identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BED1863A60AD56B79439BE3B05604323.text	BED1863A60AD56B79439BE3B05604323.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alluropus calcaratus (Pocock 1891)	<div><p>Alluropus calcaratus (Pocock, 1891)</p><p>Figs 1, 2, 3</p><p>Rhysida calcarata Pocock, 1891: 61; Kraepelin 1903: 146; Attems 1930: 191; Attems 1938: 338; Schileyko 2007: 83, fig. 6; Siriwut et al. 2018: 1029, figs 14–23.</p><p>Alluropus demangei Silvestri, 1912; Attems 1930: 198; Schileyko 2007: 92; Tran et al. 2013: 221;</p><p>Rhysida marginata Schileyko, 2007: 92; 148; Tran et al. 2013: 227;</p><p>Alluropus calcaratus Siriwut et al. 2018: 1029; Schileyko et al. 2020: 50.</p><p>Examined materials.</p><p>Vietnam • 3 specimens; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.44694&amp;materialsCitation.latitude=10.62583" title="Search Plazi for locations around (long 107.44694/lat 10.62583)">Ba Ria</a> – Vung Tau Province, residential area; 10.62583°N, 107.44694°E; 47 m a. s. l.; 25 Oct. 2016; A. D. Nguyen leg.; IEBR.VT.2016.1, IEBR.VT.2016.2, IEBR.VT.2016.3 . • 1 specimen; Dong Thap Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.52186&amp;materialsCitation.latitude=10.71746" title="Search Plazi for locations around (long 105.52186/lat 10.71746)">Tram Chim NP</a>, Melaleuca forests; 10.71746°N, 105.52186°E; 47 m a. s. l.; 25 Oct. 2016; D. H. Dang leg.; SVR.TrC.010 .</p><p>Diagnosis.</p><p>Body with 21 LBS, and LBS 7 th with spiracles (Fig. 2 E). Antennae (Fig. 2 A, B) composed of 20–21 articles, of which three basal ones glabrous dorsally. Cephalic plate smooth, lateral and posterior marginations absent; the posterior edge straight, and covered by tergite 1 (Fig. 2 A). Forcipular trochanteroprefermur with median tooth; tooth-plates with 4 + 4 teeth clearly, sharply (Fig. 2 B). Tergite 1 wider than the cephalic plate and twice as long as tergite 2 (Fig. 2 A); tergites 3–19 (20) with complete paramedian sutures, and tergites 12 (13) – 20 with lateral marginations incomplete or nearly complete (Figs 2 E, 3 A, B). Sternites 3–20 with complete paramedian sutures (Fig. 2 C, D). Coxopleuron with 1 apical, 2 subapical, and 1 lateral spine, without dorsal spine (Fig. 3 D). Leg 1 with prefemoral and femoral spurs (Fig. 2 B). Ultimate legs (Fig. 3 C, D) long and slender, with ratio between the length and width of prefermur 4.5: 1; prefemoral spines on ultimate leg: 4 VL, 2–3 VM, 3–4 M, and 1–2 DM spines.</p><p>Remarks.</p><p>Compared to Kraepelin (1903), Attems (1930), and Siriwut et al. (2018), our specimens have slight differences, e. g., shorter body length (maximum 32 mm vs. 57 mm in Kraepelin (1903) and 52 mm in Siriwut et al. (2018)). This is probably due to the low number of specimens collected and probably not an actual difference. The lateral marginations start from tergite 9 (or 11) in descriptions of Attems (1930) and Kraepelin (1903), but these start from tergite 12 (or 13) in our specimens and the description of Siriwut et al. (2018).</p><p>Our specimens do not agree with the description of Siriwut et al. (2018) in the ratio between tarsus 1 and tarsus 2 (1.6: 1 vs. 0.3: 1) but more or less similar to Schileyko (2007) (terminal tarsus 1 more than twice longer than tarsus 2).</p><p>Distribution.</p><p>This species has only been recorded in Southeast Asia, including Thailand, Laos, Cambodia, and Vietnam (Tran et al. 2013; Siriwut et al. 2018). In Vietnam it was recorded from Ha Nam Province (Phu Ly) and Ca Mau Province (Attems 1930; Tran et al. 2013), and Ba Ria – Vung Tau Province and Dong Thap Province in this study. The distribution of A. calcaratus was further updated (Fig. 1).</p></div>	https://treatment.plazi.org/id/BED1863A60AD56B79439BE3B05604323	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
1406D32EE8DC5912AC88C72B65C99D02.text	1406D32EE8DC5912AC88C72B65C99D02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alluropus Silvestri 1912	<div><p>Genus Alluropus Silvestri, 1912</p><p>Type species.</p><p>Alluropus demangei Silvestri, 1911, by monotypy.</p><p>Diagnosis.</p><p>According to Schileyko et al. (2020: p. 50), the genus can be recognized by the following characters: median tooth of labrum well developed; forcipular tooth-plates present; trochantero-prefemur with well-developed process; tergites never with longitudinal keels; sternites with incomplete paramedian sutures (in some species much shortened, sometimes also with median and lateral depressions of various sizes / shapes; LBS 7 with spiracles; legs with tarsal spur (s), legs 1 with prefemoral spur; coxopleural process ranging from short to very long and much enlarged with spines (at least with apical ones); prefemur of the ultimate leg with spines; ultimate pretarsus well-developed, with accessory spines.</p><p>Distribution.</p><p>Vietnam, Cambodia, Laos, and Thailand.</p></div>	https://treatment.plazi.org/id/1406D32EE8DC5912AC88C72B65C99D02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
C89ADC33546457969E237C952A505C35.text	C89ADC33546457969E237C952A505C35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhysida immarginata (Porat 1876)	<div><p>Rhysida immarginata (Porat, 1876)</p><p>Figs 1, 6, 7</p><p>Rhysida immarginata Pocock, 1891 - Kraepelin 1903: 143; Lewis 2001: 45; Siriwut et al. 2018: 1022, figs 9–11.</p><p>Rhysida nuda immarginata Attems, 1930: 190.</p><p>Rhysida nuda Attems, 1930: 189, figs 236–237; Attems 1938: 337; Tran et al. 2013: 227.</p><p>Examined materials.</p><p>Vietnam • 1 specimen; Ha Giang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.85867&amp;materialsCitation.latitude=22.78121" title="Search Plazi for locations around (long 104.85867/lat 22.78121)">Tay Con Linh NR</a>, natural forest; 22.78121°N, 104.85867°E; 1,113 m a. s. l.; 2019; col. A. D. Nguyen leg; HG.TCL.02 . • 3 specimens; Dien Bien Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.39833&amp;materialsCitation.latitude=21.21278" title="Search Plazi for locations around (long 103.39833/lat 21.21278)">Muong Nhe Natural Reserve</a>; residential area, 21.21278°N, 103.39833°E; 610 m a. s. l.; May 2018; S. X. Le &amp; H. D. Nguyen leg.; DB 019, DB 013, DB 018 ; • 2 specimens; same location, residential area; 22.30056°N, 102.39833°E; 558 m a. s. l.; Nov. 2017; S. X. Le leg.; MN 018.1, MN 106 . • 1 specimen; Son La Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.66972&amp;materialsCitation.latitude=21.35972" title="Search Plazi for locations around (long 104.66972/lat 21.35972)">Ta Xua NR</a>, residential area, 21.35972°N, 104.66972°E; 673 m a. s. l.; 11 Oct. 2017; S. X. Le and H. T. Vu leg.; HNUE.TX.D4.001 • 1 specimen; same location, bamboo forest; 21.33833°N, 104.69139°E; 429 m a. s. l.; 14 Oct. 2017; L. K. Ha leg.; HNUE.TX.142 . • 2 specimens; Hoa Binh Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.46142&amp;materialsCitation.latitude=20.66127" title="Search Plazi for locations around (long 105.46142/lat 20.66127)">Thuong Tien NR</a>, residential area; 20.66127°N, 105.46142°E; 175 m a. s. l., Jul. 2020; H. D. Nguyen leg.; HNUE.TT.202 to 203 . • 4 specimens; Phu Tho Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.95845&amp;materialsCitation.latitude=21.11426" title="Search Plazi for locations around (long 104.95845/lat 21.11426)">Xuan Son NP</a>, natural forest; 21.11426°N, 104.95845°E; 511 m a. s. l.; 13 Jun. 2014; S. X. Le leg.; VRTC.XS.015 to 016, VRTC.XS.018 to 019 • 4 specimens; same location, natural forest; 21.118219°N, 104.95159°E; 492 m a. s. l.; 10 Jun. 2014; S. X. Le leg.; VRTC.XS.006, VRTC.XS.009 to 010, VRTC.XS.021 • 4 specimens; same location, mixed wood – bamboo; 21.136413°N, 104.95244°E; 524 m a. s. l.; 12 Jun. 2014; S. X. Le leg.; VRTC.XS.011 to 014 • 1 specimen; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.92145&amp;materialsCitation.latitude=21.12622" title="Search Plazi for locations around (long 104.92145/lat 21.12622)">Xuan Son NP</a>, natural forest; 21.12622°N, 104.92145°E; 556 m a. s. l.; 14 Jun. 2014, S. X. Le leg.; VRTC.XS.025 . • 1 specimen; Vinh Phuc Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.64222&amp;materialsCitation.latitude=21.45627" title="Search Plazi for locations around (long 105.64222/lat 21.45627)">Tam Dao NP</a>, residential area; 21.45627°N, 105.64222°E; 1.001 m a. s. l.; Oct. 2022; A. D. Nguyen leg.; IEBR.VP.132 . • 1 specimen; Hai Phong city, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.68219&amp;materialsCitation.latitude=20.8254" title="Search Plazi for locations around (long 106.68219/lat 20.8254)">Duong Kinh</a>, residential area; 20.82540°N, 106.68219°E; 5 m a. s. l.; 16 May 2021; S. X. Le leg.; VRTC.HP.01 . • 8 specimens; Thai Binh Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.59306&amp;materialsCitation.latitude=20.25306" title="Search Plazi for locations around (long 106.59306/lat 20.25306)">Tien Hai NR</a>, coniferous forest; 20.25306°N, 106.59306°E; 8 m a. s. l.; 2021; H. D. Nguyen leg.; HNUE CHI 1066 to 1067, HNUE CHI 1081, HNUE CHI 1088, HNUE CHI 1092, HNUE CHI 1096, HNUE CHI 1098, HNUE CHI 1100 . • 1 specimen; Ha Noi city, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.35786&amp;materialsCitation.latitude=21.08191" title="Search Plazi for locations around (long 105.35786/lat 21.08191)">Ba Vi NP</a>, natural forest; 21.08191°N, 105.35786°E; 487 m a. s. l.; Aug. 2016; S. X. Le leg.; VRTC.BV.104 • 5 specimens; same location, residential area; 21.04000°N, 105.13389°E; 13 m a. s. l.; 8 Jan. 2019; H. D. Nguyen leg.; VRTC.HN.004 to 008 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.80093&amp;materialsCitation.latitude=21.04673" title="Search Plazi for locations around (long 105.80093/lat 21.04673)">Cau Giay district</a>, residential area; 21.04673°N, 105.80093°E; 9 m a. s. l.; 10 Sep. 2018; H. T. Vu leg.; IEBR.SP.01 . • 4 specimens; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.78258&amp;materialsCitation.latitude=20.946386" title="Search Plazi for locations around (long 105.78258/lat 20.946386)">Ha Dong district</a>, residential area; 20.946386°N, 105.78258°E; 8 m a. s. l.; 5 Dec. 2021; S. X. Le leg.; VRTC.KH.001 to 004 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.81969&amp;materialsCitation.latitude=20.99984" title="Search Plazi for locations around (long 105.81969/lat 20.99984)">Thanh Xuan district</a>, residential area; 20.99984°N, 105.81969°E; 9 m a. s. l.; 6 Jun. 2021; A. D. Nguyen leg.; VRTC.TX.001 • 7 specimens; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.836&amp;materialsCitation.latitude=21.07597" title="Search Plazi for locations around (long 105.836/lat 21.07597)">Tay Ho district</a>, agricultural area; 21.075970°N, 105.83600°E; 12 m a. s. l.; 12 Jan. 2022; S. X. Le and T. T. Do leg.; VRTC.HN.003, VRTC.HN.005 to 008, VRTC.HN.015, VRTC.HN.020 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.79577&amp;materialsCitation.latitude=21.06906" title="Search Plazi for locations around (long 105.79577/lat 21.06906)">Cau Giay district</a>, residential area; 21.069060°N, 105.79577°E; 15 m a. s. l.; May 2023, D. X. Le leg.; VRTC.HN.021 . • 1 specimen; Nghe An Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.23344&amp;materialsCitation.latitude=18.93889" title="Search Plazi for locations around (long 105.23344/lat 18.93889)">Pu Mat NP</a>, natural forest; 18.93889°N, 105.23344°E; 437 m a. s. l.; 17 Apr. 2018; S. X. Le leg.; VRTC.PM.033 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.18347&amp;materialsCitation.latitude=18.95917" title="Search Plazi for locations around (long 104.18347/lat 18.95917)">Pu Mat NP</a>, natural forest; 18.95917°N, 104.18347°E; 254 m a. s. l.; 23 Apr. 2018; S. X. Le leg.; VRTC.PM.034 . • 4 specimens; Dong Thap Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.52186&amp;materialsCitation.latitude=10.71746" title="Search Plazi for locations around (long 105.52186/lat 10.71746)">Tram Chim NP</a>, natural forest, 10.71746°N, 105.52186°E, 7 m a. s. l., 6 Oct. 2018, D. H. Lam leg.; VRTC.TrC.007 to 009, VRTC.TrC.011 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.52884&amp;materialsCitation.latitude=10.69942" title="Search Plazi for locations around (long 105.52884/lat 10.69942)">Tram Chim NP</a>, natural forest; 10.69942°N, 105.52884°E; 11 m a. s. l.; 18 Mar. 2023; S. X. Le and H. D. Nguyen leg.; VRTC. TrC 012 • 1 specimen; same location, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.53567&amp;materialsCitation.latitude=10.71075" title="Search Plazi for locations around (long 105.53567/lat 10.71075)">Tram Chim NP</a>, natural forest; 10.71075°N, 105.53567°E; 7 m a. s. l.; 20 Mar. 2023; S. X. Le and H. D. Nguyen leg.; VRTC.TrC 033 .</p><p>Diagnosis.</p><p>Body with 21 leg-bearing segments. LBS 7 th with spiracles. Antennae (Fig. 6 A) composed of 18–20 articles, of which three basal ones glabrous dorsally. Cephalic plate without lateral and posterior marginations; posterior edge straight, and covered by tergite 1 (Fig. 6 A). Tooth-plates with 4 (5) + 4 (5) teeth prominently; trochanteroprefemora of forcipular strong, with 3 + 4 prominent teeth (one apical and two-three lateral teeth)); coxosternite with short median suture starting from toothplate (Fig. 6 C). Tergites (Figs 6 B, 7 C) without punctate and short median sulcus; lateral margination absent (except for tergite 21, complete); tergites 6–20 with complete paramedian sutures and incomplete on tergites 3–5. Sternites 2–19 with incomplete (only 10–15 % length of sternites) paramedian sutures on anterior and posterior parts (Fig. 6 D). Coxopleural process with 1 (2) apical and 1 (2) subapical spine, lateral and dorsal spines absent (Fig. 7 B). Leg 1 with 1 fermoral, 1 tibial, and 2 tarsal spur (s) (Fig. 7 A). Ultimate leg (Fig. 7 D) prefemora with 1–2 VL, 0–1 VM, 0–1 M, and 0–2 DM spines.</p><p>Remarks.</p><p>R. immarginata is clearly distinguished from other Rhysida species in Vietnam by the absence of lateral tergite margination, except for tergite 21. This is also the characteristic supporting the division of Rhysida species into two main groups, the longipes group and the immarginata group, as mentioned in the study of Joshi et al. (2019).</p><p>Our examined specimens were consistent with the description in Siriwut et al. (2018) in most characteristics. However, the descriptions by Siriwut et al. (2018) do not mention the presence of a median suture on the coxosternite, which we observed to be present, albeit very briefly (Fig. 6 C).</p><p>Additionally, the length of the antennae reaches the boundary between tergites 5 and 6, which aligns with Siriwut et al. (2018) and is confirmed by Schileyko and Stoev (2016). This differs from Attems (1930) and Waldock and Lewis (2014), where the antennae are shorter, reaching only the posterior margin of tergite 3, and even shorter in Koch (1985).</p><p>Distribution.</p><p>This species occurs in Africa (Congo, Sudan), South America (Bolivia, Cuba, El Salvador, Guatemala, Peru, Venezuela, and Uruguay), and South and Southeast Asia (Taiwan, Cambodia, Indonesia, Kalimantan, Maluku, Myanmar, the Philippines, Singapore, Sri Lanka, Sumatera, Papua New Guinea, Malaysia, India, Nepal, Thailand, and Laos) (Bonato et al. 2016; Schileyko and Stoev 2016; Siriwut et al. 2018).</p><p>In Vietnam it was recorded in Quang Ninh Province (Ha Long Archipelago) and Nghe An Province (Vinh city) by Attems (1938), and Ha Giang, Dien Bien, Son La, Hoa Binh, Phu Tho, Vinh Phuc, Hai Phong, Thai Binh, Ha Noi, and Dong Thap in this study. The distribution of R. immarginata was further updated (Fig. 1).</p></div>	https://treatment.plazi.org/id/C89ADC33546457969E237C952A505C35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
4CFCAE0B8E105EDF82BC2CF559D24BA1.text	4CFCAE0B8E105EDF82BC2CF559D24BA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhysida longipes (Newport 1845)	<div><p>Rhysida longipes (Newport, 1845)</p><p>Figs 1, 4, 5</p><p>Rhysida longipes Kraepelin 1903: 148, fig. 91; Attems 1930: 193; Schileyko 2007: 82; Tran et al. 2013: 227; Schileyko and Stoev 2016: 257, figs 18, 22–24; Siriwut et al. 2018: 1017, figs 5 B, 6–8; Joshi et al. 2019: 835, figs 8.</p><p>Branchiostoma affine Kohlrausch, 1878: 67; Kraepelin 1903: 148.</p><p>Otostigmus simplex Attems, 1930: 153; Lewis 2002: 1690, figs 8–11.</p><p>Examined materials.</p><p>Vietnam • 1 specimen; Dien Bien Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.39833&amp;materialsCitation.latitude=21.21278" title="Search Plazi for locations around (long 103.39833/lat 21.21278)">Muong Nhe NR</a>, residential area; 21.21278°N, 103.39833°E; 610 m a. s. l., May 2018; H. D. Nguyen leg.; DB 020 ; • 23 specimens; Thai Binh Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.59306&amp;materialsCitation.latitude=20.25306" title="Search Plazi for locations around (long 106.59306/lat 20.25306)">Tien Hai NR</a>, coniferous forest; 20.25306°N, 106.59306°E; 08 m a. s. l.; 2021; H. D. Nguyen leg.; HNUE CHI 1068 to 1070, 1072, 1073, 1075 to 1080, 1082, 1083, 1085, 1087, 1088, 1090, 1091, 1095, 1096, 1099, 1101, 1103 ; • 1 specimen; Ha Nam Province, agricultural land; 20.59323°N, 105.97150°E; 6 m a. s. l.; 2017; H. T. T. Nguyen leg.; Hna.001; • 1 specimen; Hanoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.3592&amp;materialsCitation.latitude=21.08346" title="Search Plazi for locations around (long 105.3592/lat 21.08346)">Ba Vi NP</a>, natural forest; 21.08346°N, 105.35920°E; 460 m a. s. l.; Aug. 2016; S. X. Le leg.; BV.SL.013 ; • 7 specimens; Hanoi, agricultural land • 3 specimens; Hanoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.13389&amp;materialsCitation.latitude=21.04" title="Search Plazi for locations around (long 105.13389/lat 21.04)">Cau Giay district</a>; 21.04000°N, 105.13389°E; 13 m a. s. l.; 2017; H. D. Nguyen &amp; B. T. T. Tran leg.; HN 001, HN 101, HN 102 ; • 2 specimens; Hanoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.79577&amp;materialsCitation.latitude=21.06906" title="Search Plazi for locations around (long 105.79577/lat 21.06906)">Tay Ho district</a>; 21.06906°N, 105.79577°E; 15 m a. s. l.; May 2023; D. X. Le leg.; VRTC.HN 022, VRTC.HN 023 • 1 specimen; Hanoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.88749&amp;materialsCitation.latitude=21.05233" title="Search Plazi for locations around (long 105.88749/lat 21.05233)">Long Bien district</a>, 21.05233°N, 105.88749°E; 10 m a. s. l.; 20 Aug. 2023; S. X. Le leg.; VRTC.HN.025 ; • 5 specimens; QUANG BINH Province, coniferous forest; S. X. Le leg.; 11 May 2022: VRTC.BTr.001 and VRTC.BTr.003: 17.49535°N, 106.62392°E, 5 m a. s. l.; VRTC.BTr.002: 17.49481°N, 106.62323°E, 5 m a. s. l.; VRTC.BTr.004: 17.49823°N, 106.62072°E, 10 m a. s. l.; VRTC.BTr.005: 17.49949°N, 106.62004°E, 8 m a. s. l.; • 3 specimens collected by S. X. Le in residential area on 11 May 2022: VRTC.BTr.006: 17.49769°N, 106.61829°E; 8 m a. s. l.; VRTC.BTr.007 and VRTC.BTr.008: 17.49795°N, 106.61936°E, 6 m a. s. l.; • 202 specimens; Khanh Hoa Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.33333&amp;materialsCitation.latitude=12.0" title="Search Plazi for locations around (long 117.33333/lat 12.0)">Truong Sa islands</a>; residential area, 6.50000 –12.0000 °N, 111.50000– 117.33333°E; below 15 m a. s. l.; 2020 and 2021; S. X. Le, T. N. Nguyen, D. T. Dinh and H. B. Truong leg.; KCB.TS.04.001 to 003, 011, 016, 019, 021 to 023, 033, 040, 046, 102, 103, 201, 203 to 204, 206 to 209, 214 to 215, 217, 219 to 229, 301 to 303, 305 to 307, 310, 402 to 411, 414 to 421, 423 to 425, 427 to 431, 501, 506, 511 to 512, 515 to 518, 520, 526 to 527, 529, 531 to 534, 536 to 537, 539 to 544, 601 to 618, 620 to 621, 623 to 632, 701, 702 to 721, 723 to 726, 728 to 740, 745 to 746, 801 to 804, 807 to 819, 821 to 830, 832, 837, 840 to 842, 844 to 846, 901 to 904, 906, 908 .</p><p>Diagnosis.</p><p>Body with 21 LBS. LBS 7 th with spiracles. Antennae (Fig. 4 A, B) composed of 18 articles, of which three basal ones glabrous dorsally. Cephalic plate (Fig. 4 A) without lateral and posterior marginations; posterior edge straight and covered by tergite 1. Forcipular segment (Fig. 4 B) smooth; tooth plates with 4 + 4 teeth obviously; coxosternite with short median suture starting from toothplate. Tergites without short median sulcus; paramedian sutures complete on tergites 4 (5) – 19 (Figs 4 C, 5 A); lateral marginations incomplete on tergites 6 (10) – 20 and complete on tergite 21 (Figs 4 C, 5 A). Sternites 3–20 with incomplete (10–15 % length of sternites) paramedian sutures (Figs 4 B – D, 5 B). Coxopleural process long with 2 apical, 1 subapical, and 1 lateral spine, without dorsal spine (Fig. 5 B). Leg 1 present or absent femoral spur (Fig. 4 A, B). Prefemoral spines on ultimate legs with 2–3 VL, 2–5 VM, 0–1 M, and 1–3 DM spines; prefemoral process of ultimate legs very short with 1 spine (Fig. 5 C).</p><p>Remarks.</p><p>Our examined specimens have similar morphological characteristics to those described by Siriwut et al. (2018) and Joshi et al. (2019). There is a small difference in the starting of lateral marginations and paramedian sutures. Our specimens showed the lateral marginations start from tergite 4 (5) and paramedian sutures start from 6 (8), but these start from tergites 5–8 and tergites 8–11, respectively, in Siriwut et al. (2018), and tergites 7–8 and tergites 4–7, respectively, in Joshi et al. (2019).</p><p>Distribution.</p><p>This species is widespread in tropical mainland and island regions. It occurs in Africa (Madagascar, Seychelles, and Mauritius), Central and South America (Mexico, Peru, Cuba, and Haiti), and Central, East, and Southeast Asia (China, Taiwan, Nepal, India, Sri Lanka, Chagos Islands, Yemen, Pakistan, Iran, Myanmar, Thailand, Cambodia, Malay Peninsula, Philippines, and Papua New Guinea) (Siriwut et al. 2018; Joshi et al. 2019).</p><p>In Vietnam, it has been recorded from Thua Thien Hue (Hai Van Pass); Dong Nai (Ma Da); Khanh Hoa (Spratly Islands); and Kien Giang (Phu Quoc Island) (Attems, 1938; Schileyko, 2007; Son et al. 2022); and in Dien Bien, Thai Binh, Hanoi, Ha Nam, and Quang Binh (in this study). The distribution of R. longipes was further updated (Fig. 1).</p></div>	https://treatment.plazi.org/id/4CFCAE0B8E105EDF82BC2CF559D24BA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.text	855610FE43C859B592E133D707581960.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhysida vietnga Le & Do & Tran & Nguyen 2025	<div><p>Rhysida vietnga sp. nov.</p><p>Figs 1, 8, 9</p><p>Examined materials.</p><p>Holotype: Vietnam • 1 specimen; Lao Cai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.77778&amp;materialsCitation.latitude=22.4156" title="Search Plazi for locations around (long 103.77778/lat 22.4156)">Hoang Lien NP</a>, mixed bamboo forest; 22.4156°N, 103.77778°E; 1,529 m a. s. l.; 15 Apr. 2019; H. D. Nguyen leg.; HLS.064.</p><p>Paratypes: • 2 specimens; same location; 22.4156°N, 103.77778°E; 1,529 m a. s. l.; 15 Apr. 2019; S. X. Le &amp; H. D. Nguyen leg.; HLS.006, HLS.092; • 2 specimens; Son La Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.67333&amp;materialsCitation.latitude=21.35722" title="Search Plazi for locations around (long 104.67333/lat 21.35722)">Ta Xua NR</a>, mixed bamboo forest; 21.35722°N, 104.67333°E; 726 m a. s. l.; 11 Oct. 2017; S. X. Le &amp; H. T. Vu leg.; TX.D4.002, TX.D4.003; 1 specimen • same location; natural forest; 21.35972°N, 104.66972°E; 722 m a. s. l.; 30 May 2018; H. D. Nguyen leg.; TX.258 .</p><p>Diagnosis.</p><p>Body with 21 leg-bearing segments. LBS 7 th with spiracles. Antennae composed of 18 articles, of which three basal ones glabrous dorsally. Tooth-plate with five short teeth. Tergites with lateral marginations and 5–7 keels and many spinulations scattered on tergites, more obvious in the posterior tergites, without complete paramedian sutures. Sternites with paramedian suture incomplete on anterior part (on some sternites). Coxopleural process with 2 apical, 1 lateral spines and without dorsal spine.</p><p>Description.</p><p>Body length 58 mm. Antennae composed of 21 articles, of which three basal ones glabrous dorsally, following segments densely covered by minute setae (Fig. 8 A, B). Antenna length reaching to 6 th body segment.</p><p>Cephalic smooth, without puncta and setae; lateral and posterior margination absent, the posterior border adjacent to anterior border of tergite 1. Four big ocelli present, nearly equal in size. The median suture on the top of cephalic plate long, shallow (Fig. 8 A).</p><p>Forcipular segment (Fig. 8 B), coxosternite, trochanteroprefemora, and the basal part of tarsungulae smooth, without punctate and setae; coxosternite without median suture. Tooth-plates slightly wider than high and nearly semicircular, with 5 short teeth and well chitinized; the distance between the tooth – plates wide; each tooth-plate with straight, transverse basal suture. Forcipular trochanteroprefemoral process bearing 3 short denticles, the latter being indistinct, position of denticles higher than tooth – plates. Maxillae with sparse short setae, and segment 2 of second maxillary telopodite with spur; chitin-line very short.</p><p>Tergites with very small punctae; lateral margination incomplete on tergites 9–20 and complete on tergite 21; paramedian sutures absent on all tergites; tergites 1–7 smooth, and tergites 8–20 with 5–7 keels and many spinulations which scattered on tergites, more obvious in the posterior tergites; tergite of ultimate leg-bearing segment sub-quadrangular and convex posteriorly (Figs 8 D, 9 A).</p><p>Sternite surface smooth (without punctae), lacking any median sulci. Sternites shield-shaped, last sternite trapezoid (or nearly triangular). Incomplete paramedian suture on anterior part of some sternites (&lt;10 % of their width) or absent, lacking depression (Figs 8 B, C, 9 B).</p><p>Coxopleuron densely covered by pores (except the apical part); coxopleural process with 2 apical and 1 lateral spine, without dorsal spine (Fig. 9 B).</p><p>Locomotory legs usually bearing small, short setae; legs 1–4 with 1 tibia spur, legs 1–18 with 2 tarsal spurs, 19–20 with 1 tarsal spur. Ultimate legs very thin and long; the ratio of lengths of prefemur, femur, tibia, tarsus 1, and tarsus 2 as follows: 2.8: 2.7: 2.5: 2.0: 1.0; ratio of length and width of prefemur 6: 1. Prefemoral spines on ultimate leg: 4 VL, 3 VM, 3 M, and 1 DM spines; prefemoral process of ultimate legs very short with 1 spine; all of segment smooth (Fig. 9 C).</p><p>Variations.</p><p>Body length reaches a maximum of 66 mm (TX.D4.002). Specimens from Ta Xua NR (Son La) show that the lateral margination starts from tergite 5 (TX.258) or tergite 6 (TX.D4.003). The keels on the tergites from specimen TX.D4.002 present on tergites 12–20. Locomotory legs with tibia spines from legs 1–4 or legs 1–6 (HLS.092). Prefemoral spines on ultimate leg can have 3–4 VL, 2–3 VM, 3–4 M, and 1–4 DM spines.</p><p>Etymology.</p><p>The species name is named after the Joint Vietnam-Russia Tropical Science and Technology Research Center (VRTC), an institution that has significantly contributed to biodiversity research in Vietnam.</p><p>Remarks.</p><p>The new species differs from R. immarginata in the clearly visible but incomplete lateral margins on the tergites (absent in R. immarginata, except for tergite 21). The absence of a paramedian suture on the sternites is a distinguishing feature that separates the new species from Alluropus calcaratus . Rhysida vietnga sp. nov. is distinguished from R. longipes and R. lithobioides by the presence of keels on the tergites. This is also a character to distinguish the new species and its congeners.</p><p>Rhysida vietnga exhibits morphological similarities to Rhysida singaporensis Verhoeff, 1937, particularly in the distinctive structure of the dorsal plate with keels. This characteristic also distinguishes these two species from other members of the genera Rhysida and Alluropus . Therefore, a morphological comparison between the new species R. vietnga and R. singaporensis is most appropriate. The distinction between these two species can be made based on the following characters: The new species, based on six specimens examined in this study, is markedly smaller than R. singaporensis, with a maximum body length of 66 mm and an average of 43 mm, whereas R. singaporensis typically ranges from 73 to 89 mm in length. According to descriptions of Lewis (2001) and Siriwut et al. (2018), R. singaporensis has a median keel on the tergites, while the lateral longitudinal keels are either not mentioned or described as short (incomplete); if present, longitudinal keels end with numerous spines. However, the illustrations in these studies do not show the presence of complete longitudinal keels. In contrast, R. vietnga has 5–7 distinct, complete longitudinal keels on tergites 8–20 (similar to those of Otostigmus scaber; see Lewis 2001, p. 26, fig. 49), along with abundant spinulation scattered across the tergites, which becomes more pronounced on the posterior tergites (Fig. 9 A). The comparative morphological data for Southeast Asian species are summarized in Table 1.</p><p>Data sources: Lewis (2001), Siriwut et al. (2018), and Joshi et al. (2019).</p><p>In addition, R. singaporensis is distributed in the archipelagic region, likely limited to the Malay Archipelago, while R. vietnga is found in the mainland area of the Indochina Peninsula.</p><p>Distribution.</p><p>Only known in Lao Cai province and Son La province in Vietnam (Fig. 2).</p></div>	https://treatment.plazi.org/id/855610FE43C859B592E133D707581960	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
A9F5EF03B7805D64815DE99341A1804D.text	A9F5EF03B7805D64815DE99341A1804D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhysida Wood 1862	<div><p>Genus Rhysida Wood, 1862</p><p>Type species.</p><p>Rhysida lithobioides (Newport, 1845) .</p><p>Diagnosis.</p><p>Body with 21 LBS. LBS 7 th with spiracles. Antennae composed of 16–21 articles, of which 2.8–3 basal ones glabrous dorsally. Cephalic plate without lateral and posterior marginations. Forcipular tooth-plates present; tooth-plates with 4–7 teeth; trochantero-prefemur with well-developed process; coxosternite with short median suture starting from tooth-plate. Sternites with incomplete paramedian sutures. Coxopleural process long with 1–2 apical, 0–2 subapical, and 0–1 lateral spines, without dorsal spines. Leg 1 with 1 femoral, 1 tibial, and 2 tarsal spurs.</p><p>Distribution.</p><p>Neotropics, Africa and Indomalaya, and Australia.</p></div>	https://treatment.plazi.org/id/A9F5EF03B7805D64815DE99341A1804D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le, Son Xuan;Do, Thinh Tat;Tran, Binh T. T.;Nguyen, Anh D.	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
