taxonID	type	description	language	source
4048BB840C9151E0AD10CF28F9F6B0C4.taxon	description	Figs 1, 2	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
4048BB840C9151E0AD10CF28F9F6B0C4.taxon	etymology	Etymology. Nwarr ngaal is Burmese and means “ little cow ”. It refers to the conspicuous color markings on the species’ head and pronotum.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
4048BB840C9151E0AD10CF28F9F6B0C4.taxon	diagnosis	Diagnosis of species. As for genus (monotypic).	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
4048BB840C9151E0AD10CF28F9F6B0C4.taxon	description	Description. A fully preserved specimen, except for left metathoracic leg. Measurements: Values given with a “ ca. ” indicate measurements based partly on estimation. Body-length ca. 1.83 mm. Head height 0.68 mm. Compound eye height 0.31 mm, width 0.21 mm. Antenna length 0.85 mm. Pronotum length 0.54 mm, height 0.4 mm. Forewing length 0.55 mm, hindwing length 1.64 mm. Prothoracic leg: femur 0.38 mm long. Tibia 0.36 mm long. Mesothoracic leg: tibia 0.5 mm long. Femur / Tibia width = 1.07 (at mid of both). Metathoracic leg: femur ca. 0.86 mm long, 0.51 mm wide. Tibia ca. 0.79 mm long, ca. 0.09 mm wide (around mid-length). Tarsus> 0.31 mm long. Cercus 0.15 mm long. Paraproctal lobe ca. 0.05 mm long. Subgenital plate 0.18 mm long. Head: Antennae 10 - segmented, moniliform. Pedicel and scape wider than antennomeres. Head with two small bumps on the vertex in front of the compound eyes. Compound eyes placed laterally on the head. Head with distinct coloration; a lighter area crosses the eye dividing it into two equal parts and continuing on the dorso-lateral portion of the head. Ocelli not seen. Thorax: Pronotum with similar light / dark contrasting colors as head, but here the base color is light and several dark spots are scattered across the surface. Pronotum without elevation. Both fore- and hindwings present. Forewings with same color pattern as pronotum, hindwings uniformly dark. Hindwing slightly longer than abdomen. Legs: Prothoracic leg: femur bearing some long hairs. Tibia slightly inflated, completely covered with shorter to longer hairs. Apically with three dactyls, the longest of which is 0.05 mm long. Tarsus slender, only distal segment can be seen, bearing two curved claws. Mesothoracic leg: femur with some short hairs on both margins. Femur rather slender towards coxa, becoming wider towards tibia. Tibia slightly inflated, but not as wide as distal part of femur. Tibia apically with dense long hair. Tarsus slender with two claws. Metathoracic leg: femur inflated along its entire length. Tiger stripe coloration, which can be seen on the metafemur of the specimen is probably due to fissures in the amber and not actual coloration. Connection to tibia damaged. Tibia with lateral carina that runs until apex. Fin with distinct serration. Subapical and apical spurs present, approximately the same length. Tarsus proportionally long and rather thin. Ventrally with eight to nine hairs. Apex of tarsus hidden behind mesotibia. Abdomen: Epiproct short, not covering cercus. Cercus one-segmented, cylindrical in shape and with many long hairs. Paraproct distinct with paraproctal lobe in ventral position. Paraproctal lobe significantly smaller than cercus (both length and width), cylindrical without hairs. Subgenital plate elongated, approximately the length of the cercus and with long hairs; one especially long hair on the apex. Subgenital plate mostly dark in coloration becoming slightly lighter towards apex. Cercus completely of lighter color.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
21416810FE7B53629B35E97A025B6C5A.taxon	type_taxon	Type species. Abaddonella nwarrngaal sp. nov.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
21416810FE7B53629B35E97A025B6C5A.taxon	etymology	Etymology. Named after Abaddon, angel of the abyss from the Book of Revelation in connection with - ella meant as a suffix functioning as diminutive. A translation would be “ little abaddon ” or “ little demon ”, referring to the small bumps on the type species’ vertex which remind a little of demon horns.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
21416810FE7B53629B35E97A025B6C5A.taxon	diagnosis	Diagnosis. Antenna 10 - segmented. Compound eyes laterally placed on head. Two small bumps on vertex in front of the compound eyes. Pronotum more or less quadrangular in shape. Forewings and hindwings present. Hindwings slightly longer than abdomen. Protibia with hairs on entire surface. Apically with three strong dactyls. Mesotibia not distinctly wider than mesofemur. Metafemur inflated along entire length. Metatibial fin with distinct serration. Subapical and apical spurs approximately of equal length, much shorter than metatarsus. Metatarsus with rather long fine hairs on ventral margin. Abdomen with cercus one-segmented and heavily setulose. Paraproctal lobes much shorter than cercus, inconspicuous and not setulose. Subgenital plate elongated, much longer than paraproctal lobes and with long hairs. Coloration on head with distinct light / dark contrast over the eye. Pronotum with light coloration and several dark colored spots.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
D0B2BC8FC1625FD196CAFAD5AD40AD80.taxon	description	Figs 3, 4, 5	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
D0B2BC8FC1625FD196CAFAD5AD40AD80.taxon	etymology	Etymology. Named after the latin word “ pinna ” (fin) because of the ventral appendages on the abdomen, reminding of the dorsal fin of some fish, for example male sea goldies or the threadfin butterflyfish.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
D0B2BC8FC1625FD196CAFAD5AD40AD80.taxon	diagnosis	Diagnosis of species. As for genus (monotypic).	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
D0B2BC8FC1625FD196CAFAD5AD40AD80.taxon	description	Description. The specimen is well preserved, but most of the head and thorax are somewhat contorted due to internal damage of the amber structure. Measurements: Body-length 2.52 mm (head to abdominal apex). Pronotum length 0.62 mm, height 0.38 mm. Prothoracic leg: femur ca. 0.35 mm long. Tibia ca. 0.27 mm long. Tarsus ca. 0.25 mm long (including claws). Forewing ca. 0.79 mm long. Hindwing ca. 1.53 mm long. Mesothoracic leg: tibia 0.39 mm long, 0.25 mm wide (max width). Tarsus 0.35 mm long. Metathoracic leg: femur 1.36 mm long. Tibia 1.41 mm long, 0.09 mm wide. Tarsus 0.32 mm long. Subapical spurs ca. 0.05 mm long. Apical spurs 0.08 mm long. Cercus 0.19 mm long. Paraproctal lobes 0.3 mm long (not including hairs). Ovipositor (longest valve) 0.17 mm long. Head: Morphology uncertain due to internal damage in the amber. Thorax: Pronotum without elevation. Very deep laterally, covering the entire height of the thorax. Fore- and hindwings present. Venation undiscernible (lines seen on the wings are fissures in the amber, not actual veins). Hindwings just as long as abdomen. Legs: Prothoracic leg: Coxa big, about 2.5 × the width of femur with sparse hair. Trochanter halfway in width between coxa and femur. Femur with numerous hairs. Tibia with numerous hairs, too. Apically with four dactyls, the longest of which is 0.05 mm long. Tarsus damaged. Slender without hair and ending in two claws. Mesothoracic leg: only tibia and tarsus can be seen. Tibia with very distinct shape, looking like it is made up of two triangles, where the distal half is inserted into the bigger (both wider and thicker) proximal half. Greatly inflated. Apically with some hairs. Tarsus two-segmented, slender, no hair and with two claws. Metathoracic leg: femur apically with sparse hair. Tapering toward apex, only somewhat inflated. Tibial fin with serration. A pair of subapical and apical spurs. Apical spurs with four conspicuous hairs on ventral margin. Tarsus blunt with two rows ventrally, one metatibial fin serrated, the other with hair. Abdomen: Proximal to abdominal apex are three fin-like appendages of which the anterior two have singe hair inserted. Posterior fin probably lost the hair during preservation. More fins may be present but are covered by the metafemur. Cercus one-segmented, cylindrical in shape and setulose. Paraproctal lobes two-segmented, longer than cercus. Distal segment swollen, clavate in shape with distinct long hairs, especially on the apex of distal segment. Ovipositor visible, with narrower pointed end.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
F6875E61AC1D578D9E1A3AD734FB1A0D.taxon	type_taxon	Type species. Naginipteryx pinna sp. nov.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
F6875E61AC1D578D9E1A3AD734FB1A0D.taxon	etymology	Etymology. Named after the naga (female nagini), a mythological creature from Asian religions such as Hinduism and Buddhism. It is also found in the mythology of Myanmar. Nagas are usually portrayed as half-human, half serpent beings, but can also be found as dragons. This is why the name has been chosen, for the triangularly shaped mesotibia of the genus, reminding of a classical dragons’ tail.	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
F6875E61AC1D578D9E1A3AD734FB1A0D.taxon	diagnosis	Diagnosis. Protibia with four dactyls. Fore- and hindwings present, hindwings just as long as abdomen. Apical metatibial spurs slightly longer than subapical. Ventral margin of apical spurs with four distinct long hairs. External genitalia very similar to Yakkhapipteryx Schall, Cao & Husemann, 2025 more specifically Y. gracilis (Gu, Zheng, Cao & Yue, 2022) (this includes morphology of the cerci, paraproctal lobes and ovipositor valves): cercus one-segmented, cylindrical and setulose with long hairs. Paraproctal lobes two-segmented and longer than cerci, apically swollen and clavate with distinct long hair. Ovipositor visible, approximately as long as cerci, valve rather thick (more similar to Y. gracilis than Y. mira (Gu, Zheng, Cao & Yue, 2022 )). Differs from Yakkhapipteryx by 1) the mesotibia being distinctly inflated and triangular in shape. 2) Ventral appendages with an inserted hair on the abdomen. 3) A blunt (rather than pointed) metatarsus with one row of serration and one row with hairs (rather than no row).	en	Schall, Ole-Kristian Odin, Heads, Sam W., Husemann, Martin (2025): New species of Tridactyloidea (Orthoptera, Caelifera) from mid-Cretaceous Burmese amber show an intermediate evolutionary state between extant lineages. Evolutionary Systematics 9 (2): 173-181, DOI: 10.3897/evolsyst.9.154534
