identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6645E0166129FFB7A0F14A1C4E31B5D8.text	6645E0166129FFB7A0F14A1C4E31B5D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myiomma Puton 1872	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Myiomma Puton, 1872</p>
            <p> Myiomma Puton, 1872: 177 (gen. nov.); Schuh, 1995: 12 (catalog); Kerzhner &amp; Josifov, 1999: 6 (catalog); Schuh, 2002 –2013: (web catalog); Liu et al., 2022: 453 (diagnosis, key). Type species:  Myiomma fieberi Puton, 1872 , monotypic. </p>
            <p> Heidemannia Uhler, 1891: 119 (gen. nov.). (syn. by McAtee &amp; Malloch, 1932: 64). Type species: Type species:  Myiomma cixiiforme Uhler, 1891 . </p>
            <p> Paramyiomma Carvalho, 1951: 381 (gen. nov.). (syn. by Smith, 1967: 41). Type species:  Paramyiomma landsburyi Carvalho, 1951 . </p>
            <p> Diagnosis.  Myiomma taxa can be recognized by the following characters: Body length 1.8–3.2, elongate oval; body coloration usually pale brown to blackish brown, often male and female possesses different coloration; dorsum weakly shining, usually covered with pale to dark brown simple setae; compound eyes large, sometimes adjacent to each other; vertex narrow, with a pair of distinct, round ocelli; antennal segment I short and stout, segment II thick, elongated and weakly clavate in male, segment III and IV short and thin; pronotum trapezoidal, anterior margin linear, posterior margin somewhat round, W-shaped; mesoscutum distinct, laterally wide; foreleg and midleg thin and linear, hindleg with thickened metafemur; male endosoma membranous, weakly sclerotized; left paramere with moderately swollen sensory lobe and elongated, apically tapering hypophysis; right paramere small, horn-shaped; female genitalia with weakly sclerotized rings and interramal structures. For more diagnostic characters and figures, see Yasunaga et al. (2017) and Liu et al. (2022). </p>
            <p>Distribution. Afrotropical, Australasia, Nearctic, Neotropical, Oriental, and Palearctic region (Schuh, 2002 – 2013; Yasunaga et al., 2017)</p>
            <p> Biology. Members of genus  Myiomma can be found on tree bark of diverse types of deciduous trees (Yasunaga &amp; Hayashi, 2002). This group can be collected with various insect trap, such as funnel traps, light traps, and malaise traps (Henry &amp; Carpintero, 2012; Yasunaga et al., 2017). Based on observational data, species of  Myiomma are presumed to be predaceous (Wheeler &amp; Henry, 1978; Henry &amp; Carpintero, 2012; Shishido et al., 2020; Yeshwanth et al., 2021). </p>
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	https://treatment.plazi.org/id/6645E0166129FFB7A0F14A1C4E31B5D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Oh, Minsuk;Kim, Wonwoong;Kim, Wongun;Lee, Seunghwan	Oh, Minsuk, Kim, Wonwoong, Kim, Wongun, Lee, Seunghwan (2025): A new species of the genus Myiomma (Heteroptera: Miridae: Isometopinae) from the Korean Peninsula. Zootaxa 5566 (1): 145-158, DOI: 10.11646/zootaxa.5566.1.6, URL: https://doi.org/10.11646/zootaxa.5566.1.6
6645E0166129FFB2A0F14E404A8AB3B4.text	6645E0166129FFB2A0F14E404A8AB3B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myiomma koreana Oh & Kim & Kim & Lee 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Myiomma koreana Oh &amp; Lee ,  sp. nov.</p>
            <p>(Figs. 1–8)</p>
            <p> Myiomma kukai Jung &amp; Lee, 2012: 53 (misidentification of  M. koreana ). </p>
            <p> Diagnosis. This species can be distinguished from its closely allied species,  M. takahashii and  M. kukai by the combination of following characteristics: dorsum covered with brownish, reclining setae; antennal segment I widely pale brown in male, chestnut brown in female; antennal segment II pale brown except apical 1/ 3 in male, widely dark brown and base and apex narrowly pale in female; hemelytra without reddish tinge, semitransparent and brownish in male and widely dark brown in female; cuneus dark brown, anterior 1/3 pale; all male coxae widely dark brown, all femora dark brown with reddish tinge medially, apically pale; all female coxae and femora widely dark brown, apex of metafemur narrowly pale; inner margin of left paramere sensory lobe with comparatively weak protrusion, outer margin wide and covered with stiff setae. </p>
            <p>Description. Male. Body elongated oval, 2.10–2.20; Coloration: Body brown to dark brown, partly tinged with red; Head dark brown, posterior margin yellowish white, posterolateral margin partly reddish brown; vertex and frons partly pale brown along inner margin of compound eye; clypeus tinged with red. Antennae brown; antennal segment I widely pale brown, segment II pale brown, apical 1/3 darkened, segment III and IV brown, apically pale. Labium dark chestnut brown, segment III medially pale. Pronotum entirely dark brown, lateral side partly brown. Mesoscutum dark brown, lateral side widely pale, tinged with orange-red coloration. Scutellum dark brown, apical 1/3 whitish, extreme apex blackish brown. Hemelytra brown, semi-transparent, basal part of corium narrowly pale, with minute reddish tinge; cuneus brown basal 1/3 pale; membrane pale smoky brown, partly darkened along inner side of vein. Legs brownish; coxae dark brown, femora widely dark brown and medially tinged with red; metafemur dorsally pale brown and ventrally dark brown with reddish tinge, apical 1/4 pale; metatibia brown, apically pale; tibial spines pale brown; tarsi brown. Abdomen dark brown, pygophore apically reddish brown. Surface and vestiture: Dorsal surface weakly shiny, covered with pale brown to dark brown simple setae. Head weakly glabrous, frons sparsely covered with reclining pale brown setae; posterolateral margin with stiff, pale brown setae; Antennal segment II covered with suberect, brown setae. Pronotum and scutellum covered with brown, reclining setae; posteromedial margin with long, reclining brown setae. Hemelytra weakly shiny, densely covered with brown, suberect simple setae. Femora sparsely covered with short, pale setae and suberect, simple setae. Structure: Head dorsoventrally elongate, waterdrop-shaped in anterior view; posterior margin of vertex V-shaped. Antennal segment I very short and incrassate, segment II thick, elongated and weakly clavate, segment III and IV short and thin. Labium long, reaching abdominal sterna V–VII. Foreleg and midleg slender, hindleg with thick metafemur and elongated, linear metatibia. Male genitalia as in Figs. 4–6; left paramere L-shaped; sensory lobe tumid, inner margin with weak protrusion and outer margin round, sparsely covered with long, stiff setae; Hypophysis smooth and elongated, distinctly curved medially; right paramere small, weakly curved and horn-shaped; basal part of sensory lobe with a minute protrusion; endosoma membranous, with a thin, elongated sclerotized structure; phallotheca conical, tapering apically and inner margin well sclerotized. Female. Body elongate oval, 2.10–2.30; body comparatively ovoid, overall coloration darker than in male; antennal segment I and II dark brown, segment III and IV brown; segment II not clavate, pale at extreme apex; apical half of scutellum with wide, V-shaped white marking; hemelytra dark brown, femora almost entirely dark brown; metafemur dark brown, apical 1/4 pale, mottled with dark brown and reddish tinge; basal 2/3 of metatibia dark brown. Not significantly different from male in surface and vestiture. Female genitalia as in Fig. 7; sclerotized rings small and mutually adjacent, weakly sclerotized and ovate, lateral margin pointed at apex; dorsal labiate plate weakly sclerotized, surface somewhat wrinkled; ventral side of bursa copulatrix with a pair of waterdrop-shaped ventral structures; interramal lobe and sclerite weakly sclerotized, surface rugose; gonapophysis I leaflike, ventral margin moderately curved; gonapophysis II smooth, apical margin similar as spoon-style ship bow.</p>
            <p>Measurements. Male (n=2)/ Female (n=5). Total body length 2.14–2.17/ 2.16–2.29; head width across eyes 0.42–0.44/ 0.42–0.44; vertex width 0.08–0.09/ 0.08–0.09; lengths of antennal segment I–IV 0.08, 0.64–0.67, 0.10– 0.11, 0.09–0.11/ 0.09–0.10, 0.50–0.52, 0.17–0.18, 0.13–0.14; labial length 1.00/ 0.88–1.02; mesal pronotal length including collar 0.30–0.33/ 0.30–0.32; basal pronotal width 0.83/ 0.83–0.90; width across hemelytron 0.98–1.01/ 0.97–1.10; cuneal length 0.36–0.39/ 0.36–0.38; cuneal width 0.23–0.24/ 0.25–0.27; lengths of metafemur, tibia and tarsus 0.66, 0.88, 0.22/ 0.66–0.73, 0.94–0.97, 0.22–0.23.</p>
            <p>Etymology. Named after the type locality, the Korean Peninsula; an adjective.</p>
            <p>Distribution. Korea (Gyeonggi-do; Gyeongsangbuk-do; Jeollabuk-do).</p>
            <p> Biology. This species was collected from the tree bark of  Quercus palustris Münchh. (  Fagaceae ),  Zelkova serrata (Thunb.) Makino (  Ulmaceae ), and  Salix sp. (  Salicaceae ). In Suwon-si and Gwacheon-si, this species coinhabited the bark surface with  I. amurensis , another isometopine species commonly found in deciduous tree bark in Korea. Observed specimens constantly probed the bark surface with its labium stretched forward (Figs. 1C, D), and rapidly scuttle away or jump when disturbed. </p>
            <p> Type material.   Holotype, ♂: South Korea, Gyeongsangbuk–do: Mt. Gaya,  Seongju-gun , from light trap, 06.viii.2022, WG Kim (SNU)  .   Paratypes: South Korea, Gyeonggi–do: 4♀ Gwanggyo park, Gwanggyosan-ro 159,  Jangan–gu ,  Suwon–si , from  Quercus palustris , 26.vi.2024, M. S. Oh &amp; W. Kim (SNU)  ;   1♀, ditto, from  Salix sp. , 26.vi.2024, M. S. Oh &amp; W. Kim (SNU)  ;   1♀,  Empty lot near Gwacheon civic center ,  Tongyeong-ro 5,  Gwacheon-si , from  Zelkova serrata , 28.vi.2024, W. Kim (SNU)  ;   1♀,  Garaeul park , Neunggok-dong 793,  Siheung-si , from  Quercus palustris , 28.vi.2024, W. Kim (SNU)  . </p>
            <p>Additional material. 1♂: South Korea, collection information unknown (SNU).</p>
            <p> Discussion. Previously, Jung &amp; Lee (2012) added molecular data of  Myiomma kukai Yasunaga &amp; Hayashi, 2002 , into their phylogenetic analysis dataset. While preparing the manuscript for  Myiomma koreana , we found the voucher specimen which was suspected to be the reference specimen of this sequence at the insect collection at Seoul National University CALS. Despite that this specimen did not bearing a collection label and the exact collection information was not available, we were able to examine it in morphological detail. Although Jung &amp; Lee (2012) did not mention the collection records for each of the analyzed species, most specimens deposited at SNU CALS were collected from South Korea by researchers at Prof. Seunghwan Lee’s lab between the late 2000s and the early 2010s, and this  Myiomma sample is also presumed to have been collected in South Korea. Based on the comparison of the external morphology and genital characters, this specimen is conspecific with our new species,  Myiomma koreana . Pairwise distance between each sequence is less than 0.2% (Table 2), which can be regarded as intraspecific variation based on former barcoding research on  Miridae (Jung et al., 2011; Kim &amp; Jung, 2018). NJ tree results also show a close relationship between ‘  Myiomma kukai ’ from Jung &amp; Lee (2012) and our new species (Fig. 9).  M. kukai is endemic to Japan (Honshu and Kyushu) (Yasunaga 2001, Yasunaga et al. 2017, Shishido et al. 2020), and has never been reported from Korea. We therefore clarify that the ‘  M. kukai ’ sequence provided by Jung &amp; Lee (2012) pertains to our new species,  M. koreana , not  M. kukai from Japan. </p>
            <p> Furthermore, all specimens that were hand-collected by the first and second author from tree bark were female, while a single male was obtained by light trap by the third author. Species of  Myiomma exhibit sexual dimorphism regarding eye size and dorsal coloration (Yasunaga et al. 2017, Sishido et al. 2020), making it hard to properly link males and females by traditional morphological means, except when the male and female are collected at the same locality have been treated as conspecific. With the help of DNA barcoding on the COI region, we were able to confirm that the two morphologically different specimens of the new species represent sexual dimorphism. </p>
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	https://treatment.plazi.org/id/6645E0166129FFB2A0F14E404A8AB3B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Oh, Minsuk;Kim, Wonwoong;Kim, Wongun;Lee, Seunghwan	Oh, Minsuk, Kim, Wonwoong, Kim, Wongun, Lee, Seunghwan (2025): A new species of the genus Myiomma (Heteroptera: Miridae: Isometopinae) from the Korean Peninsula. Zootaxa 5566 (1): 145-158, DOI: 10.11646/zootaxa.5566.1.6, URL: https://doi.org/10.11646/zootaxa.5566.1.6
6645E0166121FFBFA0F14F384A8BB6B1.text	6645E0166121FFBFA0F14F384A8BB6B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myiomma Puton 1872	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to species of East Asian  Myiomma</p>
            <p> 1. Pronotum entirely brown to black, without a pale spot (lateral margin reddish brown in  M. zhengi ).................... 2 </p>
            <p> - Pronotum with a pale spot, lateral margin with pale portion (often entirely dark brown in  M. kukai male)............... 8 </p>
            <p>2. Inner margin of compound eye conjoined.................................................................. 3</p>
            <p>- Inner margin of compound eye narrowly separated from each other............................................. 5</p>
            <p> 3. Body large, about 2.70 mm .......................................................................  M. altica</p>
            <p>- Body comparatively small, about 2.1–2.4 mm .............................................................. 4</p>
            <p> 4. Male procoxa and all femora pale brown. Female with mostly pale hemelytra, partly darkened at corial margin along clavus and inner-apical margin. Inner margin of left paramere sensory lobe with distinct protrusion, outer margin smooth.....................................................................................................  M. takahashii</p>
            <p> - Male coxae mostly dark brown and femora dark brown with a reddish tinge and pale apices. Female with blackish brown hemelytra. Inner margin of left paramere sensory lobe with weak protrusion, outer margin angulate with border of hypophysis....................................................................................  M. koreana sp. nov.</p>
            <p> 5. Basal half of hemelytra with distinct entirely pale portion...............................................  M. choui</p>
            <p>- Hemelytra widely dark brown, often with narrow, pale markings at base.......................................... 6</p>
            <p> 6. Cuneus with a pale or whitish band basally......................................................  M. ussuriensis</p>
            <p>- Cuneus not basally pale or whitish, unicolorous............................................................. 7</p>
            <p> 7. Labium reaching posterior margin of abdominal sternum I.........................................  M. qinlingensis</p>
            <p> - Labium reaching abdominal sternum IV............................................................  M. zhengi</p>
            <p>8. Basal half of metafemur darkened........................................................................ 9</p>
            <p>- Basal half of metafemur pale yellow to pale brown......................................................... 10</p>
            <p> 9. Hypophysis of left paramere with pointed, beak-shaped apex............................................  M. kukai</p>
            <p> - Hypophysis of left paramere with somewhat blunt apex, thickness almost uniform......................  M. samuelsoni</p>
            <p> 10. Apical half of metafemur with a brown to dark brown spot.........................................  M. kentingense</p>
            <p>- Apical half of metafemur without a distinct spot........................................................... 11</p>
            <p> 11. Mesoscutum widely dark, lateral margin narrowly pale. At least anterior 1/3 of pronotum blackish........  M. austroccidens</p>
            <p> - Mesoscutum medially dark, laterally tinged with orange. Pronotum pale, partly darkened along callus.........  M. minutum</p>
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	https://treatment.plazi.org/id/6645E0166121FFBFA0F14F384A8BB6B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Oh, Minsuk;Kim, Wonwoong;Kim, Wongun;Lee, Seunghwan	Oh, Minsuk, Kim, Wonwoong, Kim, Wongun, Lee, Seunghwan (2025): A new species of the genus Myiomma (Heteroptera: Miridae: Isometopinae) from the Korean Peninsula. Zootaxa 5566 (1): 145-158, DOI: 10.11646/zootaxa.5566.1.6, URL: https://doi.org/10.11646/zootaxa.5566.1.6
