taxonID	type	description	language	source
73560112CE39A928C6F25BE1B7E6FCE7.taxon	type_taxon	Type species: Tegenaria argentata Doleschall, 1857, from Indonesia.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A928C6F25BE1B7E6FCE7.taxon	diagnosis	Diagnosis. For description and diagnosis of the genus see Bayer (2012).	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	description	Figs 1 – 3, 8	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	materials_examined	Type material. Holotype ♀ (NZC-ZSI- 9428 / 18) from INDIA: Nagaland: Phek District, Chizami (25 ° 35 ’ 49 ” N 94 ° 23 ’ 09 ” E), 1400 m a. s. l., 02. III. 2024, leg. P. P. Sudhin.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	etymology	Etymology. The species is named after its type locality, Chizami, a small village in Nagaland, North East India. The name is treated as a noun in apposition.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	diagnosis	Diagnosis. Psechrus chizami sp. nov. can be easily distinguished from those of all other Psechrus species by its unique epigynal morphology: epigyne possesses a broad median septum (MS), with its anterio-central section forming an atrium bordered by a sclerotized rim, giving it an ear-muff-like appearance; copulatory ducts long, forming a loop before entering the spermathecae posteriorly; spermathecae rounded (Figs 1 C – F, 2, 3).	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	description	Description. Female. (Holotype, NZC-ZSI- 9431 / 18) (Figs 1 – 3) Measurements: body length 20.92; carapace length 9.82, width 6.67, anterior width of carapace 4.28; opisthosoma length 11.10, width 6.04. Eye diameters: AME 0.40, ALE 0.51, PME 0.59, PLE 0.52. Eye interdistances: AME – AME 0.24, AME – ALE 0.07, ALE – ALE 1.02, PLE – PLE 1.84, PME – PME 0.24, PME – PLE 0.36, AME – PME 0.54, ALE – PLE 0.46. Clypeus height at AME 0.81, clypeus height at ALE 0.63. Chelicerae 3.12 long, with 2 promarginal and 4 retromarginal teeth. Measurement of palp and legs: Palp: 10.73 [4.13, 1.45, 1.96, 3.19], legs I 55.82 [14.68, 3.99, 16.42, 14.08, 6.65], II 45.1 [12.38, 3.20, 12.86, 11.08, 5.58], III 30.18 [9.46, 2.16, 7.56, 7.28, 3.27], IV 45.43 [13.54, 3.18, 12.09, 11.24, 5.38]. Leg formula: 1423. Spination. Palp: femur pl 1 rl 1 do 3, patella pl 1 do 1, tibia pl 2 do 1, tarsus pl 2 rl 2 plv 1. Legs: femur I – II pl 5 rl 6 do 5, III – IV pl 5 rl 5 do 5; tibia I – II pl 3 rl 3 plv 4 rlv 4, III pl 3 rl 3 plv 2 rlv 2, IV pl 3 rl 3 plv 3 rlv 3; metatarsus I – II pl 1 rl 1 plv 4 rlv 4 v 1, III – IV pl 2 rl 2 plv 3 rlv 3 v 1; tarsus I – IV spineless. Carapace pale yellow, covered with black setae, with a broad light brown median band, and narrow light brown lateral bands; median band with a broad pale-yellow marking anteriorly, and triangle-shaped pale-yellow marking posteriorly, sides of median band undulating; lateral bands extending from behind PLE (Fig. 1 A). Eye field light reddish-yellow to brown, covered with setae (Fig. 1 A). Clypeus pale yellow, with light brown sides. Chelicerae dark reddish-brown, covered with setae (Fig. 1 B). Endites light reddish-brown, scopulate, with pale yellow inner margins; sides covered with long black setae (Fig. 1 B). Labium light reddish-brown, tip with pale yellow margin, covered with black setae (Fig. 1 B). Sternum light brown, covered with long erect black setae, and short black setae; sides yellow, covered with short white setae (Fig. 1 B). Legs pale yellow, reddish-brown towards the distal segments. Opisthosoma dorsally pale yellow to greyish-brown, covered with light brown setae, anteriorly with median spindle-shaped pale yellow marking; laterally with dark-brown and white longitudinal stripes (Fig. 1 A). Venter light greyish-brown, medially with pale-yellow longitudinal stripe, extending the entire length of opisthosoma, more prominent anteriorly (Fig. 1 B). Spinnerets grey to greyish-brown (Fig. 1 B). Genitalia as shown in Figs 1 C – F, 2, 3. Epigynal field wider than long; median septum (MS) wider than long, concave with several wrinkles, and its anterio-central section forming an atrium (A) bordered by a sclerotized rim, giving it an ear-muff-like appearance (Figs 1 C, E, 2 A, 3 A); finger-like fertilization ducts visible laterally on the posterior borderline of the epigyne (Figs 1 C, E, 3 A); copulatory openings (CO) small, widely separated (Figs 1 C, E, 3 A); copulatory ducts (CD) long, forming a loop before entering the spermathecae posteriorly (Figs 1 D, F, 2 B – H, 3 B); spermathecae (S) small, rounded (Figs 1 D, F, 2 C – H, 3 B); fertilization ducts (FD) long, finger-like, and inwardly oriented (Figs 1 D, F, 2 B, G, 3 B). Male. Unknown. Note. Based on the unique structure of its female genitalia (see diagnosis), this new species cannot be assigned in any currently known species group.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE39A92AC6F25AD9B039F93A.taxon	distribution	Distribution. Known only from the type locality in India (Nagaland) (Fig. 8).	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	description	Figs 4 – 8	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	materials_examined	Type material. Holotype ♂ (NZC-ZSI- 9429 / 18) from INDIA: Nagaland: Peren District, Ngwalwa (25 ° 39 ’ 38 ” N 93 ° 47 ’ 06 ” E), 547 m a. s. l., 28. II. 2024, leg. P. P. Sudhin. Paratypes: 2 ♀♀ (NZC-ZSI- 9430 / 18), same data as for holotype. 2 ♂ ♂ & 1 ♀ (NZC-ZSI- 9431 / 18), Meghalaya: Ri Bhoi District, Anderson Tea Estate (25 ° 47 ’ 42 ” N, 91 ° 53 ’ 03 ” E), 810 m a. s. l., 13. VI. 2024, leg. P. P. Sudhin.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	etymology	Etymology. The specific epithet is a noun in apposition, dedicated to our friend Mr. Nathanael P. A. Newmai, in recognition of his unwavering support during our surveys in Nagaland and Meghalaya, as well as his passion for spiders and their conservation in northeastern India.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	diagnosis	Diagnosis. The male copulatory organ of P. nathanael sp. nov. is most similar to that of P. himalayanus Simon, 1906 in having a similar shape of conductor and U-shaped sperm duct, but it can be distinguished by the following combination of characters: embolus in ventral view broader than in P. himalayanus, slightly curved medially with distal third directed anteroretrolaterally (embolus narrower and almost straight in P. himalayanus); conductor in retrolateral view at its proximal section broader than in P. himalayanus; sperm duct covers two-thirds of the area of tegulum (half of the tegulum in P. himalayanus) (cf. Figs 5 A, 6 A vs Bayer 2012: figs 31 b, 85 g). The female genitalia of P. nathanael sp. nov. is most similar to that of P. marsyandi Levi, 1982 in general shape of epigyne and vulva, from which it can be distinguished by the following combination of characters: median septum anterolaterally with prominent lobe at each side (less prominent in P. marsyandi); copulatory openings relatively large and located medially on epigyne (small, and located mediolaterally in P. marsyandi); copulatory ducts longer than in P. marsyandi; spermathecal heads with long stalk (with very short or without any stalk in P. marsyandi) (cf. Figs 5 C – E, 7 A – B vs Bayer 2012: figs 34 A – D, 88 i, 91 i). The vulva of P. nathanael sp. nov. can also be differentiated from those of P. demiror Bayer, 2012 and P. pakawini Bayer, 2012 by the spermathecal heads with (at least slightly) longer stalk (cf. Figs 5 E, 7 B vs Bayer 2012: figs 91 j, l).	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	description	Description. Male. (Holotype, NZC-ZSI- 9429 / 18) (Figs 4 A – B, 5 A – B, 6) Measurements: body length 20.36; carapace length 8.55, width 6.13, anterior width of carapace 3.34; opisthosoma length 11.81, width 4.61. Eye diameters: AME 0.34, ALE 0.38, PME 0.49, PLE 0.49. Eye interdistances: AME – AME 0.18, AME – ALE 0.04, ALE – ALE 1.02, PLE – PLE 1.72, PME – PME 0.17, PME – PLE 0.19, AME – PME 0.54, ALE – PLE 0.30. Clypeus height at AME 0.87, clypeus height at ALE 0.71. Chelicerae 2.62 long, with three promarginal and four retromarginal teeth. Measurement of palp and legs: Palp 11.39 [4.11, 1.40, 2.19, 3.69], leg I 52.89 [14.01, 4.08, 15.48, 14.18, 5.14], II 41.42 [11.46, 3.73, 12.36, 10.01, 3.86], III 29.38 [8.46, 2.90, 7.77, 6.85, 3.40], IV 42.14 [12.48, 3.41, 11.39, 10.22, 4.84]. Leg formula: 1423. Spination. Palp: femur pl 1 rl 1 do 3. Legs: femur I – II pl 6 rl 6 do 4, III – IV pl 5 rl 5 do 4; tibia I pl 3 rl 3 plv 3 rlv 3, II pl 3 rl 3 plv 4 rlv 4, III pl 2 rl 3 plv 2 rlv 1, IV pl 3 rl 3 plv 3 rlv 3; metatarsus I pl 5 rl 5, II pl 4 rl 4 v 2, III – IV pl 3 rl 3 plv 2 rlv 2 v 1; tarsus I – IV spineless. Carapace pale yellow, covered with dark brown setae, with a broad, light brown median band, and narrow light brown lateral bands; the median band with a broad pale-yellow marking anteriorly, and a triangle-shaped pale-yellow marking posteriorly, the sides of median band undulating; the lateral bands extending from behind PLE (Fig. 4 A). Eye field light greyish-brown, covered with setae (Fig. 4 A). Clypeus pale yellow. Chelicerae pale yellow to reddish-yellow, covered with setae. Endites pale yellow, scopulate, outer margins with black lines, sparsely covered with long black setae (Fig. 4 B). Labium pale yellow, covered with black setae (Fig. 4 B). Sternum light brown, covered with long erect black setae and short black setae; sides yellowish, covered with short white setae (Fig. 4 B). Legs pale yellow, reddish-brown towards the distal segments; coxae of leg I ventrally with macrosetae. Opisthosoma dorsally greyish-brown, suffused with grey patches, anteriorly with a median spindle-shaped pale-yellow marking (Fig. 4 A); laterally with several longitudinal grey dotted lines on brown background. Venter light greyish-brown, medially with a pale-yellow longitudinal stripe extending the entire length of opisthosoma, more prominent in anterior region (Fig. 4 B). Spinnerets grey to greyish-brown. Palp as shown in Figs 5 A – B, 6: tegulum (T) and cymbium light reddish-yellow, rest of segments pale yellow (Fig. 5 A – B); tibia provided with three fields of long, strong, and densely arranged black setae (Figs 5 A – B, 6); cymbium dorsally covered with dense scopula, covering approximately two-thirds of its surface (Figs 5 B, 6 B); tegulum nearly oval, with a U-shaped sperm duct (SD) reaching posterior third of the tegulum in ventral view (Figs 5 A, 6 A); subtegulum clearly visible in ventral view; conductor (C) relatively broad and long, with anteroretrolateral orientation, arising in 10 o’clock position on tegulum (Figs 5 A, 6 A); embolus (E) medium-sized, distally third slightly curved retrolaterally, narrowing towards the distal region, arising retrolaterally at distal half of tegulum (Figs 5 A, 6 A). Female. (paratype) (Figs 4 C – D, 5 C – E, 7): body length 20.36; carapace length 10.83, width 7.32, anterior width of carapace 3.94; opisthosoma length 16.21, width 8.58. Eye diameters: AME 0.49, ALE 0.60, PME 0.52, PLE 0.63. Eye interdistances: AME – AME 0.20, AME – ALE 0.12, ALE – ALE 1.02, PLE – PLE 2.06, PME – PME 0.31, PME – PLE 0.22, AME – PME 0.39, ALE – PLE 0.29. Clypeus height at AME 1.23, clypeus height at ALE 1.10. Chelicerae 3.32 long. Measurement of palp and legs: Palp: 11.39 [4.11, 1.40, 2.19, 3.69], leg I 52.88 [14.00, 4.08, 15.48, 14.18, 5.14], II 41.41 [11.46, 3.73, 12.36, 10.00, 3.86], III 29.38 [8.46, 2.90, 7.77, 6.85, 3.40], IV 42.34 [12.48, 3.41, 11.39, 10.22, 4.84]. Leg formula: 1423. Spination. Palp: femur pl 1 rl 1 do 3. Legs: femur I – II pl 5 rl 6 do 5, III – IV pl 5 rl 5 do 5; tibia I pl 2 rl 3 plv 4 rlv 4, II pl 4 rl 4 plv 4 rlv 4, III pl 3 rl 3 plv 2 rlv 2, IV pl 3 rl 3 plv 2 rlv 1; metatarsus I – II pl 3 rl 3 plv 2 rlv 2 v 1, III pl 4 rl 3 plv 3 rlv 3 v 1, IV pl 3 rl 3 plv 2 rlv 1; tarsus I – IV spineless. In all details as male, except the following: female longer and more robust than male; opisthosoma slightly lighter, with separated lateral longitudinal grey dotted lines; chelicerae, endites, and labium darker. coxae of leg I ventrally without macrosetae. Genitalia as shown in Figs 5 C – E, 7. Epigynal field wider than long; slit sense organs (SO) and epigynal muscle sigilla anteriorly beyond epigynal field (Figs 5 C, 7 A); median septum (MS) large, broader than long, anterolaterally with lateral humps on each sides (Figs 5 C – D, 7 A); copulatory openings large, nearly oval, separated from each other, positioned medially beyond the lateral humps mentioned above (Figs 5 C – D, 7 A); copulatory ducts relatively short, broad, and transparent (Figs 5 E, 7 B); spermathecal heads (SH) with relatively long stalk (Figs 5 E, 7 B); fertilization ducts curved, oriented posteriorly, positioned at the posterior region of spermathecal base (SBA) (Figs 5 E, 7 B). Note. The himalayanus group currently comprises 11 species distributed across northern India, Nepal, Bhutan, China, Myanmar, Thailand, Laos, Cambodia, and Vietnam (Bayer, 2014). This group is characterized by the presence of a distinct field of macrosetae on the ventral side of the male coxae of legs I (Bayer 2012). The presence of these macrosetae, along with the general morphology of the male and female copulatory organs, strongly supports the placement of this new species within the himalayanus group.	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
73560112CE3BA920C6F25E66B72CFEBA.taxon	distribution	Distribution. India (Nagaland and Meghalaya) (Fig. 8).	en	Sudhin, Puthoor Pattammal, Sarma, Shouvik Mali Pratyush Das, Sen, Souvik (2025): Two new species of the spider genus Psechrus Thorell, 1878 (Araneae: Psechridae) from northeastern India. Zootaxa 5609 (3): 419-429, DOI: 10.11646/zootaxa.5609.3.7, URL: https://doi.org/10.11646/zootaxa.5609.3.7
