taxonID	type	description	language	source
F5676FAA51145933A2A3E01DFB57E21F.taxon	description	Figs 3 C, 4 E, F, 5 B, 9	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
F5676FAA51145933A2A3E01DFB57E21F.taxon	diagnosis	Diagnosis. Same as the genus.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
F5676FAA51145933A2A3E01DFB57E21F.taxon	description	Description. Shell conical, dextral, small to medium-sized (SH 22.2 – 32.5 mm), relatively thick, solid, opaque, glossy, with 6.0 – 7.0 whorls in adults. Whorls inflated. Suture rather deep. Early teleoconch with dense spiral lines invisible to the naked eye, apex blunt and low. Teleoconch almost smooth, without keels or hairs, with dense growth lines. Aperture waved, ovate, oblique, more than half of shell in height. Inner shell surface white. Outer lip black and thickened, inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Shell multicolored, teleoconch yellowish-green to brown, apex region darker. Operculum much smaller than aperture (OH / AH = 0.55 – 0.61), corneous, ovate, thin, reddish-brown, exterior surface of operculum rough, inner surface smooth. Radular central tooth with a relatively sharp central denticle and four small and sharp cusps on both sides; lateral tooth with a medium and blunt central denticle and three small and sharp cusps on both sides; inner marginal tooth with a relatively small and blunt central denticle and four small and sharp cusps on both sides; outer marginal tooth with 18 – 19 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
F5676FAA51145933A2A3E01DFB57E21F.taxon	description	Measurements. Holotype: SH 29.4 mm, SW 22.8 mm; AH 18.2 mm, AW 13.3 mm; OH 10.6, OW 7.3; SW / SH = 0.78, AW / AH = 0.73, AH / SH = 0.62, OH / AH = 0.58. Paratypes: SH 22.2 – 32.5 mm, SW 18.5 – 23.7 mm; AH 15.4 – 19.2 mm, AW 11.1 – 14.2 mm; OH 8.4 – 11.6, OW 6.3 – 7.3; SW / SH = 0.75 – 0.83, AW / AH = 0.67 – 0.75, AH / SH = 0.56 – 0.69, OH / AH = 0.55 – 0.61.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
F5676FAA51145933A2A3E01DFB57E21F.taxon	distribution	Distribution and ecology. Same as the genus.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	description	Figs 3 C, 4 E, F, 5 B, 9	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	type_taxon	Type species. Bapaludina dazhouensis Z. - G. Chen & Y. - S. Guo, sp. nov.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	diagnosis	Diagnosis. Shell small to medium-sized (SH 22.2 – 32.5 mm), relatively thick and solid; apex blunt. Whorls inflated. Suture rather deep. Teleoconch almost smooth, without keels. Inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Operculum thin, much smaller than aperture (OH / AH = 0.55 – 0.61), exterior surface of operculum rough, inner surface of operculum smooth. Lateral and inner marginal teeth with a small to medium, blunt central denticle and some small and sharp cusps on both sides.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	description	Description. See the species description.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	etymology	Etymology. The genus name is made from “ Ba, ” the abbreviation for eastern Sichuan and Chongqing, and “ paludina ” for river snails.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
A55E0727E8E1585E9A2D928F66322099.taxon	distribution	Distribution and ecology. Known from the tributaries of the upper Qujiang River (a tributary of Jialingjiang River) in the northern Dazhou (Fig. 6). Living in the pebble-bottom sections of the cold and flowing tributaries with seasonal floods (Fig. 6). No living animals were found in the main streams or the muddy or still-water sections of the tributaries.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	description	Figs 3 B, 4 C, D, 8	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	diagnosis	Diagnosis. Early teleoconch almost smooth, with very weak and sparse spiral lines. Teleoconch without a weak but distinct keel at suture, but with several weak and indistinct keels. Aperture ovate, elongated. Lateral tooth with a large, broad, and blunt central denticle and five small and sharp cusps on both sides; inner marginal tooth with a large, broad, and blunt central denticle and five small and sharp cusps on both sides; outer marginal tooth with 19 – 22 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	description	Description. Shell conical, dextral, small to medium-sized (SH 20.3 – 40.4 mm), relatively thick, solid, opaque, glossy, with 5.5 – 6.0 whorls in adults. Whorls inflated. Suture rather deep. Early teleoconch almost smooth, with very weak and sparse spiral lines, apex blunt and low. Teleoconch without a weak but distinct keel at suture, but with several weak and indistinct keels, without hairs, with dense growth lines. Aperture waved, ovate, elongated, oblique, more than half of shell in height. Inner shell surface orange to white. Outer lip brown, not thickened, inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Shell multicolored, teleoconch brown, apex region darker. Operculum much smaller than aperture (OH / AH = 0.77 – 0.80), corneous, ovate, rather thick, dark brown, exterior surface of operculum rough, inner surface of operculum with a smooth outer ring and a rough inner ring, the nuclear region with shiny grains. Radular central tooth with a relatively sharp central denticle and six small and sharp cusps on both sides; lateral tooth with a large, broad, and blunt central denticle and five small and sharp cusps on both sides; inner marginal tooth with a large, broad, and blunt central denticle and five small and sharp cusps on both sides; outer marginal tooth with 19 – 22 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	description	Measurements. Holotype: SH 30.8 mm, SW 23.5 mm; AH 19.3 mm, AW 14.0 mm; OH 15.1, OW 10.5; SW / SH = 0.76, AW / AH = 0.73, AH / SH = 0.63, OH / AH = 0.78. Paratypes: SH 20.3 – 32.3 mm, SW 17.6 – 23.6 mm; AH 14.9 – 20.1 mm, AW 9.6 – 14.2 mm; OH 11.5 – 15.8, OW 7.4 – 10.9; SW / SH = 0.73 – 0.87, AW / AH = 0.64 – 0.73, AH / SH = 0.61 – 0.73, OH / AH = 0.77 – 0.80.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	etymology	Etymology. The species is named after its type locality, Luzhou.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
03B472DC35B05E7A90D857FB21407E1C.taxon	distribution	Distribution and ecology. Known from the Huagaoxi River only, a tributary of the Yongninghe River in southern Luzhou (Fig. 6). Living in the pebble-bottom sections of the cold and flowing tributaries with seasonal floods in sympatry with Hua sp., Lithoglyphopsis sp., and Tricula sp. (Fig. 6). No living animals were found in the main streams or the muddy or still-water sections of the tributaries.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
4D584C3A740A5FBD9D9474EC71FC0343.taxon	description	Figs 2, 3 A, 4 A, B, 5 A	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
4D584C3A740A5FBD9D9474EC71FC0343.taxon	diagnosis	Diagnosis. Early teleoconch with dense spiral lines. Teleoconch with a weak but distinct keel at suture, above and below suture with several weak and indistinct keels. Aperture ovate. Lateral tooth with a large, broad, and blunt central denticle and four small and sharp cusps on both sides; inner marginal tooth with a large, broad, and blunt central denticle and four small and sharp cusps on both sides; outer marginal tooth with 15 – 16 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
4D584C3A740A5FBD9D9474EC71FC0343.taxon	description	Description. Shell conical, dextral, small to medium-sized (SH 23.8 – 40.4 mm), relatively thick, solid, opaque, glossy, with 6.0 – 7.0 whorls in adults. Whorls inflated. Suture rather deep. Early teleoconch with dense spiral lines invisible to the naked eye, apex blunt and low. Teleoconch with a weak but distinct keel at suture, above and below suture with several weaker and indistinct keels, without hairs, with dense growth lines. Aperture waved, ovate, oblique, more than half of shell in height. Inner shell surface orange to white. Outer lip black and thickened, inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Shell multicolored, teleoconch yellowish-green to reddish-brown, apex region darker. Operculum much smaller than aperture (OH / AH = 0.72 – 0.83), corneous, ovate, rather thick, dark brown, exterior surface of operculum rough, inner surface of operculum with a smooth outer ring and a rough inner ring, the nuclear region with shiny grains. Radular central tooth with a relatively sharp central denticle and six small and sharp cusps on both sides; lateral tooth with a large, broad, and blunt central denticle and four small and sharp cusps on both sides; inner marginal tooth with a large, broad, and blunt central denticle and four small and sharp cusps on both sides; outer marginal tooth with 15 – 16 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
4D584C3A740A5FBD9D9474EC71FC0343.taxon	description	Measurements. Syntype: SH 25.1 – 30.9 mm, SW 18.9 – 23.7 mm; AH 14.0 – 17.8 mm, AW 10.3 – 12.4 mm; SW / SH = 0.75 – 0.77, AW / AH = 0.70 – 0.74, AH / SH = 0.56 – 0.57. New materials: SH 23.8 – 40.4 mm, SW 20.2 – 29.6 mm; AH 15.8 – 22.1 mm, AW 11.6 – 16.7 mm; OH 12.9 – 16.1, OW 8.3 – 10.8; SW / SH = 0.72 – 0.85, AW / AH = 0.71 – 0.80, AH / SH = 0.55 – 0.66, OH / AH = 0.72 – 0.83.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
4D584C3A740A5FBD9D9474EC71FC0343.taxon	distribution	Distribution and ecology. Currently only found in the tributaries of the Minjiang River and the Tuojiang River in northwestern Chengdu (Fig. 6). Maybe distributed in the Qujiang River Basin from eastern Sichuan to northern Chongqing. Living in the pebble-bottom sections of the cold and flowing tributaries with seasonal floods in sympatry with Hua bailleti (Bavay & Dautzenberg, 1910), Lithoglyphopsis spp., and Tricula spp. (Fig. 6). No living animals were found in the main streams or the muddy or still-water sections of the tributaries.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	description	Figs 2, 3 A, B, 4 A – D, 5 A, 8	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	type_taxon	Type species. Paludina magnaciana Heude, 1889.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	diagnosis	Diagnosis. Shell small to medium-sized (SH 20.3 – 40.4 mm), relatively thick and solid; apex blunt. Whorls inflated. Suture rather deep. Teleoconch with several weak and indistinct keels. Inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Operculum rather thick, much smaller than aperture (OH / AH = 0.72 – 0.83), exterior surface of operculum rough, inner surface of operculum with a smooth outer ring and a rough inner ring, the nuclear region with shiny grains. Lateral and inner marginal teeth with a large, broad, and blunt central denticle and some small and sharp cusps on both sides.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	description	Description. Shell conical, dextral, small to medium-sized (SH 20.3 – 40.4 mm), relatively thick, solid, opaque, glossy, with 5.5 – 7.0 whorls in adults. Whorls inflated. Suture rather deep. Early teleoconch almost smooth or with spiral lines invisible to the naked eye, apex blunt and low. Teleoconch with several weaker and indistinct keels, without hairs, with dense growth lines. Aperture waved, ovate, oblique, more than half of shell in height. Inner shell surface orange to white. Outer lip black and thickened, inner lip of the aperture is folded and tightly overlapping on the columellar margin, completely covering the umbilicus and forming with growth a crescent-shaped fortification, even in juvenile specimens. Shell multicolored, teleoconch yellowish-green, brown, or reddish-brown, apex region darker. Operculum much smaller than aperture (OH / AH = 0.72 – 0.83), corneous, ovate, rather thick, dark brown, exterior surface of operculum rough, inner surface of operculum with a smooth outer ring and a rough inner ring, the nuclear region with shiny grains. Radular central tooth with a relatively sharp central denticle and six small and sharp cusps on both sides; lateral tooth with a large, broad, and blunt central denticle and four to five small and sharp cusps on both sides; inner marginal tooth with a large, broad, and blunt central denticle and four to five small and sharp cusps on both sides; outer marginal tooth with 15 – 22 long and very sharp cusps.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	etymology	Etymology. The genus name is made from the Shu abbreviation for Sichuan and paludina for river snails.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
5190098CCD655FE5A8F13A4CE8F4AB13.taxon	distribution	Distribution and ecology. Known from the tributaries of the Minjiang River and Tuojiang River in northwestern Chengdu and the Yongninghe River in southern Luzhou (Fig. 6). Maybe distributed in the Qujiang River from eastern Sichuan to northern Chongqing. The genus may have a wider potential distribution. The first author identified similar specimens during fish surveys in the Qingyijiang River at Yaan (Fig. 6) from 2017 to 2020 but did not preserve the specimens. Living in the pebble-bottom sections of the cold and flowing tributaries with seasonal floods in sympatry with Hua spp., Lithoglyphopsis spp., and Tricula spp. (Fig. 6). No living animals were found in the main streams or the muddy or still-water sections of the tributaries.	en	Chen, Zhong-Guang, Dai, Yu-Ting, Chen, Hui, Jiang, Jiao, Liu, Xin, Guo, Yan-Shu, Wang, Ping, Cheng, Yu-Qi, Wu, Jia-Wei, Ouyang, Shan, Fang, Yi-Feng, Wu, Xiao-Ping (2025): Two new genera and species of river snails from the upper Changjiang River Basin in China (Gastropoda, Viviparidae). Zoosystematics and Evolution 101 (4): 1857-1871, DOI: 10.3897/zse.101.153058
