taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
5B308C1FFFED17743781A95EFAD3A893.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819079/files/figure.png	https://doi.org/10.5281/zenodo.15819079	FIGURE 2. Live specimens of Metaphotina bimaculata (A–F) and Metaphotina austri (G–J) retrieved from iNaturalist platform, license CC-BY-NC. Images may have been cropped and/or rotated for better observation. (A) Observation 163494503, by goncrisdi [GonzaloDias],Argentina, 06/May/2023.(B) Bernardo Ferraz, Brazil (MG).(C) Observation 199319301, by jcsullivan [John Sullivan], Bolívia, Jan/2024. (D) Observation 223539397, by lucas_fornero [Lucas Fornero], Argentina, 11/May/2024. (E) Bernardo Ferraz, Brazil (MG). (F) Observation 137464405, by entomofobia [Juan Zara], Argentina, 16/Apr/2022. (G) Observation 167907017, by fredericosonntag [Frederico Acaz Sonntag], Brazil (PB), 17/Jun/2023. (H) Observation 147265948, by offvinn, Brazil (CE) 15/Jan/2023. (I) Observation 137450242, by fredericosonntag [Frederico Acaz Sonntag], Brazil (PB), 26/Jan/2000. (J) Observation 62552485, by alenilson [Alenilson], Brazil (AL), 19/Aug/2020.	FIGURE 2. Live specimens of Metaphotina bimaculata (A–F) and Metaphotina austri (G–J) retrieved from iNaturalist platform, license CC-BY-NC. Images may have been cropped and/or rotated for better observation. (A) Observation 163494503, by goncrisdi [GonzaloDias],Argentina, 06/May/2023.(B) Bernardo Ferraz, Brazil (MG).(C) Observation 199319301, by jcsullivan [John Sullivan], Bolívia, Jan/2024. (D) Observation 223539397, by lucas_fornero [Lucas Fornero], Argentina, 11/May/2024. (E) Bernardo Ferraz, Brazil (MG). (F) Observation 137464405, by entomofobia [Juan Zara], Argentina, 16/Apr/2022. (G) Observation 167907017, by fredericosonntag [Frederico Acaz Sonntag], Brazil (PB), 17/Jun/2023. (H) Observation 147265948, by offvinn, Brazil (CE) 15/Jan/2023. (I) Observation 137450242, by fredericosonntag [Frederico Acaz Sonntag], Brazil (PB), 26/Jan/2000. (J) Observation 62552485, by alenilson [Alenilson], Brazil (AL), 19/Aug/2020.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFED17743781A95EFAD3A893.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819081/files/figure.png	https://doi.org/10.5281/zenodo.15819081	FIGURE 3. Type material of Metaphotina spp. (A) True holotype of A. bimaculata described by Saussure (1870), deposited at MNHN. (B) Illustration of the holotype of A. brevipennis Saussure, 1872 and a reference to it from the precedent page: “Fig. 20. Acontista brevipennis, Sauss. Femelle de grandeur naturelle.” (C) A. bimaculata cited in Saussure & Zehntner (1894), mislabeled as “syntypus” (Pfäuti & Hollier 2012), deposited at MHNG. (D) A. bimaculata specimen cited in Saussure (1871) and mislabeled as “Type”, deposited at MNHN (adapted from Rivera & Svenson 2020). (E) Holotype of M. piracicabensis, deposited at ESALQ. (F) M. piracicabensis labeled (incorrectly) as “allotype”, deposited at ESALQ. (G) Holotype of A. rehni, deposited at ANSP. Scale: 10 mm.	FIGURE 3. Type material of Metaphotina spp. (A) True holotype of A. bimaculata described by Saussure (1870), deposited at MNHN. (B) Illustration of the holotype of A. brevipennis Saussure, 1872 and a reference to it from the precedent page: “Fig. 20. Acontista brevipennis, Sauss. Femelle de grandeur naturelle.” (C) A. bimaculata cited in Saussure & Zehntner (1894), mislabeled as “syntypus” (Pfäuti & Hollier 2012), deposited at MHNG. (D) A. bimaculata specimen cited in Saussure (1871) and mislabeled as “Type”, deposited at MNHN (adapted from Rivera & Svenson 2020). (E) Holotype of M. piracicabensis, deposited at ESALQ. (F) M. piracicabensis labeled (incorrectly) as “allotype”, deposited at ESALQ. (G) Holotype of A. rehni, deposited at ANSP. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFED17743781A95EFAD3A893.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819121/files/figure.png	https://doi.org/10.5281/zenodo.15819121	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819083/files/figure.png	https://doi.org/10.5281/zenodo.15819083	FIGURE 4. Metaphotina bimaculata. (A) Male holotype, deposited at MNHN. (B) Genitalia of holotype, slide deposited at MNHN. (C) Illustration of holotype’s genitalia. Scale: 10 mm.	FIGURE 4. Metaphotina bimaculata. (A) Male holotype, deposited at MNHN. (B) Genitalia of holotype, slide deposited at MNHN. (C) Illustration of holotype’s genitalia. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819085/files/figure.png	https://doi.org/10.5281/zenodo.15819085	FIGURE 5. Metaphotina bimaculata (Saussure, 1870). (A) Male, habitus (B) Female, habitus. (C) Head of male, frontal view. (D) Head of female, frontal view. (E) Head of female, vertex and ocellar projections. (F) Pronotum of male, dorsal view. (G) Pronotum of female, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm.	FIGURE 5. Metaphotina bimaculata (Saussure, 1870). (A) Male, habitus (B) Female, habitus. (C) Head of male, frontal view. (D) Head of female, frontal view. (E) Head of female, vertex and ocellar projections. (F) Pronotum of male, dorsal view. (G) Pronotum of female, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819087/files/figure.png	https://doi.org/10.5281/zenodo.15819087	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819089/files/figure.png	https://doi.org/10.5281/zenodo.15819089	FIGURE 7. (A) Habitus of a Metaphotina bimaculata protonymph. (B) Head, frontal view. (C) Head and prothorax, dorsal view. (D) Prothoracic leg’s external face, lateral view. (E) Enlargement of discoidal spines. *Damaged discoidal spine of a newly hatched individual, with only its insertion region remaining. Scales: 1 mm.	FIGURE 7. (A) Habitus of a Metaphotina bimaculata protonymph. (B) Head, frontal view. (C) Head and prothorax, dorsal view. (D) Prothoracic leg’s external face, lateral view. (E) Enlargement of discoidal spines. *Damaged discoidal spine of a newly hatched individual, with only its insertion region remaining. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819091/files/figure.png	https://doi.org/10.5281/zenodo.15819091	FIGURE 8. Ootheca of Metaphotina bimaculata. (A) Sagittal section, external view, external coating partially removed. (B) Sagittal section, internal view (mirrored), external coating partially removed. (C) Cross-section, upper region. (D) Cross-section, lower region. Scales: A–B, 10 mm; C–D, 1 mm.	FIGURE 8. Ootheca of Metaphotina bimaculata. (A) Sagittal section, external view, external coating partially removed. (B) Sagittal section, internal view (mirrored), external coating partially removed. (C) Cross-section, upper region. (D) Cross-section, lower region. Scales: A–B, 10 mm; C–D, 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819093/files/figure.png	https://doi.org/10.5281/zenodo.15819093	FIGURE 9. Variation in Metaphotina bimaculata male wings. (A) Aquidauana, Mato Grosso do Sul. (B) Aquidauana, Mato Grosso do Sul. (C) Serra do Espinhaço, Minas Gerais. (D) Serra do Caraça, Minas Gerais. (E) Salobra, Mato Grosso do Sul. (F) Mirador, Maranhão. (G) Salto, São Paulo. (H) Tibagi, Paraná. (I) Palmas, Paraná.	FIGURE 9. Variation in Metaphotina bimaculata male wings. (A) Aquidauana, Mato Grosso do Sul. (B) Aquidauana, Mato Grosso do Sul. (C) Serra do Espinhaço, Minas Gerais. (D) Serra do Caraça, Minas Gerais. (E) Salobra, Mato Grosso do Sul. (F) Mirador, Maranhão. (G) Salto, São Paulo. (H) Tibagi, Paraná. (I) Palmas, Paraná.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819095/files/figure.png	https://doi.org/10.5281/zenodo.15819095	FIGURE 10. Metaphotina bimaculata, variation in the ventral phallomere of males, letters match genitalia for Figs. 10 and 11. (A–I) Mato Grosso do Sul. (J–K) Mato Grosso. (L) Goiás. (M) Tocantins. (N-T) Minas Gerais, P.1* damaged during dissection. (U–W). São Paulo. (X) Rio Grande do Sul. (Y‒AC) Paraná. (AD) Uruguay. Not to scale.	FIGURE 10. Metaphotina bimaculata, variation in the ventral phallomere of males, letters match genitalia for Figs. 10 and 11. (A–I) Mato Grosso do Sul. (J–K) Mato Grosso. (L) Goiás. (M) Tocantins. (N-T) Minas Gerais, P.1* damaged during dissection. (U–W). São Paulo. (X) Rio Grande do Sul. (Y‒AC) Paraná. (AD) Uruguay. Not to scale.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819097/files/figure.png	https://doi.org/10.5281/zenodo.15819097	FIGURE 11. Metaphotina bimaculata, variation in the left phallomere of males, letters match genitalia for Figs. 10 and 11. (A-I) Mato Grosso do Sul. (J-K) Mato Grosso. (L) Goiás. (M) Tocantins. (N-T) Minas Gerais. (U-W) São Paulo. (X) Rio Grande do Sul. (Y-AC) Paraná. (AD) Uruguay. Not to scale.	FIGURE 11. Metaphotina bimaculata, variation in the left phallomere of males, letters match genitalia for Figs. 10 and 11. (A-I) Mato Grosso do Sul. (J-K) Mato Grosso. (L) Goiás. (M) Tocantins. (N-T) Minas Gerais. (U-W) São Paulo. (X) Rio Grande do Sul. (Y-AC) Paraná. (AD) Uruguay. Not to scale.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819099/files/figure.png	https://doi.org/10.5281/zenodo.15819099	FIGURE 12. Schematic variation in Metaphotina bimaculata female coloration. (A) Salto, São Paulo. (B) Salto, São Paulo. (C) Tibagi, Paraná. (D) Ponta Grossa, Paraná. (E) Salto, São Paulo. (F) Unknown locality. (G) Jaraguari, Mato Grosso do Sul. (H) San Juan, Bolívia. Coloration of live specimens typically matches B, E and H.	FIGURE 12. Schematic variation in Metaphotina bimaculata female coloration. (A) Salto, São Paulo. (B) Salto, São Paulo. (C) Tibagi, Paraná. (D) Ponta Grossa, Paraná. (E) Salto, São Paulo. (F) Unknown locality. (G) Jaraguari, Mato Grosso do Sul. (H) San Juan, Bolívia. Coloration of live specimens typically matches B, E and H.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819113/files/figure.png	https://doi.org/10.5281/zenodo.15819113	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819115/files/figure.png	https://doi.org/10.5281/zenodo.15819115	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819117/files/figure.png	https://doi.org/10.5281/zenodo.15819117	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819119/files/figure.png	https://doi.org/10.5281/zenodo.15819119	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819121/files/figure.png	https://doi.org/10.5281/zenodo.15819121	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFE017673781A8EEFD0BAEE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819081/files/figure.png	https://doi.org/10.5281/zenodo.15819081	FIGURE 3. Type material of Metaphotina spp. (A) True holotype of A. bimaculata described by Saussure (1870), deposited at MNHN. (B) Illustration of the holotype of A. brevipennis Saussure, 1872 and a reference to it from the precedent page: “Fig. 20. Acontista brevipennis, Sauss. Femelle de grandeur naturelle.” (C) A. bimaculata cited in Saussure & Zehntner (1894), mislabeled as “syntypus” (Pfäuti & Hollier 2012), deposited at MHNG. (D) A. bimaculata specimen cited in Saussure (1871) and mislabeled as “Type”, deposited at MNHN (adapted from Rivera & Svenson 2020). (E) Holotype of M. piracicabensis, deposited at ESALQ. (F) M. piracicabensis labeled (incorrectly) as “allotype”, deposited at ESALQ. (G) Holotype of A. rehni, deposited at ANSP. Scale: 10 mm.	FIGURE 3. Type material of Metaphotina spp. (A) True holotype of A. bimaculata described by Saussure (1870), deposited at MNHN. (B) Illustration of the holotype of A. brevipennis Saussure, 1872 and a reference to it from the precedent page: “Fig. 20. Acontista brevipennis, Sauss. Femelle de grandeur naturelle.” (C) A. bimaculata cited in Saussure & Zehntner (1894), mislabeled as “syntypus” (Pfäuti & Hollier 2012), deposited at MHNG. (D) A. bimaculata specimen cited in Saussure (1871) and mislabeled as “Type”, deposited at MNHN (adapted from Rivera & Svenson 2020). (E) Holotype of M. piracicabensis, deposited at ESALQ. (F) M. piracicabensis labeled (incorrectly) as “allotype”, deposited at ESALQ. (G) Holotype of A. rehni, deposited at ANSP. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819101/files/figure.png	https://doi.org/10.5281/zenodo.15819101	FIGURE 13. Metaphotina austri Ferraz et al. 2023. (A) Male holotype, habitus (B) Female allotype, habitus. (C) Head of male holotype, frontal view. (D) Head of female allotype, frontal view. (E) Head of female allotype, vertex and ocellar projections. (F) Pronotum of male holotype, dorsal view. (G) Pronotum of female allotype, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm. Figure adapted from Ferraz et al. (2023).	FIGURE 13. Metaphotina austri Ferraz et al. 2023. (A) Male holotype, habitus (B) Female allotype, habitus. (C) Head of male holotype, frontal view. (D) Head of female allotype, frontal view. (E) Head of female allotype, vertex and ocellar projections. (F) Pronotum of male holotype, dorsal view. (G) Pronotum of female allotype, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm. Figure adapted from Ferraz et al. (2023).	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819103/files/figure.png	https://doi.org/10.5281/zenodo.15819103	FIGURE 14. Metaphotina austri, male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm. Figure adapted from Ferraz et al. (2023).	FIGURE 14. Metaphotina austri, male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm. Figure adapted from Ferraz et al. (2023).	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819105/files/figure.png	https://doi.org/10.5281/zenodo.15819105	FIGURE 15. Metaphotina austri egg cases from iNaturalist records. All licenses CC-BY-NC. Images may have been cropped and/or rotated for better observation. (A) Observation 154869944, by biologocelio, Brazil (CE) 19/Mar/2023. (B) Observation 128305433, by biologocelio, Brazil (CE), 14/Jun/2022.	FIGURE 15. Metaphotina austri egg cases from iNaturalist records. All licenses CC-BY-NC. Images may have been cropped and/or rotated for better observation. (A) Observation 154869944, by biologocelio, Brazil (CE) 19/Mar/2023. (B) Observation 128305433, by biologocelio, Brazil (CE), 14/Jun/2022.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819113/files/figure.png	https://doi.org/10.5281/zenodo.15819113	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819115/files/figure.png	https://doi.org/10.5281/zenodo.15819115	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819117/files/figure.png	https://doi.org/10.5281/zenodo.15819117	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819119/files/figure.png	https://doi.org/10.5281/zenodo.15819119	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819121/files/figure.png	https://doi.org/10.5281/zenodo.15819121	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF217633781A987FDE1AE9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819087/files/figure.png	https://doi.org/10.5281/zenodo.15819087	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819107/files/figure.png	https://doi.org/10.5281/zenodo.15819107	FIGURE 16. Metaphotina akaraje sp. nov. (A) Male holotype, habitus (B) Female allotype, habitus. (C) Head of male holotype, frontal view.(D) Head of female allotype, frontal view. (E) Head of female allotype, vertex and ocellar projections.(F) Pronotum of male holotype, dorsal view. (G) Pronotum of female allotype, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm.	FIGURE 16. Metaphotina akaraje sp. nov. (A) Male holotype, habitus (B) Female allotype, habitus. (C) Head of male holotype, frontal view.(D) Head of female allotype, frontal view. (E) Head of female allotype, vertex and ocellar projections.(F) Pronotum of male holotype, dorsal view. (G) Pronotum of female allotype, dorsal view. Scale bars: (A–B) 10mm; (C–G) 1mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819109/files/figure.png	https://doi.org/10.5281/zenodo.15819109	FIGURE 17. Metaphotina akaraje sp. nov., male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	FIGURE 17. Metaphotina akaraje sp. nov., male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819111/files/figure.png	https://doi.org/10.5281/zenodo.15819111	FIGURE 18. Metaphotina akaraje sp. nov., variation in the associated ventral (upper row) and left (lower row) phallomeres. (A) Holotype. (B–C) Paratypes. Scale: 1 mm.	FIGURE 18. Metaphotina akaraje sp. nov., variation in the associated ventral (upper row) and left (lower row) phallomeres. (A) Holotype. (B–C) Paratypes. Scale: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819113/files/figure.png	https://doi.org/10.5281/zenodo.15819113	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819115/files/figure.png	https://doi.org/10.5281/zenodo.15819115	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819117/files/figure.png	https://doi.org/10.5281/zenodo.15819117	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819119/files/figure.png	https://doi.org/10.5281/zenodo.15819119	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819121/files/figure.png	https://doi.org/10.5281/zenodo.15819121	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	FIGURE 23. Metaphotina spp. habitus, dorsal view. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male holotype. (D) Metaphotina austri, female allotype. (E) Metaphotina akaraje sp. nov., male holotype. (F) Metaphotina akaraje sp. nov., female allotype. Scale: 10 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819087/files/figure.png	https://doi.org/10.5281/zenodo.15819087	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	FIGURE 6. Metaphotina bimaculata, male genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFF617593781AA6EFDB8AD83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819103/files/figure.png	https://doi.org/10.5281/zenodo.15819103	FIGURE 14. Metaphotina austri, male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm. Figure adapted from Ferraz et al. (2023).	FIGURE 14. Metaphotina austri, male holotype genitalia. (A) Left phallomere, dorsal view. (B) Ventral phallomere, dorsal view. (C) Right phallomere, ventral view. (D) Dorsal view. (E) Lateral view. Scales: 1 mm. Figure adapted from Ferraz et al. (2023).	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819113/files/figure.png	https://doi.org/10.5281/zenodo.15819113	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	FIGURE 19. Heads of Metaphotina spp. (A) M. bimaculata, male. (B) Holotype of M. austri, male. (C) Holotype of M. akaraje sp. nov., male. (D) M. bimaculata, female. (E) Allotype of M. austri, female. ((F) Allotype of M. akaraje sp. nov., female. (G) Close-up of female M. bimaculata vertex. (H) Close-up of female (allotype) M. austri vertex. (I) Close-up of female (allotype) M. akaraje sp. nov. vertex. Scale: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819115/files/figure.png	https://doi.org/10.5281/zenodo.15819115	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	FIGURE 20. Genitalia of male Metaphotina spp. (A) M. bimaculata. (B) M. austri. (C) M. akaraje sp. nov.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819117/files/figure.png	https://doi.org/10.5281/zenodo.15819117	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	FIGURE 21. Female genitalia of female Metaphotina spp. (A) Metaphotina bimaculata, ventral view (note cxdl distinguishably sclerotized). (B) Metaphotina austri, ventral view. (C) Metaphotina akaraje sp. nov., ventral view. (D) Metaphotina bimaculata, lateral view. (E) Metaphotina austri, lateral view. (F) Metaphotina akaraje sp. nov., lateral view. (G) Metaphotina bimaculata CG8, close-up. (H) Metaphotina austri CG8, close-up. (I) Metaphotina akaraje sp. nov. CG8, close-up. Scales: 1 mm.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819123/files/figure.png	https://doi.org/10.5281/zenodo.15819123	FIGURE 24. Known distribution of Metaphotina bimaculata in South America.	FIGURE 24. Known distribution of Metaphotina bimaculata in South America.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819127/files/figure.png	https://doi.org/10.5281/zenodo.15819127	FIGURE 26. Distribution of Metaphotina spp. across SouthAmerican biomes.The pie charts illustrate the relative distribution of specimens analyzed per species per biome, with the total number of specimens indicated in parentheses.All shown records were derived from specimen label data, in addition to verified records of M. austri on iNaturalist. Biome and ecoregion delimitations adapted from Olson et al. (2001) and Morrone et al. (2022).	FIGURE 26. Distribution of Metaphotina spp. across SouthAmerican biomes.The pie charts illustrate the relative distribution of specimens analyzed per species per biome, with the total number of specimens indicated in parentheses.All shown records were derived from specimen label data, in addition to verified records of M. austri on iNaturalist. Biome and ecoregion delimitations adapted from Olson et al. (2001) and Morrone et al. (2022).	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819125/files/figure.png	https://doi.org/10.5281/zenodo.15819125	FIGURE 25. Known distribution of Metaphotina austri and Metaphotina akaraje sp. nov. in Northeastern Brazil. Yellow star and triangle refers to species’ type localities.	FIGURE 25. Known distribution of Metaphotina austri and Metaphotina akaraje sp. nov. in Northeastern Brazil. Yellow star and triangle refers to species’ type localities.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
5B308C1FFFCC17593781AB66FA5DAAC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15819119/files/figure.png	https://doi.org/10.5281/zenodo.15819119	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	FIGURE 22. Wings of Metaphotina spp. (A) Metaphotina bimaculata, male. (B) Metaphotina bimaculata, female. (C) Metaphotina austri, male. (D) Metaphotina austri, female. (E) Metaphotina akaraje sp. nov., male. (F) Metaphotina akaraje sp. nov., female.	2025-06-10	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio		Zenodo	biologists	Ferraz, Bernardo R.;Queiroga, Drielly;Souza-Dias, Pedro G. B.;Rivera, Julio			
