taxonID	type	description	language	source
4D413F2D0F2E9E3BFF2FF8E268E94B67.taxon	materials_examined	Holotype: NHMUK ZD 1909.1. 4.70 Phyllorhina brachyota, Central India, Staff Surgeon F. P. Staples; skull extracted; in good condition. Other material: NHMOU. CHI. 54.2016; NHMOU. CHI. 7.2014; NHMOU. CHI. 31.2014, NHMOU. CHI. 32.2014; NHMOU. CHI. 35.2014; NHMOU. CHI. 1.2003; ZSI 20551, ZSI 20552	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F2E9E3BFF2FF8E268E94B67.taxon	diagnosis	Diagnosis: A medium-sized bat with a forearm length ranging between 45.6 – 49.6 mm. Ears triangular. The noseleaf is broad, with two pairs of supplementary leaflets. The posterior leaf has three septa, clearly divided into four cells. Skull short, condylo-canine length 14.73 ± 0.33 mm. Maxillary toothrow (CM 3) 5.4 – 6.3 mm. The first upper premolar (PM 2) minute is wedged between the canine and PM 4, sometimes extruded or in toothrow. First lower premolar (PM 2) half the height of PM 4, one-fourth the height of the canine. The baculum is small, with a slightly broader base and slender shaft ending with a blunt tip, faintly curved in lateral view. External characters: A medium-sized hipposiderid with an average forearm length of 47.17 ± 1.33 mm (45.6 – 49.6 mm). Ears are short (12.94 ± 1.81 mm; 10.0 – 15.3 mm), wider at the base, with thick fur covering half the height of the pinna on the dorsal surface (Figure 6). Tip narrowly pointed and long. The outer border of the pinna shows concavity just before the tip. The inner border is covered with thick hair except for the apical region. Eight transverse ridges present on the outer border of the pinna. The noseleaf is broad, covers the entire muzzle and is greyish-brown (Figure 6). The anterior leaf is broader than the posterior and intermediate leaves. Just below the anterior leaf, two pairs of supplementary leaflets are present; the second pair of supplementary leaflets is slightly longer than the first pair (Figure 6). The internarial septum is located between nares and does not touch the intermediate leaf; it is roughly triangular and ends with a blunt tip. Nares are transverse and long with well-developed narial lappets on the sides of the nares. The intermediate leaf is thick, with concavity toward the middle, and is shorter than the anterior and posterior leaves. The posterior leaf has three septa, dividing it into four distinct cells. Wing and interfemoral membrane attached on either side of the ankle. In the wing, both the third and fourth metacarpals are almost equal, while 5 th metacarpal is shorter than 3 rd and 4 th metacarpals. The first and second phalanges are 41.1 % and 52.3 % of the third metacarpal respectively. Craniodental Characters: Skull is small and delicate with an average greatest total length of 17.36 ± 0.27 mm (17.0 – 17.69 mm). Sagittal crest is low, slightly pronounced in the frontal region but not so through the rest of its length and extends up to the parietal region (Figure 7 A). The lambdoid crests are not well developed. The Interorbital region is narrow. Rostrum is bulged, with four well-developed nasal inflations; the bottom one is the largest, and the median one is thin and almost dumbbell-shaped, nasal inflations are kidney-shaped and located on either side of the rostrum (Figure 7 B). Zygoma are robust and lack dorsal process (Figure 7 A). Upper incisors are simple, small and bicuspidate (Figure 7 E). Canines are robust and tall. The first upper premolar (PM 2) is minute, slightly extruded from the toothrow, wedged between the second upper premolar (PM 4) and the canine, and in contact with PM 4 (Figure 7 A). Metastyle of all three upper molars is well developed; paracone of M 1 is slightly longer than metacone; in M 2 paracone and metacone are equal in length. Hypoconal flange is well-developed in all three upper molars but not touching each other or PM 4. Mesostyle and commissures are thick. Protocone of both M 1 and M 2 is well-developed. In M 3, however, the metastyle and the hypoconal flange are lacking. Canines of the lower toothrow are delicate and tall (Figure 7 C). The first and second lower premolars (PM 2 and PM 4) are not in contact. The second premolar is one-third the height of the canine and half the height of the fourth premolar. Two pairs of tricuspidate lower incisors are present in a single line (Figure 7 F). Pelage color: Fur is dark brown to fawn on the dorsal surface. Around the muzzle the fur is darker and colored similar to the dorsal surface. On the head, hairs have pale bases with fawn tips. The area under the face is similarly colored as the head, while the belly hairs are off-white. Wing and interfemoral membranes are dark and hairless with fur of the body not extending onto the interfemoral membrane. Baculum structure: The baculum is simple, and small (0.7 mm; n = 1) with a slightly wide base, and a slender shaft ending with a bluntly rounded tip (Figure 4 A). Laterally, the shaft shows a slight concavity. Common name: Dobson’s Leaf-nosed Bat	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F2E9E3BFF2FF8E268E94B67.taxon	biology_ecology	Ecology: This species is rare in comparison to other leaf-nosed bats. It has been observed to roost in small colonies of 2 – 3 to up to 10 individuals, or sometimes singly. It roosts in narrow caves located on the top of hills, among large boulders, old disused rooms of houses, and old disused temples, mines, dungeons, or even in cracks in the walls of houses (Khajuria 1980; present study).	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F2E9E3BFF2FF8E268E94B67.taxon	distribution	Distribution: India — Maharashtra: Ajanta and Ellora caves, Bedsar caves, Chikalda, Shivthatghal near Mahad, and Kanheri caves. Karnataka: Avani, Badami, Honawar, Lingsugur, Shivagange, and Ukkarahalli. Kerala: Alampara forest cave, Veliangad. Telangana: Akkamahadevibilam. Andhra Pradesh: Donkarai. Odisha: Badudi Pahad, Hadgarh Wildlife Sanctuary, Hemagiri, Khandadhar, Koilighogar, Gupteswar caves, Similipal Biosphere Reserve, and Sunabeda Wildlife Sanctuary. Gujarat: Danta, Palanpur, Tera Mines, and Jambugoda Wildlife Sanctuary. Madhya Pradesh: Gwarighat, Jabalpur, and Kanha National Park. Bihar: Singar. Bangladesh: Chittagong, Dhaka, and Sylhet (Bates & Harrison 1997; Debta & Palita 2017; Ghosh 2008; Srinivasulu 2004; Srinivasulu & Srinivasulu 2005, 2017) (Figure 1).	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F299E27FF2FFB146F784B2F.taxon	materials_examined	Holotype: NMSL 2023.08.01 NH, adult male preserved in alcohol with the skull removed and clean, collected by Tharaka Kusuminda and Sahan Siriwardana on 06 May 2018. Type locality: Makulella (Bandarawela), Badulla district, Uva Province, Sri Lanka. Other material: ZSI 20553; ZSI 20554; ZSI 20555; ZSI 20556; BNHS 3687; HZM 9.28779; HZM 10.28780; HZM 12.30955; HZM 13.31080; HZM 15.40112; NMSL 15 A, NMSL 15 O, NMSL 15 L, NMSL 15 H, NMSL 15 K, NMSL 15 N.	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F299E27FF2FFB146F784B2F.taxon	diagnosis	Diagnosis: Medium-sized bat with forearm length ranging between 45.07 – 53.1 mm. Ears tall, broad, with moderately developed antitragus. Tragus absent. Noseleaf broad, covering the whole muzzle; two pairs of supplementary leaflets. Posterior leaf with three septa clearly dividing it into four cells. Skull narrow, long, condylocanine length 15.66 ± 0.45 mm. Maxillary toothrow (CM 3) 6.05 ± 0.11 mm. First upper premolar (PM 2) small but within toothrow. First lower premolar (PM 2) half the height of PM 4, one-third the height of canine. Baculum small, base and apex trilobed with straight parallel-sided shaft in lateral view. External characters (based on holotype): A medium-sized hipposiderid with a forearm length of 49.99 mm. Ear moderately tall (16.57 mm) and broad, ending with a pointed tip; antitragus is moderately developed, roughly rectangular, one-third the height of the pinna; tragus absent (Figure 8 A, C). The outer border of the pinna narrows gradually to a narrowly rounded tip, with slight concavity just before the tip of the pinna. The outer border with nine transverse sparsely-haired ridges, inner border of pinna hairy. The noseleaf is broad covering the whole of the muzzle; anterior leaf is as broad as the posterior leaf; two pairs of supplementary leaflets are seen immediately beneath the anterior leaf; first pair of supplementary leaflets are longer than the second, almost touching near the median margin of the anterior leaf; second pair of supplementary leaflets three-quarter the length of first pair (Figure 8 B). The anterior leaf, supplementary leaflets, and the posterior leaf are greyish black. The internarial septum is greyish black, parallel-sided, long, tapers to a point, touching the intermediate leaf, and lower than the margin of nares (unlike in other hipposiderids wherein the internarial septum is slightly raised from the rest of the noseleaf). Nares are long and transverse; narial lappets are well developed, and located on the sides of nares. The intermediate leaf is heart-shaped with a concave median superior border; and is slightly smaller than the anterior leaf. The posterior leaf has a slight median projection at the same position as the concavity in the intermediate leaf. Three septa divide the posterior leaf into four distinct cells. The wing is attached to the base of the toe, and the interfemoral membrane is attached to the ankle (Fig. 8 E). Fourth metacarpal longest, while the fifth metacarpal is the shortest. The first and the second phalanges are 40.50 % and 48.86 % of the third metacarpal respectively. The shaft of the penis is straight, cylindrical, and tapering towards the tip (Figure 8 D). Craniodental characters: The skull is narrow and delicate (GTL: 19.18 mm). The sagittal crest is visible but not pronounced, extending up to the parietal region (Figure 9 A). The interorbital region is broad. Rostrum is bulged, with three well-developed nasal inflations; the bottommost one is the largest, and the median one the smallest, and round; kidney-shaped nasal inflations are located on either side of the rostrum (Figure 9 B). The zygoma are delicate (Figure 9 A). Upper incisors are simple, small, and bicuspid (Figure 9 E). Canines are delicate and tall; the first upper premolar (PM 2) is small, within the toothrow, located between the second upper premolar (PM 4) and canine (Figure 9 A). The parastyle of the first and second upper molars (M 1 and M 2) is well developed; the metacone of M 1 is longer than the paracone; the paracone of M 2 is slightly longer than the metacone. The hypoconal flange is well-developed in M 1, both toward PM 4 and M 2, which is why the hypoconal flange of M 2 is in contact with the mesostyle; the commissures are thick. Metastyle of protocone of all three molars well-developed. The metacone and paracone of M 3 are well developed but lack metastyle and hypoconal flange. The canine of the lower toothrow is narrow and tall. The first and second lower premolars (PM 2 and PM 4) are in contact (Figure 9 C). The second premolar is one-third the height of the canine, and half the height of the fourth premolar. Two pairs of tricuspidate lower incisors present in a single line (Figure 9 F). Pelage color: Fur dark brown to fawn on the dorsal surface. Around the muzzle, the fur is darker, colored similar to the dorsal surface. On the head, hairs have pale bases with fawn tips. The area under the face is similarly colored as the head, while belly hairs are off-white. Wing and interfemoral membranes are dark, hairless with the fur of the body not extending onto the interfemoral membrane. Baculum structure: The baculum is small (0.85 mm, N = 1), with a parallel-sided shaft. The proximal end has a median rounded portion with two projections on either side, and the distal end has two prongs with space in between (Figure 4 b). Common name: Sri Lankan Leaf-nosed Bat	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F299E27FF2FFB146F784B2F.taxon	biology_ecology	Ecology: This species has been recorded to roost in small colonies comprising a male, a few females and young, in small caves, old houses, churches, beneath overhanging rocks, and among boulders (Phillips 1923; Kusuminda et al. 2022 a). It has been known to frequent open areas for foraging and is found from the low country to elevations of 1067 m (3500 ft: Phillips 1923).	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F299E27FF2FFB146F784B2F.taxon	distribution	Distribution: This species is endemic to Sri Lanka, and has been recorded from North Western Province: Kala Oya, Puttalam, Wilpattu National Park and Kurunegala. Central Province: Gammaduwa, Kumbalgamuwa and Ginigathhena. Western Province: Bulathsinghala, Anasigalla, Kalutara, Labugama and Pilikuttuwa. Uva Province: Makulella (Bandarawela), Walhaputenna and Passara. Northern Province: Mannar. Sabaragamuwa Province: Balangoda, Kitulgala, Kegalle, Deraniyagala and Wavulpane. Southern Province: Pitabeddera (Phillips 1980; Bates & Harrison 1997; Yapa 2017; Kusuminda et al. 2022 a; Present study) (Figure 1). Comparisons: Cantor (1846) described the noseleaf to be large in H. galeritus with the transverse leaf (= posterior leaf) being as broad as the anterior leaf and almost in contact with the ears. Dobson (1874, 1876), in his description of Phyllorhina brachyota (= H. brachyotus) mentioned that the transverse leaf is rectangular, with the superior margin being less convex, and with a concave front portion with three prominent vertical ridges. In fresh specimens of H. brachyotus the posterior leaf is rectangular, divided into three ridges but does not show any concavity. The intermediate leaf shows a slight concavity and a slight projection in the median area and does not exceed the width of the anterior leaf (Figure 10 A). The noseleaf of H. srilankaensis is broad and large, covering the whole of the muzzle and is almost in contact with the broad ears. The posterior leaf shows a well-developed medial projection while being divided by three ridges, while the intermediate leaf shows a pronounced “ V ” shaped concavity in the median region (Figure 10 B). The frontal pore was mentioned to be small and indistinct by Dobson (1874), which is so in fresh specimens of H. brachyotus. In H. srilankaensis, however, the frontal pore is clearly evident, though it is absent in H. galeritus (Cantor, 1846). The noseleaf of H. galeritus from South Thailand is as broad as long (Figure 10 C). The posterior leaf is tall, similar to that of H. srilankaensis. In the noseleaf of H. galeritus from NE Thailand the internarial septum is short, parallel-sided, and ends with a bluntly rounded tip (Figure 10 D), rounder than that in H. brachyotus. The internarial septum is spindle-shaped in the South Thailand specimen, while in H. srilankaensis the internarial septum has a broad base and tapers to a point, touching the intermediate leaf. The ears of H. brachyotus are very small, as broad as long, with the anterior margin notably convex and the outer margin slightly concave beneath the tip (Dobson 1874). In live specimens, the ears are evidently broader than long. The ears of H. galeritus were mentioned to be sub-erect, broader than long, their breadth equaling the length of the head (Cantor 1846). In H. srilankaensis, the ears are broad and erect. In H. galeritus more than two-third of the back of the ear is covered with fur, with only a narrow hairless line along the external margin of the ear (Cantor 1846), a character which is present in both H. barchyotus and H. srilankaensis. The zygoma are robust and lack the dorsal process in H. brachyotus (Figure 11 A), whereas in H. srilankaensis they are slender and delicate, and lack the dorsal process (Figure 11 B), while in H. galeritus from Tarutalos, South Thailand the zygoma are slender with a prominent dorsal process (Figure 11 C). The sagittal crest is slightly pronounced in the frontal region in H. brachyotus, visible but not pronounced in H. srilankaensis, and slightly pronounced in H. galeritus. The skull is comparatively bulbous in H. srilankaensis, more than in H. brachyotus and H. galeritus (Figure 11 A, B, C). The palate is wider and longer in H. srilankaensis in comparison to H. brachyotus. The mesopterygoid space in H. brachyotus is an almost O-shape, while in H. srilankaensis it is a blunt V-shape. The canine of the lower jaw is tall and slender in H. srilankaensis, while in H. brachyotus it is not as tall, while the 2 nd premolar reaches three-quarters the height of the canine.	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F339E21FF2FFF6169064ECB.taxon	materials_examined	INDIA: Andhra Pradesh, East Godavari District, Donkarai (17.928 N, 81.805 E) (NHMOU. CHI. 7.2014, male); Karnataka, Kolar District, Ukkarahalli, near KGF (12.915 N, 78.237 E) (NHMOU. CHI. 31.2014, female; NHMOU. CHI. 32.2014, female); Karnataka, Bengaluru Rural District, Shivagange (13.169 N, 77.222 E) (NHMOU. CHI. 35.2014, male); Karnataka, Raichur district, Lingsugur (16.17 N, 76.52 E) (ZSI 20551, female); Karnataka, Dharwar district, Honkan (14.6440200 N, 75.2010800 E) (ZSI 20552, unknown); Kerala, Kozhikode District, Alampara Forest Cave (11.634 N, 75.868 E) (NHMOU. CHI. 54.2016, male); Telangana State, Akkamahadevibilam cave (16.1577 N, 78.834 E) (NHMOU. CHI. 1.2003, male).	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F339E21FF2FFC6D6EA64AFF.taxon	materials_examined	SOUTHEAST ASIA: Cambodia, Kampot Province, Kampot Limestone Cave (10.594242 N, 104.164032 E) (HZM 13.33423, female); Cambodia, locality unknown (HZM 14. A. 36201, male); Thailand, Kalasin Province, Pha Pheung Cave (19.432 N, 101.067 E), Phu Si Than WS (PSUZC-MM 2007.248, male); Thailand, Kalasin Province, Tham Bing (17.200 N, 103.979 E), Phu Si Than WS, (PSUZC-MM 2007.247, female); Thailand, Pha Taem NP (15.398 N, 105.507 E), Ubon Ratchathani (PSUZC-MM 2007.246, female); Thailand, Khao Singto (13.983 N, 102.016 E), Sa kaeo (PSUZC-MM 2007.236, female); Thailand, Bo Thong St., Khao Ang Ru Nai WS (13.416 N, 101.877 E), Chacherngsao (PSUZC-MM 2005.65, male); Thailand, Silawan Cave (10.708 N, 99.243 E), Chumphon (PSUZC-MM 2007.238, male); Thailand, Knad Dai Cave (10.032 N, 98.919 E), Ranong (PSUZC-MM 2007.239, male); Thailand, Pra Kayang Cave (10.326 N, 98.764 E), Ranong (PSUZC-MM 2007.237, male); Thailand, Koh Mai Pai (8.342 N, 98.529 E), Phang Nga (PSUZC-MM 2008.99, male); Thailand, Ban Nai Khao (7.70819 N, 99.6901 E), Trang (PSUZC-MM 2007.352, male); Thailand, Ao Son, Tarutao Is. (6.629 N, 99.998 E), Satun (PSUZC-MM 2005.66, male); Thailand, KM 8 to Talowow, Tarutao Is. (6.621 N, 99.679 E), Satun (PSUZC-MM 2008.53, female); Thailand, Bala (5.823 N, 101.840 E), Narathiwat (PSUZC-MM 2012.17, female).	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
4D413F2D0F339E21FF2FFD9D68F54C9B.taxon	materials_examined	SRI LANKA: Northwestern Province, Kala Oya (8.20481 N, 80.10375 E) (ZSI 20553, female; ZSI 20554; female; ZSI 20555; Male, ZSI 20556, female; BNHS 3687, male); Uva Province, Makulella, Bandarawela (6.8394 N, 81.0488 E) (NMSL 2023.08.01 NH, male) (Holotype); Uva Province, Walahaputharina (now Walhaputhenna, 6.764077 N, 80.871051 E) (HZM 9.28779, male); Uva Province, Walhaputhenna, Haldumulla (6.765278 N, 80.889167 E) (HZM 10.28780, juvenile female); Uva Province, Walhaputhenna, near Beragala (6.763985 N, 80.914925 E) (HZM 15.40112); Sabaragamuwa, Dantea Watta, near Ruwanawella (7.03361111 N, 80.25027778 E) (HZM 13.31080, female; HZM 12.30955, female); Northwestern Province, Kurunegala (7.483333 N, 80.366667 E) (NMSL 15 A, male); Northwestern Province, Puttalam (8.034167 N, 79.835278 E) (NMSL 15 K, male; NMSL 15 L, male); Central Province, Gammaduwa (7.566667 N, 80.7 E) (NMSL 15 H, female); Central Province, Kumbalgamuwa (7.11297 N, 80.85497 E) (NMSL 15 N, male; NMSL 15 O, male)	en	Srinivasulu, Bhargavi, Kusuminda, Tharaka, Srinivasulu, Aditya, Ukuwela, Kanishka D. B., Amarasinghe, Chamara, Siriwardana, Sahan, Kaur, Harpreet, Mannakkara, Amani, Soisook, Pipat, Kamalakannan, Manokaran, Yapa, Wipula Bandara, Srinivasulu, Chelmala (2025): Taxonomic revision of the South Asian allies of Hipposideros galeritus Cantor, 1846. Zootaxa 5590 (4): 507-530, DOI: 10.11646/zootaxa.5590.4.3, URL: https://doi.org/10.11646/zootaxa.5590.4.3
