taxonID	type	description	language	source
8316879B9F73FF9DFF128D9147D5FBB4.taxon	discussion	Morphological key to species of Yoyetta cicadas Specimens must be set with wings spread to begin using this key. A vernier calliper is required to check measurements. Male specimens must often be examined microscopically and may require dissection of the genitalia in some instances. The key is adapted from Emery et al. (2025).	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	description	(Plates 1 – 2; Figs. 1 – 3)	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	materials_examined	Types. Holotype ♂, Kangaroo River State Forest, Towallum, 30 ° 07 ' 42 " S 152 ° 51 ' 09 " E, 22. xi. 2022, T. J. Bush (EME 0785 - 001) (K. 629041; AM). Paratypes. NEW SOUTH WALES: 1 ♂, 1 ♀, same location as holotype, 22 - 24. xi. 2022, T. J. Bush, (EME 0785 - 002, 005) (K. 629042, K. 629043; AM); 8 ♂♂, same location as holotype, 22 - 24. xi. 2022, T. J. Bush, (EME 0785 - 003 – 004, 006 – 009; - 003, genitalia prep 785 - 1); 6 ♂♂, Kangaroo River State Forest, Towallum, 30 ° 07 ' 45 " S 152 ° 51 ' 06 ‘ E, 8. xii. 2022, T. J. Bush (EME 0785 - 012 – 017); 2 ♀♀, Towallum, 30 ° 07 ' 44 " S 152 ° 51 ' 01 " E, 15. xii. 2022, T. J. Bush (EME 0785 - 021 – 022) (DE); 1 ♂, same location as holotype, 23. xi. 2022, T. J. Bush; 1 ♂, same location as holotype, 28. xi. 2022, T. J. Bush; 2 ♂♂, Towallum, 30 ° 07 ' 44 " S 152 ° 51 ' 01 " E, 15. xii. 2022, T. J. Bush; 1 ♀, Towallum, 30 ° 07 ' 45 " S 152 ° 51 ' 06 " E, 8. xii. 2022, T. J. Bush; 1 ♂, same location as holotype, 20. x. 2023, T. J. Bush (TJB); 3 ♂♂, AUSTRALIA-NSW, Kangaroo Ck., Towallum, 30 ° 07 ' 41 " S 152 ° 51 ' 11 " E, 8. xii. 2022, T. J. Bush; 3 ♂♂, same location as holotype, 6. xii. 2023, T. J. Bush (EME 0785 - 018 – 020) (LWP); 2 ♂♂, same location as holotype, 22 - 24. xi. 2022, T. J. Bush, (EME 0785 - 010 – 011) (MM). Audio records (TJB). NEW SOUTH WALES: Wild Cattle Creek, 30 ° 11 ' 29 " S 152 ° 48 ' 59 " E, 16. xii. 2022; Deadmans Range Rd, Clouds Creek, 30 ° 04 ' 47 " S 152 ° 38 ' 19 " E, 29. xii. 2022; Clouds Creek, 30 ° 08 ' 02 " S 152 ° 36 ' 07 " E, 17. xi. 2023; Girard State Forest, 28 ° 56 ' 14 " 152 ° 18 ' 03 ", 8. xii. 2023; Girard State Forest, 28 ° 56 ' 37 " S 152 ° 17 ' 58 " E, 8. xii. 2023; Towallum, 30 ° 07 ' 42 " S 152 ° 51 ' 09 " E, 17. x. 2024; Clouds Creek, 30 ° 08 ' 02 " S 152 ° 36 ' 07 " E, 24. x. 2024; Towallum, 30 ° 07 ' 42 " S 152 ° 51 ' 09 " E, 14. xi. 2024; Girard State Forest, 28 ° 56 ' 14 " 152 ° 18 ' 05 ", 6. xii. 2024. Aural records (TJB, DE). Moonpar Forest Drive, 30 ° 13 ' 45 " S 152 ° 39 ' 06 " E; 1. xii. 2023.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	distribution	Distribution, habitat and seasonality. Plate 2 E – F; Fig. 1). Currently, Y. bushi sp. nov. is found predominantly in pockets of dry Blackbutt (Eucalyptus pilularis) or in Sydney Blue Gum (Eucalyptus saligna) forests at elevations greater than 500 m in Girard State Forest and in the region of Nymboi-Binderay National Park (Fig. 1). In Kangaroo River State Forest (30 ° 07 ' S 152 ° 51 ' E), specimens are found almost exclusively within dry Blackbutt forest. More extensive analysis of bushland habitat by TJB revealed additional sites with the dry Blackbutt habitat in Wild Cattle Creek State Forest (30 ° 11 ' S 152 ° 48 ' E) around 7 km SSW of Towallum and at Clouds Creek (30 ° 08 ' S 152 ° 36 ' E) around 24 km WNW of Towallum where Y. bushi sp. nov. was also found and the song recorded. The species has also been located and recorded in Sydney Blue Gum forests within Girard and Moonpar State Forests. The areas inhabited by this species consist of small plateaus and ridgetops with forest which is more open than the surrounding denser bush (Plate 2 E – F). Adult males show a preference for the dense crown of juvenile trees in the understorey or in more established forests with few juvenile trees, males call from the upper branches of eucalypts. Females have been observed on Forest Oak (Allocasuarina torulosa) and amongst the foliage of Blackbutt saplings but have been located most often sitting on the bark of small to medium sized dead trees within the understorey. Specimens have been observed from mid-October until early January. Populations of Y. bushi sp. nov. co-locate with Y. timothyi Emery, Emery & Popple, which was heard calling in the canopy at Girard State Forest, Towallum and Clouds Creek. Within Girard State Forest, they also co-locate with Y. ignita and another undescribed Yoyetta where New England Blackbutt (E. campanulata) forest converges with Sydney Blue Gum forest.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	etymology	Etymology This species is named after Tony Bush, who initially located specimens on his birthday, then captured and recorded all the specimens currently in collections.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	description	Description of adult male (Plates 1 A – B, E – F, 2 A – C; Fig. 2) Head including eyes about as wide as lateral margins of pronotal collar, wider than mesonotum, predominantly black around ocelli and eyes, an ochraceous triangular spot on the central posterior margin; postclypeus ochraceous dorsally, black ventrally, brown along posterior margin, transverse ridges black with ochraceous lateral margins; anteclypeus, lorum and gena black. Rostrum black proximally, brown centrally, becoming black apically, extending midway between middle and hind coxae. Antennae black, antennomeres black, pedicles black; eyes black, dark brown when alive, ocelli red. Thorax. Pronotum ochraceous brown along midline and raised areas, black areas either side of midline at posterior margin, additional black markings along paramedian and lateral fissures; pronotal collar dark brown, suffused with black over lateral third, lateral angles rounded, black. Mesonotum black, with ochraceous-brown areas around sigilla; scutal depressions black, cruciform elevation and lateral ridges black, depressions orange-brown; metanotum black centrally and at extremities, dull orange-brown over remainder. Thorax below black. Legs. Coxae black; trochanters black, coxal membranes orange-red; femora black with lighter lateral and distal areas, primary spine light brown, angled 45 o; tibiae dark brown, becoming lighter distally; fore- and mid-tarsi dark brown becoming light at tips, hind-tarsi cream; claws brown to black. Meracantha triangulate, pale, partially overlapping opercula (Plate 2 B). Wings. Hyaline; forewings with 8 apical cells; costal vein parallel-sided, orange to node, becoming darker distally, CuA orange, other venation black, reddish pterostigma present; basal cell tinged orange along anterior margin, translucent over remainder; basal membrane dull orange-black. Hind wings with distinct orange-grey plaga over jugum and proximal half of 2 A; venation black, with 6 apical cells. Opercula (Plate 2 B). Black at base, becoming pale cream over remainder, ovate, covering rim of distal margin of tympanal cavity, almost reaching the to the distal margin of tergite 2, not meeting. Timbals (Plate 2 A) with five long ribs; ribs 1 - 3 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 4 extending almost the full length of the timbal membrane; rib 5 unattached, extending over the medial two-thirds of the timbal membrane; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre. Abdomen (Plate 2 C). Tergite 1 black; tergite 2 black with small dark orange-brown circular markings either side of midline; tergites 3 – 7 predominantly dark orange each with a distinct black mark on dorsal midline, this mark broadest on tergite 3 and equivalently narrower on tergites 4 – 7, with black annular markings on the posterior third, expanding ventrally to cover the posterior two-thirds, widest laterally on tergites 4 – 6; tergite 8 black; epipleurites black along medial border, orange-grey over remainder, flexed ventrally. Sternites I – II black; sternites III-VII smudged orange-grey over anterior two-thirds, light orange posteriorly, sternite VII with a central black spot on the posterior half; sternite VIII black anteriorly, becoming pale cream over the posterior half. Genitalia (Fig. 2). Pygofer black, upper lobe of moderate size with a broadly rounded apex, in ventral view angled medially; basal pygofer lobe smaller, weakly developed, slightly rounded, a distinct groove angled around 45 o antero-ventrally between both lobes; median lobe of uncus of moderate length, in lateral view ‘ hipped’ posteriorly at base, evenly tapering to a rounded apex (Fig. 2 A); claspers robust, ‘ duck-beaked’ in lateral view, separated, in ventral view their inner margins diverging laterally and apically, outer margins notched near base, gently turned outwards to a bluntly pointed apex (Fig. 2 B); basal plate ‘ teat-like’, gently curved and widest at base, narrowing from hinge region to a rounded apex (Fig. 2 B); aedeagus recurved distally through 150 o with ornamentation distally from the recurvature, the vesica broadened and distinctly toothed either side towards the apex, apex terminating in a short, slender ‘ beak’ pointing outwards, downturned 30 o viewed laterally, with a pair of very small flat ear-like flanges directed medially and minutely toothed caudally (Figs. 2 D, E); a ‘ viking helmet, wings and ears’ shape viewed laterally (Fig. 2 D); pseudoparameres very long and slender, reaching the distal recurvature of the aedeagus, each with a rounded apex (Fig. 2 C). Description of adult female (Plates 1 C – D, G – H; 2 D. Colour and markings similar to male but lighter brown; black coloration present lateral to cruciform elevation. Abdominal tergites with dark brown posterior margins; abdominal segment 9 orange-brown, tending orange ventrally. Ovipositor sheath black; ovipositor striped dark brown and black, black at tip, extending 2 – 2.5 mm beyond posterior abdomen. Measurements (in mm; range with mean in parentheses for 10 males and 4 females, including smallest and largest specimens). Length of body including head: male 17.4 – 19.2 (18.5); female 21.3 – 22.6 (21.5). Length of fore wing: male 20.8 – 23.2 (21.9); female 24.3 – 26.0 (25.0). Forewing width: male 6.6 – 7.3 (6.9); female 7.9 – 8.2 (8.1). Width of head (including eyes): male 5.1 – 7.7 (5.4); female 5.3 – 5.8 (5.6). Width of pronotum (across lateral angles): male 5.1 – 5.6 (5.3); female 5.8 – 6.2 (6.0). Width of abdomen: male 4.7 – 5.3 (5.1); female 5.1 – 5.3 (5.2). Length of ovipositor: 7.1 – 7.5 (7.3). Morphological variations between and within population. Across its distribution, both male and female specimens of Yoyetta bushi sp. nov. exhibit coloration ranging from light and darker brown to blackish over the mesonotum and dorsal abdomen (Plate 1 E – H). Morphological distinguishing features. Males of Yoyetta bushi sp. nov. can be differentiated from males of Yoyetta species by three main features: (1) a body length (BL) 17.4 – 19.2 mm; (2) hindwing with orange-grey plaga over jugum and proximal half of 2 A; (3) lateral edges of tergites diffuse orange, with distinct and extensive dark markings along posterior margin. Females of Y. bushi sp. nov. can be distinguished from other female Yoyetta species by 3 criteria: (1), hindwing with orange-grey plaga over jugum and proximal half of 2 A; (2) BL between 21.3 – 22.6; and, (3) an ovipositor extending 2 – 2.5 mm beyond abdominal segment 9.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F70FF96FF128E0546ECF9D0.taxon	biology_ecology	Calling song (Fig 3). The male calling song is a series of clicking that ends in a short or long buzz. An examination of the song structure (all statistics, n = 9 recordings from Towallum and Wild Cattle Creek localities) as illustrated in Fig. 3 shows each ‘ click’ is a syllable comprising of 4 - 6 pulses (0.006 – 0.026 s). Amongst recordings, the number of clicks in a sequence varied considerably from 9 to 41. The ‘ short buzz’ at the end of a clicking sequence is a single syllable immediately followed by a short macrosyllable (0.030 – 0.071 s; Fig. 3 C, E). By comparison, the ‘ long buzz’ is an echeme that decreases in amplitude throughout (0.116 – 0.348 s; Fig. 3 B, D). The frequency plateau of the male calling song ranges from approximately 10.3 to 16.3 kHz, with a dominant frequency between 11.6 and 14.7 kHz (n = 3, RS 1; Fig. 3 F). It was noted in the field by TJB that the length of the buzz component of the song will be switched back and forth without any apparent stimulus, but the longer buzz frequently precedes the cessation of calling. Y. bushi sp. nov. predominantly calls at rest and observed instances of males calling in flight are an exceptionally rare phenomenon. Calling occurs in sunlight with cloud cover significantly reducing activity. Males commence calling between 0830 h and 0900 h (Australian Eastern Daylight Savings Time (AEDT) GMT + 11 hours) frequently moving between perches. By 1130 h (AEDT) much of the activity ceases, males move higher into the trees and become more sedentary but will continue to call sporadically until late in the afternoon. Finger snapping can sometimes elicit a response from calling males when timed correctly, but more often than not, this technique fails to attract their attention. The calling song of Yoyetta bushi sp. nov. closely resembles the high-frequency songs of Yoyetta fluviatilis Emery, Emery & Popple and Y. robertsonae Moulds, Popple & Emery. However, its structure differs: Y. fluviatilis produces a short buzz followed by a click, whereas Y. bushi sp. nov. emits a series of distinct clicks before a short buzz. Similarly, the calling song of Y. robertsonae is much lower frequency and features evenly spaced groups of three clicks but lacks the final short buzz present in Y. bushi sp. nov.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	description	(Plates 3 – 4, Figs. 1, 4 – 6) Types. Holotype ♂, Goonoowigal State Conservation Area (SCA), Inverell, 28 o 48 ' 58 " S 151 o 07 ' 03 " E, 2. xii. 2022, T. Bush & D. Emery (EME 0504 - 001) (K. 629040; AM).	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	materials_examined	Paratypes. NEW SOUTH WALES: 2 ♂♂, same data as holotype (EME 0504 - 002 – 3); 1 ♂ 1 ♀, same location as holotype, 3. xii. 2022, T. Bush & D. Emery (EME 0504 - 006 - 7; genitalia prep 504 - 2); 2 ♂♂, same location as holotype, 4. xii. 2022, T. Bush (EME 0504 - 008 – 009); 1 ♂, same location as holotype, 22. xi. 2024, D. Emery (EME 0504 - 021); 3 ♂♂, 20 km W Armidale, 30 o 29 ' 07 " S 151 o 26 ' 37 " E, 10. i. 2006, D. Emery (EME 0504 - 012 – 015); 5 ♂♂, 16 km W Armidale, 30 o 29 ' 57 " S 151 o 29 ' 55 " E, 3. i. 2013, N., C. & D. Emery (EME 0504 - 016 – 020; 016, genitalia prep 504 - 1); 2 ♂♂, same location as holotype, 29. xi. 2024, T. J. Bush (EME 0504 - 017 – 8) (DE); 2 ♂♂, same location as holotype, 4. xii. 2022, T. Bush (TJB); 1 ♂, AUSTRALIA NSW, 16 km W Armidale, 959 m, 30 ° 29.57 ' S 151 ° 29.55 E, 5. i. 2013, N. C. & D. Emery; 1 ♂, AUSTRALIA-NSW, Goonoowigall CRes, 29 ° 48 ' 58 " S 151 ° 07 ' 01 " E, 2. xii. 2022, T. Bush & D. Emery; 1 ♂, AUSTRALIA-NSW, Goonoowigall CRes, 29 ° 48 ' 58 " S 151 ° 07 ' 01 " E, 3. xii. 2022, T. Bush & D. Emery (LWP); 2 ♂♂, same location as holotype, 4. xii. 2022, T. Bush (EME 0504 - 010 – 011) (MM); 1 ♂, Copeton Dam, 29 ° 54 ' 52 " S 151 ° 00 ' 56 " E, 10. xi. 2024, K. Warner (KW). Audio records. Howell Rd., Copeton Dam, 3. xii. 2022, D. Emery; Apex Lookout Rd., Bingara, 22. xii. 2024 B. McBurney.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	distribution	Distribution, habitat and seasonality (Plate 4 E – F, Fig. 1). Yoyetta nathani sp. nov. is located in the New England region of NSW, from around and west of Armidale to Yarrowyck and Bundarra, and northwards to Bingara, Copeton Dam and Inverell (Fig. 1). Males appear to favour the trunks and branches of partially smooth barked eucalypts (such as Angophora floribunda and Eucalyptus blakelyi) at heights of 3 - 8 m (depending on the height of local trees). They also frequent understorey shrubs (Slender Tea-tree, Gaudium brevipes) during ticking flights. The only female to be found was located within 1 m of the ground in small G. brevipes. Most specimens have been collected between mid-November and mid-January, but males have been recorded into early February.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	etymology	Etymology. The species is named after Dr. Nathan Emery, who located the majority of early specimens of Y. nathani sp. nov. from the Armidale region.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	description	Description of adult male (Plates 3 A – B, E, 4 A – C; Fig. 4) Head slightly narrower than the lateral margins of pronotum; mostly light to dark orange-brown with limited triangular black colouration surrounding the ocelli, branching laterally along the posterior margins of the dorsal postclypeus and broadening towards the anterior margins of the eyes, a triangular orange-brown fascia with a base on the midline of the posterior border narrows to an apex midway between the ocelli; supra-antennal plates black, edged dull orange-brown around base of antennae; ocelli bright red; eyes dull red to light brown; postclypeus orange-brown dorsally with a black transverse stripe along the anterior margin, ventral surface black with an orange medial line and orange coloration along the lateral and posterior margins; anteclypeus shiny black; lorum black, gena black, dull orange anteriorly; rostrum reaching anterior margins of hind coxae, dark brown proximally, becoming dark brown to black distally; antennae black. Thorax mostly brown to orange-brown. Pronotum with a narrow orange-brown fascia along midline surrounded by black, resembling a keyhole; interior of pronotum orange to dark brown, with irregular black areas, especially along paramedian and lateral fissures and adjacent ridges, also bordering the lateral halves of the pronotal collar; pronotal collar orange-brown, black along anterior margins of the lateral halves. Mesonotum orange- brown, lateral margins burnt red-brown, submedian and lateral sigilla black, lateral margins black; scutal depressions black; cruciform elevation orange-brown, black along midline and laterally along anterior margin of anterior ridge; wing grooves mainly dark orange-brown. Metanotum dark brown to black centrally, with black ridges. Thorax below mainly black. Legs brown with variable longitudinal orange-brown to black markings between specimens. Coxae about half orange-brown and black; fore femora with inner surface black, black posteriorly along femoral spines, outer surface predominantly dull orange-brown, femoral spines dark brown basally and black tipped, upright; mid and hind femora dark brown to black anteriorly, paler orange-brown posteriorly; fore tibiae black brown proximally, becoming brown distally; mid and hind tibiae orange-brown, black at joints; tarsi orange; claws black; meracantha variably black on base around medial half, remainder pale cream-brown, reaching anterior margin of opercula (Plate 4 B). Wings with forewing costal margin and CuA dull orange-brown; venation generally dark brown to black; basal cell translucent; basal membrane bright orange; smoky infusions throughout a 1 – a 8. Hind wing venation black; v-shaped plagas tinged orange-grey visible along vein 2 a and 3 a and within anal cell. Opercula (Plate 4 B); variably black basally, plates pale cream, spatulate, curved medially, flexed ventrally, not meeting; approximately 1.5 x larger than adjacent hind coxae. Timbals (Plate 4 A) with five long ribs; ribs 1 - 3 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 4 unattached, spanning entire timbal membrane, rib 5 unattached, extending over central half of membrane; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre. Abdomen with tergite 1 black; tergite 2 black laterally and along anterior margin, dark orange centrally, a black rectangular spot over posterior midline, curved around lateral margin of exposed timbals; tergites 3 – 7 predominantly dusky orange-brown each with a distinct black rectangular mark on dorsal midline, this mark broadest on tergite 3 and equivalently narrower on tergites 4 – 7, variable annular black infusions adjacent to posterior margin, posterior margins bright orange; tergite 8 black, dark orange-brown marks on posterior midline and anterolateral aspect (Plate 4 C); epipleurites orange, flexed ventrally, with mottled blackish suffusions. Sternite I black; sternite II black, orange ridges along timbal margins; sternites III-VII orange, sternite VII orange with an indistinct black spot on the posterior midline; sternite VIII orange-brown, infused with black along lateral aspects. Genitalia (Fig. 4) with pygofer orange dorsally, dorsal beak black, black over remainder, in lateral view, upper lobe of moderate size with a broadly rounded apex, in ventral view angled medially; basal pygofer lobe small weakly developed, broadly rounded over anterior margin of pygofer; median lobe of uncus relatively narrow and elongated with rounded apex and length around 1.0 mm (Fig. 4 A), slightly convex in ventral view; claspers robust, ‘ duckbilled’ in lateral view, partially meeting, in ventral view their inner margins diverging anteriorly from the midline, outer margins gently turned outwards to a bluntly pointed apex (Fig. 4 B); basal plate triangulate in ventral view with concave anterior border, relatively flat in lateral view (Fig. 4 C); aedeagus recurved distally through 180 o with ornamentation distally from the recurvature, the vesica expanding and distinctly toothed either side towards apex; apex terminating in typical ‘ viking helmet with ears’ in lateral view Figs. 4 C, D), with a short, slender ‘ beak’ curved ventrally, downturned around 45 o in lateral view, with a pair of very small flat ear-like flanges directed medially and minutely toothed (Fig. 4 E); pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, hair-like, each with a narrow but rounded apex (Fig. 4 C). Description of adult female (Plates 3 C – D, F, 4 D). Head, wings and legs similar to male. Thorax. Darker brown (“ burnt orange ”) coloration than male, with black line along midline, black markings on cruciform elevation extend into anterior depression and along posterior ridges. Abdomen. Tergite 1 black; tergite 2 dark burnt orange, suffused variably with black coloration; tergites 3 – 7 predominantly dark orange-brown each with a distinct black rectangular mark on midline, most pronounced on tergites 3 – 4, reducing slightly in width on successive tergites 5 – 7; annular black markings along posterior margins of each tergite with a distinct black marking centrally located on each ventrolateral aspect (Plate 4 D); tergite 8 orange, with a black spot on the posterior border of the lateral third. Sternite I black; sternites II-VII orange with a black infusion along midline; sternite VIII darker orange-brown with a distinct black spot either side of posterior midline; abdominal segment 9 orange dorsally, black over anterior half, bordered by a black semicircular marking, tending ochraceous laterally, with a black spot on lateral aspect; ovipositor sheath black, ovipositor dark brown, becoming black posteriorly, not extending beyond abdominal segment 9. Measurements (in mm; range with mean in parentheses for 14 males and 1 female, including smallest and largest of available specimens). Length of body including head: male, 21.9 – 25.2 (23.0); female, 21.1. Length of forewing: male, 25.8 – 29.8 (28.1); female, 28.8. Width of forewing: male, 8.5 – 10.1 (9.4); female, 9.5. Width of head (including eyes): male, 6.1 – 6.7 (6.4); female, 6.6 – 7.5 (7.0). Width of pronotum (across lateral angles): male, 6.1 – 6.6 (6.3); female, 6.4. Width of abdomen: male, 5.8 – 6.8 (6.3); female, 6.4. Length of ovipositor: female, 6.5. Morphological distinguishing features. Males of Y. nathani sp. nov. can be readily differentiated from other male Yoyetta by a combination of 6 characteristics: (1) BL> 22 mm; (2) v-shaped plagas tinged orange-grey visible along vein 2 a and 3 a and within anal cell of the hindwing; (3) 5 long timbal ribs; (4) a ‘ viking helmet with ears’ thecal apex (not a ‘ knob’ or lanceolate termination) with a strong downturn of the thecal apex; (5) a short medial lobe of the uncus; and (6) lack of smoky forewings. Females of Y. nathani sp. nov. can be distinguished from the other 20 Yoyetta species which have orange-grey visible hindwing plagas by the following combination of 5 characters: (1) BL> 21.0 (7 species); (2) deep burntorange coloration and annular black markings on the lateral tergites; (3) ovipositor not extending beyond abdominal segment 9; (4) lack of smoky wings; and, (5) black smudged markings along the midline of sternites.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F7BFF92FF128EF0403FF878.taxon	biology_ecology	Calling song (Figs. 5 – 6). The male calling song of Y. nathani sp. nov. at rest is series of 3 – 4 clicks, which is occasionally followed by a short buzz. Closer examination of the calling song (all statistics, n = 5 recordings from Goonoowigal SCA and Copeton Dam localities) is shown in Fig. 5, indicating each ‘ click’ is a macrosyllable comprising between 4 and 9 coalesced syllables (0.011 – 0.031 s; Fig. 5 C). A brief gap between 0.116 and 0.173 s duration punctuates each macrosyllable. The ‘ short buzz’ is an echeme that varies from 0.047 and 0.184 s duration. The gap between each calling song sequence (i. e., clicking and short buzz) ranges from 0.370 to 0.457 s duration. Males may also flick their wings after 1 - 3 clicks as illustrated in Figs. 5 and 6. Noting the limitations on iPhone recordings in the Methods and Terminology section, the approximate frequency plateau ranges from 5.2 – 12.6 kHz, with an approximate dominant frequency between 5.8 and 10.9 kHz (Fig. 5 E). Yoyetta nathani sp. nov. calls at rest, and occasionally, in flight. At Goonoowigal SCA, calling did not commence until around 1000 - 1100 h (AEDT), where males sang in a chorus lasting around 0.5 - 1 minute at intervals of around 10 minutes in sunlight. Cloud cover delayed and reduced singing, though males have been observed calling in warm, overcast conditions, during light precipitation and immediately following storms. At Copeton Dam and west of Armidale, individuals called for the same length of time but around every 5 minutes, constantly moving positions. Males could be attracted by finger snapping. The male calling song of Yoyetta nathani sp. nov. is similar to that of Yoyetta tristrigata (Goding & Froggatt) but the latter differs in its higher frequency and the absence of a short buzz at the end of its 3 - 4 tick phrase.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	description	(Plates 5 – 6; Figs. 1, 7 – 8)	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	materials_examined	Types. Holotype ♂, Cooper Drive, Clandulla, 32 o 54 ' 27 " S 149 o 49 ' 00 " E, 30. xi. 2020, D. Emery (EME 0713 - 001) (K. 629044; AM). Paratypes. NEW SOUTH WALES: 1 ♂ 1 ♀, same data as holotype (EME 0713 - 013, - 014) (K. 629045, K. 629046, AM); 1 ♂, 2 km S Kandos, 32 o 52 ' 26 " S 149 o 57 ' 57 " E, 7. i. 2010, D. Emery & L. Popple; 7 ♂♂ 2 ♀♀, same data as holotype (EME 0713 - 003 – 007, - 010 – 012; - 003 genitalia prep 713 - 1); 6 ♂♂ 1 ♀, same location as previous, 15. i. 2021, N. & D. Emery (EME 0713 - 015 – 019, - 021 – 022) (DE); 1 ♂, AUSTRALIA NSW, 5 km S. of Kandos, Capertee Valley, open woodland, 8. i. 2010, Popple & Emery, 32 ° 54 ' 06 " Sx 149 ° 57 ' 32 " E. 473 - 0006; 2 ♂♂, same data as holotype, 30. xi. 2020, D. Emery (EME 0713 - 008, - 009); 1 ♀, same location as holotype, 9. i. 2021, N., C. & D. Emery (EME 0713 - 020) (LWP); 2 ♂♂, same location as holotype, 24. i. 2023, D. Emery (EME 0713 - 023 – 024) (MM).	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	distribution	Distribution, habitat and seasonality. (Plate 6 E – F, Fig. 1). At present, Yoyetta crepita sp. nov. has only been captured around Clandulla and Kandos in the Bylong Valley, NSW (Fig. 1). This species appears to favour smooth barked eucalypts, particularly Eucalyptus melliodora (Yellow Box) and the rough-barked E. cannonii (Capertee Stringybark). Individuals are located in the upper reaches of the eucalypts> 10 m from ground level, but may descend briefly to lower levels during ticking flights (see “ calling song ”). At Clandulla, Y. crepita sp. nov. cohabits with Y. humphreyae Moulds & Popple, Y. timothyi Emery, Emery & Popple, Y. psammitica Emery, Emery & Popple and an additional undescribed Yoyetta species. Most specimens have been present from mid-November to late-January.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	etymology	Etymology. The Latin “ crepita ” (medical term, from the verb “ crepo ”, meaning to crackle or creak) is designated for this species due to the crackling chorus produced by the males at intervals on sunny days.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	description	Description of adult male (Plates 5 A – B, E, 6 A – C. Fig. 7) Head approximately as wide as to slightly wider than mesonotum; dorsally mainly black, with a thin, brown fascia posterior to ocelli; ocelli pinkish-red on live specimens; postclypeus dorsally ochraceous, suffused with black, with a thin brown median fascia, ventral surface black, lateral ridges black, lateral margins pale brown; anteclypeus black; rostrum dark brown, black at apex, reaching anterior edge of hind coxae; lorum and gena dull black with pale brown anterior margins; eyes variably dull black to dark brown; antennae black, supra-antennal plates black. Thorax predominantly shiny black with variable orange-brown fascia. Pronotum dorsally with brown midline fascia immediately posterior to pronotal collar extending along the anterior half of the midline; a black diamond shaped fascia posteriorly; with extensive orange-brown markings between sutures; pronotal collar black anteriorly and across lateral angles, orange-brown posteriorly and an “ open U ” - shaped fascias on midline directed anteriorly. Mesonotum black with dark orange-brown colouration between submedian and lateral sigilla; scutal depressions black; cruciform elevation orange with black midline and ridges; metanotum black along midline and lateral halves, darker orange-brown over medial half. Legs. Coxae black with orange basal and apical margins; fore femora with anterior and posterior surfaces black, striped orange-brown laterally and medially, orange at joints, femoral spines black, directed 45 o posteriorly; mid and hind femora with outer side black, inner side dark brown to brown; fore and mid tibiae black to dark ochraceous, with spines variably dull ochraceous and black at tips; hind tibiae paler brown; tarsi black proximally, ochraceous to claws, claws ochraceous, black at tips. Wings with forewing costal margin black to dark orange-brown; venation generally dark brown to node, black distally; basal cell translucent; basal membrane bright orange; smoky infusions throughout apical cells a 1 – a 8. Hind wing venation dark brown to black, bright orange at base of M vein; v-shaped plagas orange along vein 2 a and 3 a and within anal cell, with 6 apical cells. Opercula (Plate 6 B) small, spatulate, following body axis ventrolaterally, depressed centrally; black over anterior (basal) third, light cream over; clearly separated. Meracanthus small, narrow, cream, black at base, pointed, overlapping anterior third of opercula. Timbals (Plate 6 A) with five distinct long ribs; long ribs 1 – 3 extending across surrounding membrane and fused dorsally along basal spur; long rib 4 with medial and lateral components connected by a narrow middle segment; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large, ridged dome on posterior timbal plate extending across two-thirds of timbal; apodeme pit oval-shaped and conspicuous. Abdomen with tergite 1 black, orange-brown at posterior intersegmental membrane; tergite 2 black with prominent reddish-orange triangular marking located on the posterior half of tergite 2 posterior to timbals (Plate 6 C), posterior margin burnt orange-brown over middle third, black laterally; tergites 3 – 7 orange-brown with black median triangular markings along midline, widest at tergite 3, decreasing in width on tergites 4 – 6, widening on posterior margin of tergite 7, posterior margins with consecutive black and burnt orange annular rings, black markings broadening slightly towards lateral extremities; tergite 8 shiny black, with a small amount of orange-brown colouration on the anterior lateral side; epipleurites orange, flexed ventrally. Sternite I black, slight orange at lateral margins and through tympanal cavity; sternite II black, with orange-brown coloration on lateral posterior margins; sternites III – VI orange; sternite VII orange with dark brown to black marking on posterior half of midline; sternite VIII black anteriorly, becoming dark orange-brown posteriorly, with slightly opalescent pubescence. Genitalia (Fig. 7). Pygofer mainly black; dorsal beak brown, anal styles orange-brown; pygofer upper lobe of moderate size with a rounded apex; basal pygofer lobe small weakly developed, broadly rounded. Uncus orange-brown; in lateral view beak-like and stumpy, median lobe of uncus moderate, lobes in ventral view bulbous, with rounded lateral termination; claspers clearly divided, with apices directed and gradually tapering laterally. Aedeagus recurved distally through 180 degrees, with transparent toothed flanges along margins of recurvature, these broadly smooth along dorsal and posterior edges, finely serrated and broadening to> 2 x width of theca ventrally, with dorsal ornamentation restricted to termination adjacent to apex of theca; apex short, sclerotised, transparent, lanceolate, without ornamentation, tip directed slightly upwards; pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, each with a small rounded apical knob. Description of adult female (Plate 5 C – D, F, 6 D). Head, thorax, wings and legs similar to male but with more abundant orange-brown markings on the pronotum and mesonotum; metanotum suffused orange-black with black stripe along midline. Abdomen. Tergite 1 orange, black centrally and laterally along posterior margin, orange over remainder; tergites 2 – 7 predominantly dark orange-brown each with a distinct black fascia along the midline, extending laterally along the third quarter of each tergite midline, most pronounced on tergites 3 – 5; the posterior margins striped bright redorange; tergite 8 with small black spot on midline, stippled black along anterior margin, burnt orange- brown over remainder. Epipleurites flexed ventrally with black infusion on each medial margin. Sternite I black, ochraceous posteriorly; sternite II black on midline and lateral margins, orange over remainder; sternites III-VII orange with smudged black infusions along midline; sternite VIII orange with distinct black oval marking directed laterally from the anterior margin to midway across the sternite; abdominal segment 9 brown along dorsal midline, black over anterior half, extending posteriorly, black ventral border expanding dorsally to a spot on the posterior margin (Plate 6 D); ovipositor sheath black, ovipositor dark brown, becoming black posteriorly, not extending beyond abdominal segment 9. Measurements (in mm; range with mean in parentheses for 10 males and 5 females, including smallest and largest specimens). Length of body including head: male, 17.7 – 20.6 (19.5); female, 18.7 – 20.4 (19.5). Length of forewing: male, 21.0 – 23.3 (22.4); female, 25.1 – 25.4 (25.3). Width of forewing: male, 7.3 – 8.2 (7.6); female, 8.0 – 8.2 (8.1). Width of head (including eyes): male, 4.5 – 5.5 (5.1); female, 5.7 – 6.0 (5.9). Width of pronotum (across lateral angles): male, 4.3 – 5.3 (5.0); female, 5.7 – 5.9 (5.8). Width of abdomen: male, 4.7 – 5.8 (5.4); female, 5.0 – 5.3 (5.1). Length of ovipositor: female, 4.8 – 5.4 (5.2). Morphological variations between and within population. Both male and female Y. crepita sp. nov. exhibit a consistent morphology at Clandulla. It remains to be determined if variation exists within this species if specimens are found at additional locations. Morphological distinguishing features. Y. crepita sp. nov. has a body length <20.6 mm and hindwing plagas orange-grey and extending along veins 2 a and 3 a and within the anal cell. This feature is shared with 12 other Yoyetta species. Males of Y. crepita sp. nov. can be distinguished from these species by the following combination of 6 characters: (1) the presence of smoky wings with infusions in forewing apical cells; (2) a prominent reddish-orange triangular marking located laterally on the posterior half of tergite 2 posterior to timbals; (3) bright orange basal membranes; (4) brown markings on the lateral areas of tergite 8; (5) reflection of the theca through 180 o and a slightly upturned lanceolate thecal apex; and, (6) when viewed laterally, a 90 o hinge on the basal plate of the aedeagus. Females of Y. crepita sp. nov. have body lengths between 18.6 – 20.6 mm that differentiate them from females of 11 of the other 16 Yoyetta species that possess hindwing plagas orange-grey and extending along veins 2 a and 3 a and within the anal cell. An additional four features of female Y. crepita sp. nov. differentiates them for these other 5 species: (1) ovipositor not extending beyond abdominal segment 9; (2) extension of the black midline markings laterally around the posterior half of the tergites; (3) two black ovoid spots on sternite VIII; and (4) smudged black along midline of orange sternites.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F62FF89FF128D7647B3FB60.taxon	biology_ecology	Calling song (Fig. 8). The male calling song of Y. crepita sp. nov. is a series of sharp, rapid ticks. Following analyses of the calling song structure (all statistics, n = 3 from Clandulla locality), Fig. 8 shows each ‘ tick’ is a single syllable between 0.005 and 0.016 s duration, and the gap between syllables ranges from 0.159 to 0.185 s duration. The frequency plateau ranges from 9 to 14.4 kHz, with a dominant frequency approximately between 10.5 and 11.9 kHz (Fig. 8 C). Groups of male Y. crepita sp. nov. emit a distinctive, rapid clicking chorus from the resident population at around 5 min intervals when sunny (and cease mid-chorus with any cloud cover). During these choral cacophonies, males may relocate, calling in flight and often resting for short intervals at lower heights before resuming flight. This behaviour is likely aimed at finding responding females, as males respond readily to finger clicking during the chorus, particularly when present in large numbers. Males may also call for brief periods when resting. Females are usually found on the main trunks around 5 m from ground and have been observed wing-flicking during the chorus. The calling song of Yoyetta crepita sp. nov. closely resembles those of Y. australicta Popple & Emery, which also produces single pulses. While both species have broad frequency plateaus, the pulse gap of Y. australicta is 0.27 – 0.67 s, greater than for Y. crepita sp. nov. (<0.19 s). In addition, males of Y. crepita sp. nov. do not emit quiet macrosyllables at the end of their bouts of ticking.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	description	(Plates 7 – 8, Figs. 9 – 10)	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	materials_examined	Types. Holotype ♂, Toolangi CFA (Country Fire Authority) Stn., 37 ° 32 ' 32 " S 145 ° 28 ' 29 " E, 23. i. 2017, N., C. & D. Emery (to light, EME 0778 - 006) (HEM- 9295; NMV). Paratypes. VICTORIA. 2 ♂♂ 2 ♀♀, Wirrawilla Rainforest Park, Toolangi, 37 ° 31 ' 39 " S 145 ° 31 ' 17 " E, 1. i. 2023, K. Green, N. & D. Emery & T. Corbin (at light, EME 0778 - 059 – 061) (HEM- 9297 – 9299; NMV); 3 ♂♂, same location as holotype, 11. i. 2017, S. Emery & T. Corbin (at light, EME 0778 - 001 – 3); 2 ♂♂, same location, 23. i. 2017, S. Emery & T. Corbin (to light, EME 0778 - 004, 006, - 004 genitalia prep); 1 ♀, Ridge Rd, Mt. Dandenong, 37 ° 49 ' 27 " S 145 ° 21 ' 32 " E, 20. i. 2017, S. Emery & T. Corbin (to light, EME 0778 - 005); 1 ♀, same location, 29. i. 2017, S. Emery & T. Corbin (EME 0778 - 007); 2 ♂♂ 2 ♀♀, Olinda golf course, 37 ° 51 ' 13 " S 145 ° 22 ' 19 " E, 29. xi- 7. xii. 2017, S. Emery & T. Corbin (emerging, EME 0778 - 011 – 14); 1 ♀, same location, 25. xii. 2018, S. Emery & T. Corbin (emerging, EME 0778 - 015); 1 ♂, Ridge Rd, Mt. Dandenong, 37 ° 49 ' 27 " S 145 ° 21 ' 32 " E, 25. xii. 2018, S. Emery & T. Corbin (EME 0778 - 016); 1 ♀, same location, 3. xi. 2019, S. Emery & T. Corbin (EME 0778 - 017); 1 ♂, same location, 30. xii. 2019, S. Emery & T. Corbin (EME 0778 - 018); 1 ♂, same location, 20. xii. 2022, S. Emery & T. Corbin (EME 0778 - 020); 1 ♀, Wirrawilla Rainforest Park, Toolangi, 37 ° 31 ' 39 " S 145 ° 31 ' 17 " E, 7. i. 2017, K. Green (at light, EME 0778 - 021); 1 ♂ 2 ♀♀, same location as previous, 26. xii. 2022, K. Green (at light, EME 0778 - 024 – 26); 43 ♂♂ 1 ♀♀, same location as previous, 1. i. 2023, K. Green, N. & D. Emery & T. Corbin (at light, EME 0778 - 027, 029 – 058, 064 – 078) (DE); 5 ♂♂ 4 ♀♀, same location, 7. i. 2017, K. Green (at light); 6 ♂♂ 3 ♀♀, same location, 26. xii. 2022, K. Green (at light) (KG). 1 ♂, Wirrawilla Rainforest Park, Toolangi, 37 ° 31 ' 39 " S 145 ° 31 ' 17 " E, 7. i. 2017, K. Green (at light, EME 0778 - 022); 1 ♀, same location as previous, 26. xii. 2022, K. Green (at light, EME 0778 - 023) 1 ♂ 1 ♀, same location as previous, 1. i. 2023, K. Green, N. & D. Emery & T. Corbin (at light, EME 0778 - 028) (TJB); 1 ♂, AUSTRALIA-VIC, Olinda Golf Course, 37 ° 51 ' 13 " S 145 ° 22 ' 19 " E, 6. ii. 2017, S. & T. Corbin, EME 0778 - 010; 1 ♀, AUSTRALIA-VIC, Ridge Rd, Mt Dandenong, 37 ° 49 ' 27 " S 145 ° 21 ' 32 " E, S. Emery / T. Corbin, EME 0778 - 008 (LWP); 2 ♂♂ 1 ♀, Wirrawilla Rainforest Park, Toolangi, 37 ° 31 ' 39 " S 145 ° 31 ' 17 " E, 1. i. 2023, K. Green, N. & D. Emery & T. Corbin (at light, EME 0778 - 076 – 078) (MM).	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	distribution	Distribution, habitat and seasonality. (Plate 8 E – F; Fig. 1). This species is often the first to emerge in eastern Melbourne, usually around late September or early October and can persist until late January. It favours the upper branches of Eucalyptus regnans (Mountain Ash) at altitudes> 600 m, and specimens are currently restricted to the Dandenong and Yarra ranges east of Melbourne. Since males appear to call constantly at rest from great heights, Y. corbinorum sp. nov. have only been captured emerging around 2100 h on shrubs, ferns and grasses close to stands of Mountain Ash. However, they are readily attracted to light, appearing in large numbers at Wirrawilla Reserve in late December 2022.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	etymology	Etymology. Noun plural; named after Drs. Samantha (nee Emery) and Thomas Corbin who have collected and recorded the majority of specimens of Y. corbinorum sp. nov., and an impressive range of other Victorian and NSW cicadas (especially Yoyetta species) over the past decade.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	description	Description of adult male. (Plate 7 A – B, E, 8 A – C, Fig. 9). Head wider than lateral margins of pronotum; mostly black with a dull brown triangular spot between lateral ocelli based on posterior margin, brown along margins of eyes in lighter specimens; supra-antennal plates black; ocelli light brown; postclypeus burnt brown dorsally, with a lighter brown stripe on midline, black below, posterior margin brown, brown spots on anterior and central midline; lorum orange-brown, gena black, anteclypeus black, rostrum black, reaching between mid- and hind coxae; antennae black. Thorax mostly black in most specimens, brown markings variably prominent across specimens. Pronotum with variable dark brown patches, green-brown laterally, a dark brown fascia along anterior midline abutting posteriorly with a black circular to diamond structure impinging on the pronotal collar centrally; pronotal collar brown, narrow centrally, brown with black along lateral margins. Mesonotum black, with dull to prominent brown “ W ” around submedian sigilla, following parapsidal sutures anteriorly, thorax brown laterally; cruciform elevation black along arms, lateral depressions brown; wing grooves black. Metanotum black, dark brown along ridge, bright orange on anterior margin. Thorax below mainly black with dark brown anepisternum 2 and epimeral lobes edged orange-red around mid and hind leg cavities. Legs black with orange-red striped markings, variable between individuals. Coxae black with reddish ridges, trochanter variably reddish-orange; fore femora brightly striped red-black, black along femoral spines; mid and hind femora black with single reddish stripe; tarsi striped red-black, lighter on hind tarsi; claws dark brown to black; meracantha black with red edges, marginally overlapping opercula. Wings with fusion of forewing veins M and CuA around half length of basal cell; forewing costal veins reddish-brown, pterostigma with orange-brown mottling; basal membranes bright orange-red; basal cell translucent; veins mainly dark brown to black, with eight apical cells; hind wing plaga pale orange-grey to cream at base and margins through anal cell and jejum, becoming transparent towards central third, with six apical cells. Opercula (Plate 8 A) medium, spatulate, following body axis ventrolaterally, depressed centrally; black over basal half, cream over remainder; clearly separated. Timbals (Plate 8 B) with four complete long ribs spanning the timbal membrane, fused dorsally on basal spur; rib 4 broken centrally; rib 5 short, detached, occupying dorsal half of timbal only; timbal plate with well developed, large ridged dome on posterior timbal plate grey, extending across two-thirds of timbal; apodeme pit oval-shaped and conspicuous. Abdomen with tergite 1 black; tergite 2 black with variable brown posterior margin; tergites 3 - 7 black with prominent red-brown to orange posterior margins, increasing on tergite 7, extending laterally and expanding along margins of epipleurites; epipleurites flexed ventrally, black with orange posterior margins; tergite 8 black, with brown posterior margin, a brown spot on lateral surface (Plate 8 C). Sternite I black; sternite II black, dull orange posteriorly; sternites III-VII bright orange anteriorly, fading to yellow posteriorly, with a central black spot black spot variably present on sternites III-VI; sternite VIII dull orange-black with yellowish pubescence. Genitalia (Fig. 9). Pygofer light brown to black; apical spine orange-brown, anal styles light orange; upper and lower lobes dark brown. Pygofer upper lobe peaked at apex; basal pygofer lobe small weakly developed, broadly rounded. Uncus black, median lobe well developed in lateral view ovoid stumpy, lobes in ventral view rounded laterally; claspers prominent, divided, drop-like, with apices gradually tapering laterally. Aedeagus with pseudoparameres not extending as far as theca; theca recurved ventrally at 180 ° towards apex, with broad transparent flanges along margin of recurvature, these smooth dorsally, strongly serrated ventrally,> 2 x diameter of theca, with marked dorsal ornamentation adjacent to apex of theca; apex grooved on internal surface, curved dorsally around 60 o towards a knob-like tip, transparent, sclerotised, with prominent cornuti on dorsal and ventral faces. Description of adult female (Plates 7 C – D, F, 8 D). Generally similar morphology to male, but larger, strikingly coloured mottled brown to more greenish-brown, especially in newly-emerged specimens. Head similar to male, ventral postclypeus with more orange-red markings. Thorax conspicuously brown to ochraceous green-brown, pronotal collar green to brown with central black spot; black along parapsidal sutures; sigilla ranging from light brown to olive green; scutal depression black. Ventral thorax variably ochraceous. Abdomen with more prominent and widespread red-brown markings on tergites, black coloration restricted to lateral margins (Plate 8 D). Tergite 8 brown to olive green, with lateral black spot; variable burnt orange markings on epipleurites 3 – 7. Sternite I mainly green to dark brown; sternites II – VII ranging from burnt to dull orange with small black markings of variable density along midline; abdominal segment 9 black with dorsomedial, brown to green stripe, expanding laterally near posterior margin, stigma dark brown to black, dorsal beak black. Anal styles black; ovipositor sheath dark brown to black; ovipositor brown, tending black towards apex, reaching apex of abdomen. Wings with greenish-brown costa, some specimens with R + Sc and RA green to blue (Plate 7 F); veins CuA and CuP + 1 A blue to green in most specimens (which fades to brown in stored specimens); hindwing plaga tinged orange-grey. Measurements (in mm; range with mean in parentheses for 15 males and 10 females, including smallest and largest specimens). Length of body including head: male 21.5 – 23.0 (22.3); female 22.7 – 25.2 (24.2). Length of forewing: male 25.0 – 28.2 (27.3); female 27.8 – 31.3 (30.3). Width of forewing: male 7.9 – 9.4 (8.5); female 8.4 – 9.8 (9.4). Width of head (including eyes): male 5.8 – 6.5 (6.0); female 6.1 – 7.4 (6.7). Width of pronotum (across lateral angles): male 5.7 – 6.3 (6.1); female 6.0 – 6.9 (6.6). Width of abdomen (across second segment): male 5.8 – 6.5 (6.1); female 6.0 – 7.2 (6.9). Ovipositor length: 6.5 – 7.2 (6.9). Morphological variation within species. The morphology and coloration of male Yoyetta corbinorum sp. nov. are consistent amongst specimens collected to date. Freshly emerged females exhibit principally ochraceous green coloration across the dorsal head, thorax and abdomen, which evolves into more orange-brown with time after eclusion, also allowing the annular banding on the abdomen to become more distinct. It was noted that females from the Dandenong Ranges had less prominent black markings along the ventral midline than those from Wirrawilla. Morphological distinguishing features. Interestingly, at an altitude> 500 m, Y. corbinorum sp. nov. cohabits with Y. grandis Emery, Emery & Popple Y. subalpina Emery, Emery & Popple, Y kershawi Goding & Froggatt and Y. hunterorum Moulds, all of which express prominent white hindwing plaga. In contrast, Y. corbinorum sp. nov. does not cohabit with any currently described Yoyetta species that have BLs <20.6 mm, and hindwing v-shaped plagas tinged orange-grey visible along vein 2 a and 3 a and within the anal cell. Males of Y. corbinorum sp. nov. can be differentiated from other Yoyetta species by 6 characters: (1) the absence of prominent white hindwing plaga; (2) BL> 20.6 mm; (3) a highly serrated, knob-like thecal apex; (4) a relatively short medial lobe of the uncus; (5) four timbal ribs with rib 4 discontinuous centrally; and (6) complete annular, orange-brown, posterior margins of the tergites. Amongst Yoyetta females with typical hindwing v-shaped plagas tinged orange-grey along vein 2 a and 3 a, some 12 species have BLs less than those of female Y. corbinorum sp. nov. (<22.7 mm) Females of this species can be differentiated from the remaining species by the following combination of characters: (1), the greenish coloration of female Y. corbinorum sp. nov.; (2) the extensive red-brown posterior margins of the tergites of Y. corbinorum sp. nov.; (3) small black markings along midline of sternites; and, (4) an ovipositor that does not extend beyond abdominal segment 9.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
8316879B9F64FF83FF128D204638F853.taxon	biology_ecology	Calling song (Fig. 10). The male calling song of Y. corbinorum sp. nov. is a series of repetitive chirps. Further examination of the calling song (all statistics, n = 5 from the Dandenong Ranges) in Fig. 10 illustrates a ‘ chirp’ is a macrosyllable ranging from 0.029 to 0.050 s duration, with a gap between each macrosyllable ranging from 0.214 to 0.245 s duration. The frequency plateau ranges between 8.1 and 14.3 kHz, with a dominant frequency between 9.3 and 10.9 kHz (Fig. 10 C). Males call from around 0800 h and appear to remain at rest for periods approaching 15 - 30 min. Males will still call under cloud cover, even when temperatures fall to around 15 C. Female wing flicking has not been detected except in captivity (approximately 0.05 s after a male chirp), and is likely due to their elevated locations in the treetops. Finger snapping to attract males has been successful only on limited occasions, again likely to the height of their preferred locations. The male calling song of Yoyetta corbinorum sp. nov. most closely resembles those of Y. robertsonae and Y. robusta Popple & Emery and Y. repetens Emery, Emery & Popple. While Y. robertsonae shares an overlapping distribution with Y. corbinorum sp. nov., its call consists of a series of three ticks, whereas Y. corbinorum sp. nov. produces a single macrosyllable. This strident chirping is similar to that of Y. robusta but occurs at a higher frequency (> 9.3 kHz for Y. corbinorum sp. nov. and around 8.3 kHz for Y. robusta), and their distributions do not overlap. Additionally, Y. corbinorum sp. nov. is not known to exhibit call variation when at rest, unlike Y. robusta. When compared with Y. repetens, the macrosyllables of Y. corbinorum sp. nov. are shorter (29 – 50 ms duration) compared with the former and their locations do not overlap.	en	Emery, David L. (2025): Description of four new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from Eastern Australia. Zootaxa 5665 (4): 509-542, DOI: 10.11646/zootaxa.5665.4.2, URL: https://doi.org/10.11646/zootaxa.5665.4.2
