identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4155C6671A73AC314243FC7BFE13FE5B.text	4155C6671A73AC314243FC7BFE13FE5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eudolium subintermedium (d'Orbigny 1852)	<div><p>Eudolium subintermedium (d’Orbigny, 1852)</p><p>(Figs 2–3)</p><p>1852 Cassis subintermedia – D’ORBIGNY, p. 90.</p><p>1966 Cassidaria cingulifera Hoernes &amp; Auinger – STRAUSZ, p. 241, fig. 116.</p><p>1969 b Cassidaria cingulifera Hoernes et Auinger – CSEPREGHY-MEZNERICS, pl. 7, fig. 2.</p><p>2009 Eudolium subintermedium (d’Orbigny) – LANDAU et al., p. 63, pl. 1, figs 1–6 (cum syn.).</p><p>Material – One fragmentary specimen (SL 20); HNHM: INV 2017.663. (Figs 2–3) .</p><p>Description – Globose shell, low spire, two convex teleoconch whorls preserved, last whorl shouldered by a slightly nodulose primary cord, constricted at the base, ovate aperture. Spiral sculpture of four narrow primary cords with three fine spiral threads in interspaces on the penultimate whorl, last whorl bears nine primary cords with secondary cords and threads in interspaces.</p><p>Remarks – The taxonomic revision of the species was accomplished by LANDAU et al. (2009), Cassidaria cingulifera was considered as a synonym of the E. subintermedium . The species is very rare in Hungarian localities. The subadult specimen figured here with weakly nodulose sculpture is closely allied in morphology to the specimen illustrated by HOERNES &amp; AUINGER (1884, pl. 17, fig. 17, refigured by LANDAU et al. 2009, pl. 1, fig. 1).</p><p>Distribution – Badenian: CP (Austria, Bulgaria, Hungary, Poland, Romania), Langhian: NEA (France), Tortonian: PMS (Italy). For Pliocene range see LANDAU et al. (2009).</p></div>	https://treatment.plazi.org/id/4155C6671A73AC314243FC7BFE13FE5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A72AC3142CCFD84FB89FB5A.text	4155C6671A72AC3142CCFD84FB89FB5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malea orbiculata (Brocchi 1814)	<div><p>Malea orbiculata (Brocchi, 1814)</p><p>(Figs 4–5)</p><p>1814 Buccinum orbiculatum – BROCCHI, p. 647, pl. 15, fig. 22.</p><p>1966 Dolium orbiculatum Brocchi – STRAUSZ, p. 253, fig. 120, pl. 64, fig. 7.</p><p>2013 Malea orbiculata (Brocchi) – LANDAU et al., p. 123, pl. 17, fig. 1 (cum syn.).</p><p>Material – Ten specimens (SL 20–42); in private collections.</p><p>Description – Globose shell, low spire, four convex teleoconch whorls, last whorl constricted at the base. Elongate aperture, denticulate outer lip thickened by labral varix, excavated columella with folds, short and wide siphonal canal, short fasciole. Spiral sculpture of broad, slightly rounded cords (16 on the last whorl), narrow interspaces, axial sculpture of fine growth lines.</p><p>Remarks – The species was frequently cited as M. denticulata (Deshayes) in the literature. According to the revision of orbiculata (LANDAU et al. 2004), Brocchi’s specimen represents a juvenile form, therefore denticulata is a junior synonym. M. orbiculata is a rare element of the mollusc assemblage of Letkés.</p><p>Distribution – Karpatian: CP (Hungary), Langhian: NEA (France), Badenian: CP (Austria, Hungary, Poland), Serravallian: PMS (Turkey), Tortonian: NEA (Portugal), PMS (Italy). For Pliocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A72AC3142CCFD84FB89FB5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A72AC3642D3FACEFE51FCDF.text	4155C6671A72AC3642D3FACEFE51FCDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cassis postmamillaris Sacco, HNHM INV 1890	<div><p>Cassis postmamillaris Sacco, 1890</p><p>(Figs 6–9)</p><p>1890 Cassis postmamillaris Sacco – SACCO, p. 16, pl. 1, fig. 11.</p><p>2013 Cassis postmamillaris Sacco – LANDAU et al., p. 123, pl. 17, figs 2–4, pl. 79, fig. 8 (cum syn.). 2017 Cassis postmamillaris Sacco – VICIÁN et al., p. 267, pl. 1, figs 13–14.</p><p>Material – 167 specimens (SL 10–83); HNHM: INV 2017.664. (Figs 6–9), INV 2017.665. (14), 152 specimens in private collections.</p><p>Description – Large, helmet-shaped shell, low spire, last whorl more than 90% of the total height, constricted at base. Elongate aperture, thickened, denticulate outer lip, columella with folds, broad, flattened, thickened parietal callus forming triangular apertural shield, deep, recurved siphonal canal. Spiral sculpture of two or three rows of rounded, slightly pointed tubercles, axial sculpture of weakly developed ribs and growth lines.</p><p>Remarks – Although the Oligocene C. mamillaris Grateloup was distinguished from the Miocene C. postmamillaris by SACCO (1890), it was overlooked in the Hungarian literature, and the Miocene species was recorded as C. mamillaris from the N Börzsöny and E Cserhát Mts (NOSZKY 1925, 1940). C. postmamillaris is relatively abundant in the mollusc assemblage studied here.</p><p>Distribution – Langhian: PMS (Italy), Badenian: CP (Austria, Czech Republic, Hungary, Romania, Slovakia), Serravallian: PMS (Greece, Turkey). Tortonian: PMS (Italy).</p></div>	https://treatment.plazi.org/id/4155C6671A72AC3642D3FACEFE51FCDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A75AC36429CFC00FDBCF9C7.text	4155C6671A75AC36429CFC00FDBCF9C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Galeodea echinophora (Linnaeus 1758)	<div><p>Galeodea echinophora (Linnaeus, 1758)</p><p>(Fig. 10)</p><p>1758 Buccinum echinophorum – LINNAEUS, p. 735.</p><p>1969 a Cassidaria echinophora pliotriseriata Sacco – CSEPREGHY-MEZNERICS, p. 78, pl. 2, figs 21–22. 2004 Galeodea echinophora (Linnaeus) – LANDAU et al., p. 43, pl. 2, fig. 2 (cum syn.).</p><p>2009 Galeodea echinophora (Linnaeus) – LANDAU et al., p. 66, pl. 3, figs 1–2 (cum syn.).</p><p>Material – Six specimens (SL 36–48); in private collections .</p><p>Description – Fusiform shell, moderately elevated spire, five subangulate teleoconch whorls with slightly concave sutural ramp, convex, expanded last whorl. Ovate aperture, denticulate outer lip thickened by labral varix, wide pseudoumbilicus. Spiral sculpture of fine cords, nodulose shoulder, two rows of nodules on the last whorl, axial sculpture of fine growth lines.</p><p>Remarks – The species has a large but fragile shell, so complete shells are very rare in the resedimented layers of the Letkés section.</p><p>Distribution – Burdigalian: PMS (Italy), NSB (Germany), Eggenburgian: CP (Germany), late Karpatian: CP (Slovakia), Badenian: CP (Austria, Bulgaria, Hungary, Romania), Tortonian: NEA (Portugal), PMS (Italy). For Pliocene range see VAN DINGENEN et al. (2016).</p></div>	https://treatment.plazi.org/id/4155C6671A75AC36429CFC00FDBCF9C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A74AC3742A7FEF4FE04FBAB.text	4155C6671A74AC3742A7FEF4FE04FBAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Echinophoria haueri (Hoernes et Auinger 1884)	<div><p>Echinophoria haueri (Hoernes et Auinger, 1884)</p><p>(Fig. 11)</p><p>1884 Cassis (b. Cassidea) Haueri – HOERNES &amp; AUINGER, p. 158, pl. 17, fig. 13.</p><p>1969 a Cassidaria haueri M. Hörnes – CSEPREGHY-MEZNERICS, p. 79, pl. 3, figs 1, 3, 5.</p><p>2009 Echinophoria haueri (Hoernes et Auinger) – LANDAU et al., p. 67, pl. 3, figs 6–8, pl. 9, fig. 8 (cum syn.).</p><p>Material – 35 specimens (SL 26–38); HNHM: INV 2017.666. (Fig. 11), INV 2017.667. (7), 27 specimens in private collections.</p><p>Description – Globose shell, low spire, protoconch of 3.5 smooth, convex whorls, four convex teleoconch whorls, last whorl 90% of the total height, constricted at the base. Ovate aperture, denticulate outer lip thickened by labral varix, short, deep, recurved siphonal canal, expanded parietal callus, slightly excavated columella with folds. Spiral sculpture of numerous narrow spiral cords, nodulose shoulder, axial sculpture of weak riblets (16–20 on the last whorl).</p><p>Remarks – E. haueri is typical of the Paratethys, it clearly differs in morphology from the similar Middle Miocene Mediterranean E. variabilis (Bellardi et Michelotti) in lack of distinct rows of tubercles on the last whorl (LANDAU et al. 2009).</p><p>Distribution – Early Miocene: PMS (Italy), Badenian: CP (Austria, Bulgaria, Hungary, Romania).</p></div>	https://treatment.plazi.org/id/4155C6671A74AC3742A7FEF4FE04FBAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A74AC3542FDFB37FBA5FE3E.text	4155C6671A74AC3542FDFB37FBA5FE3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypraecassis cypraeiformis (Borson 1820)	<div><p>Cypraecassis cypraeiformis (Borson, 1820)</p><p>(Figs 12–13)</p><p>1820–1825 Cassis Cypraeiformis – BORSON, p. 229, pl. 1, fig. 20.</p><p>1966 Cypraecassis crumena cypraeiformis Borson – STRAUSZ, p. 246, pl. 64, figs 10–11.</p><p>2013 Cypraecassis cypraeiformis (Borson) – LANDAU et al., p. 124, pl. 17, fig. 5 (cum syn.).</p><p>Material – 27 specimens (SL 21–35); HNHM: INV 2017.668. (Figs 12–13), INV 2017.669. (2), 24 specimens in private collections.</p><p>Description – Ovate shell, low spire, four teleoconch whorls, slightly convex last whorl more than 80% of the total height, constricted at the base. Narrow, elongate aperture, denticulate outer lip thickened by labral varix, short, narrow siphonal canal, short, recurved fasciole. Smooth shell with axial sculpture of fine growth lines.</p><p>Remarks – Both the genus and the species was thoroughly discussed by LANDAU et al. (2004). C. cypraeiformis is widespred in the European Middle Miocene to Middle Pliocene, but it is a rare element of the mollusc assemblages.</p><p>Distribution – Burdigalian-Langhian: PMS (Italy), Badenian: CP (Austria, Czech Republic, Hungary, Poland, Romania), Serravallian: PMS (Turkey), Tortonian: NEA (Portugal), PMS (Italy). For Pliocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A74AC3542FDFB37FBA5FE3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A76AC3542E5FDA0FE32FA4A.text	4155C6671A76AC3542E5FDA0FE32FA4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Semicassis laevigata (Defrance 1817)	<div><p>Semicassis laevigata (Defrance, 1817)</p><p>(Figs 14–17)</p><p>1817 Cassis laevigata – DEFRANCE, vol. 7, p. 210.</p><p>1966 Phalium (Semicassis) saburon miolaevigatum Sacco – STRAUSZ, p. 244, pl. 64, fig. 4, pl. 72, figs 15–17, pl. 73, figs 1–5.</p><p>2013 Semicassis laevigata (Defrance) – LANDAU et al., p. 125, pl. 17, fig. 6 (cum syn.).</p><p>Material – 351 specimens (SL 12–44); HNHM: M.62.829–831, INV 2017.670. (Figs 14–15), INV 2017. 671. (Figs 16–17), INV 2017.672 (9), 337 specimens in private collections.</p><p>Description – Globose shell, low spire, slightly convex spire whorls, convex, smooth last whorl more than 80% of the total height, constricted at the base. Wide aperture, outer lip thickened by labral varix, smooth with small denticles, short, deep, recurved siphonal canal, plicate columella, short, recurved fasciole. Spiral sculpture of fine grooves on the spiral whorls, two fine grooves on the concave sutural ramp of the last whorl, axial sculpture of growth lines.</p><p>Remarks – S. laevigata is widely distributed in the Badenian localities of Hungary but generally it is a rare element of the mollusc assemblages. Its abundance is noteworthy; it was the most dominant tonnoidean species in the Letkés material.</p><p>Distribution – Karpatian: CP (Austria, Czech Republic, Hungary), Burdigalian-Langhian: NSB (the Netherlands, Germany), PMS (Italy), Badenian: CP (Austria, Bosnia, Bulgaria, Czech Republic, Hungary, Poland, Romania, Serbia, Slovenia, Ukraine), Serravallian: NEA (France), PMS (Turkey), Tortonian: NSB (Denmark), NEA (Portugal, S Spain), PMS (Italy). For Pliocene range see VAN DINGENEN et al. (2016).</p></div>	https://treatment.plazi.org/id/4155C6671A76AC3542E5FDA0FE32FA4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A76AC3B411BFA5AFD26FACB.text	4155C6671A76AC3B411BFA5AFD26FACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Semicassis szilviae Kovács & Vicián 2017	<div><p>Semicassis szilviae n. sp.</p><p>(Figs 18–24)</p><p>Holotype – PAL2017.58.1., Hungarian Natural History Museum, Department of Palaeontology and Geology, (SL 21.5, MD 14.5) (donation of Tamás Hirmetzl) (Figs 21–22).</p><p>Paratypes – 1st: Coll. Z. Vicián (SL 17, MD 11.5) (Figs 23–24), 2nd: Coll. T. Hirmetzl (SL 26, MD 18) (Figs 18–20), 3rd: HNHM PAL 2017.59.1., 4th: PAL 2017.60.1 .</p><p>Type strata – Lower Badenian (Middle Miocene) clayey sand (Sámsonháza Formation).</p><p>Type locality – Letkés, W Börzsöny Mts, N Hungary.</p><p>Derivation of name – In honour of Szilvia Józan (Budapest).</p><p>Material – 14 specimens; HNHM: PAL 2017.58.1., 59.1, 60.1 ., 11 specimens in private collections.</p><p>Diagnosis – Subovate shell, moderately elevated spire, multispiral protoconch, convex last whorl, wide aperture, denticulate outer lip, short, recurved siphonal canal, excavated columella with irregular folds, four narrow spiral cords on the ramp, one strong cord at the shoulder, deep, widely spaced grooves on the last whorl, weak varices and growth lines.</p><p>Description – Subovate shell (largest specimen: SL 27 mm). Moderately elevated spire with concave outline, protoconch of 2.5 convex, smooth whorls, boundary delimited by slightly prosocline scar. Teleoconch of five whorls, subangulate spire whorls with concave sutural ramp. Convex last whorl (85% of the total height) with concave ramp, constricted at the base. Wide aperture, outer lip thickened by labral varix, 17 regular denticles on the inner edge, parietal callus expanded, thin. Short, deep, recurved siphonal canal, broadly excavated columella with irregular folds, short fasciole, abaxially recurved. Spiral sculpture of four narrow cords on the spire ramp and one strong cord at the shoulder, and deep, narrow, widely spaced grooves on the entire last whorl, weakening with ontogeny, axial sculpture of weak varices and growth lines with small tubercles at intersections.</p><p>Remarks – Based on overall morphology the new species is assigned to genus Semicassis . It is characterized by intraspecific variability in sculpture: some specimens have almost smooth last whorl. The species differs from S. laevigata by smaller and narrower shell, by higher, narrower, step-like spire, by less thickened labral varix, and by sculpture with axial ribs on the spire and regular spiral grooves on the last whorl. It also differs from the Early Miocene S. grateloupi (Deshayes) by narrower shell and spire, and by lack of broad spiral cords on the last whorl. The closest form is the small S. subsulcosa pedemontana Sacco from the Early Miocene of the Torino Hills (see FERRERO MORTARA et al. 1984, pl. 1, fig. 5); however, it differs by somewhat broader convex shell, thicker labral varix and stronger cords on the last whorl.</p><p>Distribution – Badenian: CP (Hungary).</p></div>	https://treatment.plazi.org/id/4155C6671A76AC3B411BFA5AFD26FACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A78AC3842CEFA5FFDAFFD04.text	4155C6671A78AC3842CEFA5FFDAFFD04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ranella olearia (Linnaeus 1758)	<div><p>Ranella olearia (Linnaeus, 1758)</p><p>(Figs 25–26)</p><p>1758 Murex olearium – LINNAEUS, p. 748.</p><p>1969 a Bursa (Ranella) gigantea Lam. – CSEPREGHY-MEZNERICS, p. 80, pl. 3, figs 13, 16–17.</p><p>2009 Ranella olearia (Linnaeus) – LANDAU et al., p. 70, pl. 5, figs 3–4 (cum syn.).</p><p>Material – Four specimens (SL 62–74); in private collections .</p><p>Description – Fusiform shell, elevated spire, six convex teleoconch whorls, concave sutural ramp, nodulose shoulder, last whorl constricted at the base. Ovate aperture, denticulate outer lip thickened by labral varix, long, recurved siphonal canal. Spiral sculpture of two primary and numerous secondary cords both on the ramp and below the shoulder on the spire whorls, strong cords and four rows of nodes on the last whorl, axial sculpture of rounded ribs (9 on the last whorl), fine growth lines, two varices per whorl.</p><p>Remarks – The wide palaeogeographical and stratigraphical ranges, as well as the morphological variability of the species was discussed in detail by LANDAU et al. (2004). R. olearia is a rare element of the studied assemblage.</p><p>Distribution – Burdigalian-Tortonian: PMS (Italy), Langhian: NEA (France), Badenian:CP (Austria, Czech Republic, Hungary, Poland, Romania).For Pliocene-Recent range see LANDAU et al. (2004).</p></div>	https://treatment.plazi.org/id/4155C6671A78AC3842CEFA5FFDAFFD04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7BAC394286FCE4FC03FEA2.text	4155C6671A7BAC394286FCE4FC03FEA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Charonia lampas (Linnaeus 1758)	<div><p>Charonia lampas (Linnaeus, 1758)</p><p>(Figs 27–28)</p><p>1758 Murex lampas – LINNAEUS, p. 748.</p><p>2004 Charonia lampas (Linnaeus) – LANDAU et al., p. 53, pl. 4, fig. 1 (cum syn.).</p><p>2013 Charonia lampas (Linnaeus) – LANDAU et al., p. 126, pl. 18, figs 1–3 (cum syn.).</p><p>Material – 28 specimens (SL 20–74); HNHM: INV 2017.673. (Fig. 27), INV 2017.674. (5), 22 specimens in private collections.</p><p>Description – Large, fusiform shell, elevated spire, slightly convex teleoconch whorls, concave sutural ramp, last whorl 72% of the total height, constricted at the base. Ovate aperture, denticulate outer lip thickened by labral varix, short, deep, recurved siphonal canal, excavated, plicate columella, long fasciole. Spiral sculpture of numerous fine cords, two rows of tubercles on spire whorls, two stronger and three weaker rows on the last whorl, axial sculpture of varices.</p><p>Remarks – The species is characterized by wide palaeogeographical and stratigraphical distributions, and extreme morphological variability; the taxonomic revision was accomplished by LANDAU et al. (2004). C. lampas was recorded from Sámsonháza (Cserhát Mts) by NOSZKY (1925) as Triton nodiferum Lamarck, but it has never been illustrated in the Hungarian literature.</p><p>Distribution – Aquitanian-Burdigalian: NEA (France), Burdigalian: PMS (Italy), Badenian: CP (Austria, Hungary, Poland), Serravallian: PMS (Turkey), Late Miocene: Caribbean (Dominican Republic), Pacific (Japan, New Zealand). For Eocene and Pliocene-Recent range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A7BAC394286FCE4FC03FEA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7DAC3E4295FEA9FC31FB83.text	4155C6671A7DAC3E4295FEA9FC31FB83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monoplex corrugatus (Lamarck 1816)	<div><p>Monoplex corrugatus (Lamarck, 1816)</p><p>(Figs 29–30)</p><p>1816 Triton corrugatum – LAMARCK, p. 5.</p><p>1966 Cymatium (Lampusia) affine Deshayes – STRAUSZ, p. 247, pl. 29, figs 8–11, pl. 30, figs 1–5.</p><p>2013 Monoplex corrugatus (Lamarck) – LANDAU et al., p. 127, pl. 18, figs 5–7 (cum syn.).</p><p>Material – 123 specimens (SL 18–74); HNHM: M.62.836., INV 2017.675. (9), 113 specimens in private collections.</p><p>Description – Narrow, fusiform shell, elevated spire, six subangulate teleoconch whorls, broad, concave sutural ramp, last whorl 70% of the total height. Ovate aperture, denticulate outer lip thickened by labral varix, long, narrow, straight siphonal canal, concave columella with folds, long fasciole. Spiral sculpture of three primary and numerous secondary cords on the spire, six broad primaries on the last whorl, axial sculpture of seven prosocline rounded ribs, one varix per whorl.</p><p>Remarks – The species is characterized by large morphological variability, so Triton affine Deshayes – that was frequently recorded from the Mediterranean and the Paratethys – is considered as a junior synonym in the recent literature. M. corrugatus is abundant in the mollusc assemblage of Letkés.</p><p>Distribution – Burdigalian: PMS (Italy), late Burdigalian-Langhian: NSB (the Netherlands), Badenian: CP (Austria, Bulgaria, Czech Republic, Hungary, Poland, Romania, Ukraine), Serravallian: PMS (Turkey), Tortonian: NEA (Portugal), PMS (Italy). For Pliocene-Recent range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A7DAC3E4295FEA9FC31FB83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7DAC3F42B0FB00FCBDFDF1.text	4155C6671A7DAC3F42B0FB00FCBDFDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monoplex subcorrugatus (d'Orbigny 1852)	<div><p>Monoplex subcorrugatus (d’Orbigny, 1852)</p><p>(Figs 31–32)</p><p>1852 Triton subcorrugatum – D’ORBIGNY, p. 77.</p><p>1969 a Cymatium (Ranularia) multicostatum Cossmann et Peyrot – CSEPREGHY-MEZNERICS, p. 80, pl. 3, figs 9–12.</p><p>2001 Cymatium subcorrugatum (d’Orbigny) – LOZOUET et al., p. 46, pl. 19, fig. 1 (cum syn.).</p><p>2009 Monoplex subcorrugatus (d’Orbigny) – LANDAU et al., p. 71 (pars).</p><p>Material – Three specimens (SL 20–26); HNHM: INV 2017.676. (2), one specimen in private collection.</p><p>Description – Broad shell, elevated spire, four subangulate teleoconch whorls, slightly concave, sloping sutural ramp, convex last whorl. Ovate aperture, denticulate outer lip thickened by labral varix, weakly plicate columella. Spiral sculpture of fine cords on the ramp, two well-developed primary cords on the spire whorls and 6 on the last whorl (adapical five divided by median grooves), one secondary cord in interspaces, axial sculpture of broad, rounded ribs (7–9 on the last whorl) and growth lines.</p><p>Remarks – M. subcorrugatus differs in morphology from M. corrugatus in smaller size, broader shell and lack of varices on the early teleoconch whorls, it is probably the ancestor of the latter. It also differs from M. heptagonus in broader shell, conical spire and less developed, broader primary cords.</p><p>Distribution – Aquitanian-Langhian: NEA (France), late Burdigalian-Langhian: NSB (the Netherlands), Badenian: CP (Hungary).</p></div>	https://treatment.plazi.org/id/4155C6671A7DAC3F42B0FB00FCBDFDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7CAC3F42E4FDD1FCF3FA4A.text	4155C6671A7CAC3F42E4FDD1FCF3FA4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monoplex heptagonus (Brocchi 1814)	<div><p>Monoplex heptagonus (Brocchi, 1814)</p><p>(Figs 33–34)</p><p>1814 Murex heptagonus – BROCCHI, p. 404, pl. 9, fig. 2.</p><p>1966 Cymatium (Ranularia) heptagonum vindobonicum Cossmann &amp; Peyrot – STRAUSZ, p. 248, pl. 29, fig. 7.</p><p>2013 Monoplex heptagonus (Brocchi) – LANDAU et al., p. 128, pl. 18, figs 8–10 (cum syn.).</p><p>Material – 106 specimens (SL 10–25); HNHM: M.62.835., INV 2017.677. (Figs 33–34), INV 2017.678. (7), 97 specimens in private collections.</p><p>Description – Small shell, moderately elevated, step-like spire, four, convex, shouldered teleoconch whorls, slightly convex, subtrigonal last whorl with sloping sutural ramp. Ovate aperture, denticulate outer lip thickened by labral varix, long, narrow, straight siphonal canal, plicate columella, thickened parietal callus, long fasciole. Spiral sculpture of three well-developed primary cords on the spire whorls and 5–6 on the last whorl, one secondary cord in interspaces, axial sculpture of broad, rounded ribs (7–8 on the last whorl), growth lines and one varix on the last whorl, tubercles at intersections.</p><p>Remarks – The morphology of the species is very similar to that of M. corrugatus (see BAŁUK 1995), but it differs in scalate spire, more strongly shouldered whorls and more raised spiral cords. The M. heptagonus material of Letkés shows moderate intraspecific variability in development of sculpture. The species was abundant in the mollusc assemblage.</p><p>Distribution – Burdigalian: PMS (Italy), Badenian: CP (Austria, Bulgaria, Hungary, Poland, Romania), Serravallian: PMS (Turkey), Tortonian: PMS (Italy). For Pliocene range see VAN DINGENEN et al. (2016).</p></div>	https://treatment.plazi.org/id/4155C6671A7CAC3F42E4FDD1FCF3FA4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7EAC3D42B0FE66FC79FBA7.text	4155C6671A7EAC3D42B0FE66FC79FBA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Turritriton grundensis (Hoernes et Auinger 1884)	<div><p>Turritriton grundensis (Hoernes et Auinger, 1884)</p><p>(Figs 37–38)</p><p>1884 Triton (d. Simpulum) Grundense – HOERNES &amp; AUINGER, p. 177, pl. 21, fig. 16.</p><p>2009 Turritriton grundensis (Hoernes &amp; Auinger) – LANDAU et al., p. 72, pl. 6, fig. 5 (cum syn.).</p><p>Material – 13 specimens (SL 17–22); HNHM: INV 2017.679. (Figs 37–38), 12 specimens in private collections.</p><p>Description – Gracile shell, high, step-like spire, spire whorls with two strong spiral cords at mid-whorl, sloping ramp, strongly shouldered last whorl. Ovate aperture, denticulate outer lip thickened by labral varix, long, narrow, straight siphonal canal, columella with two folds, thickened parietal callus. Spiral sculpture of well-developed primary and secondary cords, axial sculpture of broad, rounded ribs, growth lines and one varix on the last whorl.</p><p>Remarks – The species is typical of the Central Paratethys. It differs in morphology from the similar M. heptagonus in slender shell, sloping sutural ramp and finely reticulate surface. T. grundensis is a new record in Hungary.</p><p>Distribution – Badenian: CP (Austria, Hungary, Poland).</p></div>	https://treatment.plazi.org/id/4155C6671A7EAC3D42B0FE66FC79FBA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A7EAC22413DFB38FE1BFDF1.text	4155C6671A7EAC22413DFB38FE1BFDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sassia apenninica (Sassi 1827)	<div><p>Sassia apenninica (Sassi, 1827)</p><p>(Figs 39–40, 45)</p><p>1827 Triton apenninicum – SASSI, p. 480.</p><p>1956 Charonia (Sassia) apenninica Sassi – CSEPREGHY-MEZNERICS, p. 397, pl. 4, figs 13–14.</p><p>2013 Sassia apenninica (Sassi) – LANDAU et al., p. 130, pl. 19, fig. 4.</p><p>Material – Ten specimens (SL 13–29); HNHM: M.62.831–834., INV 2017.680. (3), three specimens in private collections .</p><p>Description – Fusiform shell, elevated spire, carinate teleoconch whorls, steep sutural ramp, rounded last whorl. Ovate aperture, denticulate outer lip thickened by labral varix, long, narrow, recurved siphonal canal, plicate columella. Spiral sculpture of strong primary and narrow secondary cords, axial sculpture of riblets, one varix per whorl.</p><p>Remarks – The taxonomy and the morphological variability of S. apenninica were discussed in detail by LANDAU et al. (2004). As the species is typical of the deeper-water deposits, it is rare in the shallow-water marly sand dominated Letkés section.</p><p>Distribution – Aquitanian-Langhian: NEA (France), Burdigalian: PMS (Italy), Badenian: CP (Austria, Bulgaria, Czech Republic, Hungary, Poland, Romania), Serravallian: PMS (Turkey), Tortonian: PMS (Italy). For Pliocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A7EAC22413DFB38FE1BFDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A61AC2242F3FDDEFB99FAF0.text	4155C6671A61AC2242F3FDDEFB99FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sassia turrita (Eichwald 1830)	<div><p>Sassia turrita (Eichwald, 1830)</p><p>(Figs 41–42)</p><p>1830 Tritonium turritum – EICHWALD, p. 225.</p><p>1966 Charonia (Sassia) tarbelliana Grateloup – STRAUSZ, p. 249, pl. 30, figs 7–8.</p><p>2013 Sassia turrita (Eichwald) – LANDAU et al., p. 130, pl. 19, figs 5–8, pl. 62, fig. 9 (cum syn.).</p><p>Material – 43 specimens (SL 15–48); HNHM: INV 2017.681. (Figs 41–42), INV 2017.682. (5), 37 specimens in private collections.</p><p>Description – Fusiform shell, elevated spire, seven convex teleoconch whorls, shouldered early spire whorls with steep sutural ramp, rounded last whorl, constricted at the base. Ovate aperture, denticulate outer lip thickened by labral varix, long, narrow siphonal canal, long fasciole. Spiral sculpture of narrow cords on the spire, fine threads on the last whorl, axial sculpture of rounded ribs and one or two broad, rounded varices per whorl.</p><p>Remarks – The nomenclature problem of the species, the priority of Eichwald’s taxon and the replacement of Triton tarbellianum Grateloup was analyzed by LANDAU et al. (2009).</p><p>Distribution – Chattian: NSB (Germany), Egerian: CP (Hungary), late Burdigalian-Langhian: NSB (the Netherlands, Belgium, Germany), Langhian-Serravallian: NEA (France), Badenian: CP (Austria, Bulgaria, Czech Republic, Hungary, Poland, Romania, Serbia, Slovakia, Ukraine), Serravallian: PMS (Turkey).</p></div>	https://treatment.plazi.org/id/4155C6671A61AC2242F3FDDEFB99FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A61AC204295FAECFD56FDD5.text	4155C6671A61AC204295FAECFD56FDD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymatiella tritonea (Grateloup 1847)	<div><p>Cymatiella tritonea (Grateloup, 1847)</p><p>(Figs 43–44)</p><p>1847 Murex tritoneum – GRATELOUP, pl. 29, fig. 23.</p><p>1966 Charonia (Sassia) parvula Michelotti – STRAUSZ, p. 250, pl. 30, fig. 9.</p><p>2009 Cymatiella tritonea (Grateloup) – LANDAU et al., p. 74, pl. 7, figs 4–9, pl. 10, fig. 3 (cum syn.).</p><p>Material – 21 specimens (SL 11–13); HNHM: INV 2017.683. (5), 16 specimens in private collections.</p><p>Description – Small, fusiform shell, elevated spire, teleoconch of five convex whorls, rounded last whorl, constricted at the base. Small, ovate aperture, denticulate outer lip thickened by labral varix, short siphonal canal and fasciole, columella with three folds. Spiral sculpture of cords, axial sculpture of rounded ribs with tubercles at the intersections, one or two varices per whorl.</p><p>Remarks – Early Miocene specimens from France and the Paratethys were revised by LANDAU et al. (2004, 2009), and Triton parvulum Michelotti, which had been frequently recorded from the Paratethys was regarded as a synonym of C. tritonea .</p><p>Distribution – Chattian-Burdigalian: NEA (France), Burdigalian: PMS (Italy), Karpatian: CP (Austria), Badenian: CP (Austria, Bulgaria, Hungary, Poland, Romania, Serbia), Tortonian: PMS (Italy).</p></div>	https://treatment.plazi.org/id/4155C6671A61AC204295FAECFD56FDD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A63AC204281FD55FE66FA23.text	4155C6671A63AC204281FD55FE66FA23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bursa scrobilator (Linnaeus 1758)	<div><p>Bursa scrobilator (Linnaeus, 1758)</p><p>(Figs 46–47)</p><p>1758 Murex scrobilator – LINNAEUS, p. 749.</p><p>1969 a Bursa (Ranella) nodosa subnodosa Sacco – CSEPREGHY-MEZNERICS, p. 81, pl. 3, figs 6, 8. 2009 Bursa scrobilator (Linnaeus) – LANDAU et al., p. 75, pl. 7, figs 10–11 (cum syn.).</p><p>Material – Three specimens (SL 18–22); HNHM: INV 2017.684. (Figs 46– 47), two specimens in private collections.</p><p>Description – Fusiform shell, elevated spire, protoconch of three smooth, convex whorls, four subangulate teleoconch whorls, convex last whorl 70% of the total height, constricted at the base. Ovate aperture, convex, denticulate outer lip thickened by labral varix, short anal canal, short, slightly recurved siphonal canal, slightly concave, plicate columella. Spiral sculpture of narrow cords, three fine nodular rows on the sutural ramp, nodulose shoulder, two fine primary nodular rows below shoulder on the last whorl, axial sculpture of two prominent varices per whorl.</p><p>Remarks – The high diversity of genus Bursa in the studied assemblage is remarkable: all three species known from the Paratethys occur at Letkés. The specimen figured here represents the “ nodosa ” form which was typical of the Miocene Paratethys.</p><p>Distribution – Badenian: CP (Austria, Hungary, Poland), Tortonian: PMS (Italy), NEA (France). For Pliocene-Pleistocene range of the “ nodosa ” form see LANDAU et al. (2004).</p></div>	https://treatment.plazi.org/id/4155C6671A63AC204281FD55FE66FA23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A63AC2142F2FA60FE6EFD04.text	4155C6671A63AC2142F2FA60FE6EFD04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bursa ranelloides (Reeve 1844)	<div><p>Bursa ranelloides (Reeve, 1844)</p><p>(Figs 48–49)</p><p>1844 Triton Ranelloides – REEVE, pl. 3, fig. 10.</p><p>2009 Bursa ranelloides (Reeve) – LANDAU et al., p. 75, pl. 8, figs 1–6, pl. 10, fig. 4 (cum syn.).</p><p>Material – Nine specimens (SL 18–23); in private collections.</p><p>Description – Fusiform shell, elevated spire, protoconch of three smooth, convex whorls, four subangulate teleoconch whorls, last whorl 74% of the total height, constricted at the base. Ovate aperture, denticulate, angulate outer lip thickened by labral varix, short anal canal, short, slightly recurved siphonal canal, straight, plicate columella. Spiral sculpture of narrow cords, nodulose shoulder, one row of small tubercles above abapical suture on the spire, three rows of tubercles on the last whorl, axial sculpture of two varices per whorl.</p><p>Remarks – The species differs in morphology from B. scrobilator by subangulate whorls with stronger sculpture, and from B. corrugata by narrower shell with sharper tubercles. B. ranelloides is a new record in Hungary.</p><p>Distribution – Burdigalian: PMS (Italy), Badenian: CP (Austria, Hungary, Romania), Middle Miocene: WA (Trinidad). For Pliocene-Recent range see LANDAU et al. (2009).</p></div>	https://treatment.plazi.org/id/4155C6671A63AC2142F2FA60FE6EFD04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A62AC21413BFC8DFCC9F9FC.text	4155C6671A62AC21413BFC8DFCC9F9FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bursa corrugata (Perry 1811)	<div><p>Bursa corrugata (Perry, 1811)</p><p>(Figs 50–54)</p><p>1811 Biplex corrugata – PERRY, pl. 5, fig. 1.</p><p>2009 Bursa corrugata (Perry) – LANDAU et al., p. 77, pl. 8, figs 7–10, pl. 10, fig. 5 (cum syn.).</p><p>Material – 87 specimens (SL 10.5–28); HNHM: INV 2017.685. (Figs 52– 53), INV 2017.686. (9), 77 specimens in private collections.</p><p>Description – Fusiform shell, elevated spire, protoconch of three smooth, convex whorls, four teleoconch whorls, angulate last whorl 78% of the total height, constricted at the base. Ovate aperture, denticulate outer lip thickened by labral varix, short, deep anal canal, short, recurved siphonal canal, straight, plicate columella, thickened columellar callus, short fasciole. Papillate surface, spiral sculpture of narrow cords, two rows of tubercles on the spire, four rows on the last whorl (strong tubercles of adapical row – 9 on the last whorl – form the shoulder), axial sculpture of two varices per whorl.</p><p>Remarks – Based on the revision of fossil and recent bursids, Bursa papillosa (Pusch) frequently recorded from the Paratethys was regarded as a junior synonym of B. corrugata by LANDAU et al. (2009). The species shows slight variability in the development of nodes. It is a new record in Hungary.</p><p>Distribution – Burdigalian: NEA (France), PMS (Italy), Badenian: CP (Austria, Bulgaria, Hungary, Poland, Romania), Middle Miocene: WA (Trinidad). For Pliocene-Recent range see LANDAU et al. (2009).</p></div>	https://treatment.plazi.org/id/4155C6671A62AC21413BFC8DFCC9F9FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A65AC2642F6FED6FB90FC13.text	4155C6671A65AC2642F6FED6FB90FC13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspa marginata (Gmelin 1791)	<div><p>Aspa marginata (Gmelin, 1791)</p><p>(Figs 55–56)</p><p>1791 Buccinum marginatum – GMELIN, p. 3486.</p><p>1966 Bursa (Aspa) marginata depressa Grateloup – STRAUSZ, p. 251, pl. 29, figs 5–6, pl. 63, figs 14–18.</p><p>2013 Aspa marginata (Gmelin) – LANDAU et al., p. 131, pl. 19, fig. 9 (cum syn.).</p><p>Material – 160 specimens (SL 14–42); HNHM: INV 2017.687. (Fig. 55), INV 2017.688. (Fig. 56), INV 2017.689. (15), 143 specimens in private collections.</p><p>Description – Ovate, dorsoventrally compressed shell, low spire, four teleoconch whorls, convex last whorl 85% of the total height. Ovate, denticulate aperture, straight, long anal canal, short, recurved siphonal canal, concave columella, thickened parietal callus, short, recurved fasciole. Spiral sculpture of fine cords, tubercles at the shoulder, axial sculpture of two varices per whorl.</p><p>Remarks – The species is widely distributed in the Badenian localities of Hungary, and it was abundant in the mollusc assemblage of Letkés.</p><p>Distribution – Burdigalian-Langhian: PMS (Italy), Ottnangian: CP (Germany), Karpatian: CP (Austria), Badenian: CP (Austria, Bosnia, Bulgaria, Hungary, Poland, Romania, Ukraine), Serravallian: PMS (Turkey), Tortonian: NEA (Portugal, France), PMS (Italy). For Pliocene-Pleistocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A65AC2642F6FED6FB90FC13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A65AC24428DFBE8FB90FE3E.text	4155C6671A65AC24428DFBE8FB90FE3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Distorsio cancellina (Lamarck 1803)	<div><p>Distorsio cancellina (Lamarck, 1803)</p><p>(Figs 57–58)</p><p>1803 Murex cancellinus – LAMARCK, p. 225.</p><p>2004 Distorsio cancellina (Lamarck) – LANDAU et al., p. 71, pl. 6, figs 1–3 (cum syn.).</p><p>2013 Distorsio cancellina (Lamarck) – LANDAU et al., p. 132, pl. 19, figs 10–12 (cum syn.).</p><p>Material – 103 specimens (SL 14–43); HNHM: INV 2017.690. (Figs 57– 58), INV 2017.691. (4), 98 specimens in private collections.</p><p>Description – Fusiform shell, moderately elevated, irregularly coiled spire, six teleoconch whorls, last whorl expanded to left of the aperture. Denticulate aperture, short, recurved siphonal canal, short fasciole. Spiral sculpture of narrow cords, axial sculpture of rounded ribs and low varices with small tubercles at the intersections.</p><p>Remarks – The revision of the taxon was accomplished by LANDAU et al. (2004). In Hungary the species was recorded from the Börzsöny and Cserhát Mts as Triton tortuosum Borson (NOSZKY 1925), but it has never been illustrated in the Hungarian literature. D. cancellina is rare at the Paratethys localities; its abundance in the mollusc assemblage of Letkés is noteworthy.</p><p>Distribution – Burdigalian-Tortonian: PMS (Italy), late Burdigalian-Langhian: NSB (the Netherlands, Germany), Aquitanian-Serravallian: NEA (France), Badenian:CP (Austria, Bulgaria, Hungary, Romania), Langhian-Serravallian:PMS (Turkey), Tortonian: NEA (Portugal). For Pliocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A65AC24428DFBE8FB90FE3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A67AC244241FDA0FCAEFB9A.text	4155C6671A67AC244241FDA0FCAEFB9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Personopsis grasi (Bellardi in d’Ancona 1872)	<div><p>Personopsis grasi (Bellardi in d’Ancona, 1872)</p><p>(Fig. 59)</p><p>1872 Triton Grasi Bellardi – D’ANCONA, p. 70.</p><p>2009 Personopsis grasi (d’Ancona) – LANDAU et al., p. 79, pl. 9, figs 5–7.</p><p>Material – One fragmentary specimen (SL 20); in private collection.</p><p>Description – Fusiform shell, elevated spire, six weakly convex teleoconch whorls, regularly convex last whorl. Subtrigonal aperture, denticulate outer lip thickened by labral varix, strongly excavated columella, thick parietal callus. Spiral sculpture of three narrow cords on the spiral whorls and 11 on the last whorl, fine threads, axial sculpture of about 16 rounded ribs, one weak varix per whorl.</p><p>Remarks – The Paratethys specimens differ in morphology from their Pliocene descendants in Italy by smaller and narrower shell with finer ornamentation. The species is very rare in the Paratethys. P.grasi is a new record in Hungary.</p><p>Distribution – Badenian: CP (Austria, Hungary, Romania), Tortonian: PMS (Italy). For Pliocene-Recent range see LANDAU et al. (2004).</p></div>	https://treatment.plazi.org/id/4155C6671A67AC244241FDA0FCAEFB9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A67AC25428BFA8EFDC1FDF1.text	4155C6671A67AC25428BFA8EFDC1FDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ficus burdigalensis (Sowerby 1824)	<div><p>Ficus burdigalensis (Sowerby, 1824)</p><p>(Figs 60–61)</p><p>1824 Pyrula Burdigalensis – SOWERBY, fig. 2.</p><p>1966 Pirula (Fulguroficus) burdigalensis Sowerby – STRAUSZ, p. 256, fig. 121.</p><p>2007 Ficus burdigalensis (Sowerby) – PFISTER &amp; WEGMÜLLER, p. 153, pl. 1, figs 10–15 (cum syn.).</p><p>Material – Six specimens (SL 32–38); in private collections.</p><p>Description – Ficiform shell, moderately low spire, eroded protoconch, three shouldered teleoconch whorls, last whorl 92% of the total height. Long aperture, smooth outer lip, long siphonal canal. Spiral sculpture of numerous strong, narrow cords, two rows of spines on the spire, four rows of spines on the last whorl, axial sculpture of fine growth lines.</p><p>Remarks – F. burdigalensis differs in morphology from its Badenian congeners in characteristic rows of spines on the teleoconch whorls. The species is typical of the Early Miocene, its occurrence in the early Badenian is noteworthy.</p><p>Distribution – Aquitanian: NEA (France), Burdigalian: NEA (Portugal), Paratethys (Switzerland), Eggenburgian-Karpatian: CP (Austria, Hungary), Badenian: CP (Hungary).</p></div>	https://treatment.plazi.org/id/4155C6671A67AC25428BFA8EFDC1FDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A66AC2542C9FDDEFD0FFA97.text	4155C6671A66AC2542C9FDDEFD0FFA97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ficus condita (Brongniart 1823)	<div><p>Ficus condita (Brongniart, 1823)</p><p>(Figs 62–64)</p><p>1823 Pyrula condita – BRONGNIART, p. 75, pl. 6, fig. 4.</p><p>1966 Pirula condita Brongniart – STRAUSZ, p. 254, pl. 57, figs 4–6.</p><p>2013 Ficus condita (Brongniart) – LANDAU et al., p. 132, pl. 19, fig. 13, pl. 62, fig. 10 (cum syn.). 2016 Ficus condita (Brongniart) – STEIN et al., p. 41, pl. 15, fig. 7.</p><p>Material – 32 specimens (SL 21–44); HNHM: INV 2017.692. (Fig. 63), INV 2017.693. (9), 22 specimens in private collections.</p><p>Description – Ficiform shell, low spire, protoconch of two convex, smooth whorls, convex teleoconch whorls, last whorl 94–97% of total height, constricted at base. Long, wide aperture, smooth outer lip, long, slightly recurved siphonal canal. Cancellate sculpture with numerous narrow and strong primary spiral cords and fine secondaries in the interspaces, and fine axial ribs.</p><p>Remarks – The morphology of the species differs from that of F. cingulata in higher spire and finer sculpture, and from F. geometra in stronger primary cords. It is widely distributed in the Early-Middle Miocene localities of Hungary, and it was the most abundant Ficus species in the mollusc assemblage of Letkés.</p><p>Distribution – Aquitanian-Serravallian: NEA (France), Burdigalian: PMS (Italy), Eggenburgian: CP (Austria, Hungary, Slovakia, Romania), Karpatian: CP (Hungary), late Burdigalian-Langhian: NSB (the Netherlands, Belgium, Germany, Denmark), Badenian: CP (Austria, Bulgaria, Hungary, Poland, Romania, Slovakia, Ukraine), Serravallian: PMS (Turkey), Tortonian: PMS (Italy). For Oligocene range see LANDAU et al. (2013).</p></div>	https://treatment.plazi.org/id/4155C6671A66AC2542C9FDDEFD0FFA97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A66AC2B413FFA3CFD15FD6F.text	4155C6671A66AC2B413FFA3CFD15FD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ficus geometra (Borson 1825)	<div><p>Ficus geometra (Borson, 1825)</p><p>(Figs 65–66)</p><p>1820–1825 Pyrula geometra – BORSON, p. 311.</p><p>1966 Pirula geometra Borson – STRAUSZ, p. 255, pl. 57, figs 1–3.</p><p>1973 Pirula geometra Borson – BORZA, pl. 2, fig. 9.</p><p>2004 Ficus geometra (Borson) – LANDAU et al., p. 79, pl. 8, fig. 3 (cum syn.).</p><p>2007 Ficus geometra (Borson) – PFISTER &amp; WEGMÜLLER, p. 155, pl. 1, figs 21–23, pl. 2, figs 1–4.</p><p>Material – Seven specimens (SL 26–31); HNHM: INV.2017.695 ., six specimens in private collections.</p><p>Description – Ficiform shell, convex teleoconch whorls, last whorl constricted at the base. Long, wide aperture, smooth outer lip, straight columella, long, slightly recurved siphonal canal. Reticulate sculpture with narrow primary spiral cords (more than 30 on last whorl) and one secondary cord in each interspace, and axial ribs as strong as the primaries.</p><p>Remarks – The species differs in morphology from F. condita and F. cingulata by regular reticulate sculpture.</p><p>Distribution – Middle Miocene: PMS (Italy), NSB (Denmark), Burdigalian: NSB (Germany), Paratethys (Switzerland), Eggenburgian: CP (Slovakia), Badenian: CP (Austria, Hungary, Poland, Romania), Tortonian: PMS (Italy). For Pliocene range see VAN DINGENEN et al. (2016).</p></div>	https://treatment.plazi.org/id/4155C6671A66AC2B413FFA3CFD15FD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
4155C6671A68AC2B42FAFD24FD6FFADD.text	4155C6671A68AC2B42FAFD24FD6FFADD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ficus cingulata (Bronn in Hörnes 1856)	<div><p>Ficus cingulata (Bronn in Hörnes, 1856)</p><p>(Figs 67–68)</p><p>1856 Pyrula cingulata Bronn – HÖRNES, p. 676, pl. 28, figs 1–3.</p><p>1954 Pirula (Ficus) condita cingulata Bronn – STRAUSZ, p. 103, pl. 7, fig. 152.</p><p>1969b Pirula cingulata Bronn – CSEPREGHY-MEZNERICS, p. 24, pl. 8, figs 5, 10.</p><p>2002 Ficus (Ficus) cingulata Bronn – HARZHAUSER, p. 90, pl. 5, fig. 16 (cum syn.).</p><p>Material – 12 specimens (SL 30–64); HNHM: INV 2017.694. (9), three specimens in private collections.</p><p>Description – Large, ficiform shell, low spire, protoconch of two convex, smooth whorls, convex teleoconch whorls, last whorl 98% of the total height, constricted at the base. Long, wide aperture, smooth outer lip, long, slightly recurved siphonal canal. Spiral sculpture of two strong primary and numerous fine secondary cords on sutural ramp of the last whorl, broad, flat primary cords (10–14 on the last whorl) with two secondaries in the interspaces, axial sculpture of fine axial ribs.</p><p>Remarks – The species is typical of the Paratethys. It differs in morphology from F. condita by larger size, lower spire and broad and flat spiral primary cords.</p><p>Distribution – Karpatian-Badenian: CP (Austria, Hungary), Badenian: CP (Bulgaria, Hungary, Poland, Romania).</p></div>	https://treatment.plazi.org/id/4155C6671A68AC2B42FAFD24FD6FFADD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kovács, Zoltán;Vicián, Zoltán	Kovács, Zoltán, Vicián, Zoltán (2017): Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkés (Hungary). Fragmenta Palaeontologica Hungarica 34: 75-104, DOI: 10.17111/FragmPalHung.2017.34.75, URL: https://doi.org/10.17111/fragmpalhung.2017.34.75
