10, 0.34, 0.91, 0.78, 0.61); leg III 2.78 (0.83, 0.37, 0.67, 0.52, 0.39); leg IV 2.87 (0.88, 0.26, 0.70, 0.60, 0.43). Prosoma cuboid, almost as long as wide, dark reddish brown; surface weakly granulated; laterally adorned with long macrosetae, each arising from a small tubercle. Chelicerae and sternum coloured as prosoma. Endites brown with white tip. Front legs brown; back legs brown except for the white femora. Opisthosoma round, dark brown with paired white bars in the middle that are almost touching each other; anteriorly with a white border; posteriorly with a white patch; dorsally covered with scutum; five large sigillae present, arranged in a triangular formation. Palp (Figs 16 C, D, 17): cymbium oval. Embolus elongated, tapering; tip with a slight flange. Tegular hood small. RTA trilobed; ventral lobed finger-shaped; median lobe triangular, with dorsal lobe axe-shaped; dorsal edge slightly folded. VTA hook-shaped, small, attached medially on the ventral lobe of RTA. Female. Unknown.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
CD696F8764635550B8478967D3B46BE2.taxon	distribution	Distribution. Only known from the type locality (Indonesia: Sumatra: Jambi) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10DC5F151D155FB7AE1FEDF552C40744.taxon	type_taxon	Type species. Nyctimus bistriatus Thorell, 1877.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
10DC5F151D155FB7AE1FEDF552C40744.taxon	diagnosis	Diagnosis. Small thomisids, 2 – 5 mm in body length. Prosoma cuboid, as long as wide, with granulated surface and bearing long macrosetae (Fig. 1). Body colouration dark with white / yellow markings, mostly in the form of spots and bars. Opisthosoma with five large sigillae arranged in a triangular formation Embolus is weakly curved and possesses a thick base. Tegulum with a “ hood ” (Fig. 3 A). Tibial apophyses are well developed, VTA present, either free (Fig. 3 A) or hook-shaped and attached to RTA (Fig. 7 A). Epigynum highly variable, spermatheca rounded or oval, single pair (Fig. 3 C, D).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
6487E2EB928A57BE84E8F71297687052.taxon	diagnosis	Diagnosis. See Lin et al. (2023).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
6487E2EB928A57BE84E8F71297687052.taxon	description	Description. See Tang et al. (2010).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
6487E2EB928A57BE84E8F71297687052.taxon	distribution	Distribution. China (Yunnan), Vietnam (Cao B ằng) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
68082F374A2F5D83A03EFE75127061E0.taxon	type_taxon	Type species. Zametopina calceata Simon, 1909.	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
68082F374A2F5D83A03EFE75127061E0.taxon	diagnosis	Diagnosis. Zametopina is similar to Nyctimus in habitus, differing in the presence of a unique denticulate process next to the embolus on the male palp (DP in Fig. 18 B). No other oriental thomisid genus possesses this structure. For a detailed diagnosis see Tang et al. (2010).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
83804B747CD85BBE9E96196B32D20079.taxon	description	Fig. 18	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
83804B747CD85BBE9E96196B32D20079.taxon	diagnosis	Diagnosis. See Lin et al. (2023).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
83804B747CD85BBE9E96196B32D20079.taxon	description	Description. Male (MCZ 6652; Fig. 18). Total length 2.44. Prosoma length 1.22; width 1.14. Opisthosoma length 1.22; width 1.40. Diameter of eyes: AME 0.09; ALE 0.15; PLE 0.12; PME 0.09. Interdistances between eyes: AME – AME 0.30; AME – ALE 0.13; ALE – ALE 0.69; PME – PME 0.37; PME – PLE 0.35; ALE – PLE 0.23; AME – PME 0.07; PLE – PLE 0.85. For a complete description, see Lin et al. (2023).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
83804B747CD85BBE9E96196B32D20079.taxon	distribution	Distribution. Vietnam (Vĩnh Phúc), Thailand (Chaiyaphum [new record]) (Fig. 19).	en	Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan, Drescher, Jochen (2025): Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia. ZooKeys 1255: 95-125, DOI: 10.3897/zookeys.1255.158380
