identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
815C87EAFFF15966FF1A0B4FEB32FD10.text	815C87EAFFF15966FF1A0B4FEB32FD10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus dawai Zhang & Rong & Nyima & Zhang & Lin 2025	<div><p>Tanytarsus dawai Lin,  sp. nov.</p><p>urn:lsid:zoobank.org:act: 031BCB5A-D91A-4C28-B79F-06BB3D4FA5FE</p><p>(Figs 2–10)</p><p>Type material.   Holotype: male (SHOU &amp; BOLD sample ID: XL157), China, Xizang, Xigaze City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.9341&amp;materialsCitation.latitude=29.31684" title="Search Plazi for locations around (long 88.9341/lat 29.31684)">Nilun Village</a>, 29.31684°N, 88.93410°E, 3798 m a.s.l., sweep net, 22.VII.2014, leg. X.L. Lin.   Paratypes: female (SHOU &amp; BOLD sample ID: XL50), China, Xizang, Chamdo City, Ranwu Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76048&amp;materialsCitation.latitude=29.50581" title="Search Plazi for locations around (long 96.76048/lat 29.50581)">Ranwu Lake</a>, 29.50581°N, 96.76048°E, 3963 m a.s.1., sweep net, 10.VII.2014, leg. X.L. Lin;   2 males (SHOU &amp; BOLD sample ID: XL117, XL118), China, Xizang, Shannan City, Gongga County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76048&amp;materialsCitation.latitude=29.50581" title="Search Plazi for locations around (long 96.76048/lat 29.50581)">Gangdui Town</a>, 29.27888°N, 90.82323°E, 3586 m a.s.1., sweep net, 18.VII.2014, leg. X.L. Lin;   4 males (SHOU: XL8, XL10, XL11; SHOU &amp; BOLD sample ID: XL9), China, Xizang, Shannan City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76048&amp;materialsCitation.latitude=29.50581" title="Search Plazi for locations around (long 96.76048/lat 29.50581)">Yamzho Yumco</a>, 29.18879°N, 90.56003°E, 4458 m a.s.l., sweep net, 18.VII.2014, leg. X.L. Lin;   5 males (SHOU &amp; BOLD sample ID: XL48; SHOU: XL49, XL51, XL52, XL53), China, Xizang, Chamdo City, Ranwu Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76048&amp;materialsCitation.latitude=29.50581" title="Search Plazi for locations around (long 96.76048/lat 29.50581)">Ranwu Lake</a>, 29.50581°N, 96.76048°E, 3963 m a.s.l., sweep net, 10.VII.2014, leg. X.L. Lin;   6 larvae (SHOU &amp; BOLD sample ID: XL431, XL432, XL434, XL435, XL437, XL440), China, Xizang, Chamdo City, Ranwu Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76048&amp;materialsCitation.latitude=29.50581" title="Search Plazi for locations around (long 96.76048/lat 29.50581)">Ruanwu Lake</a>, 29.50581°N, 96.76048°E, 3963 m a.s.l., 10.VII.2014, leg. X.L. Lin.</p><p>Etymology. The specific epithet is dedicated to Associate Professor Dawa from the Tibet Plateau Institute of Biology in Lhasa, Xizang, China, for his outstanding contribution to the knowledge of insects in Xizang; noun in genitive case.</p><p>Diagnostic characters. The adult male of the new species can be distinguished from known species of the  Tanytarsus norvegicus species group by the following combination of characters: Tergite I-V with horizontal dark brown bands at the posterior quarter. AR 0.86–1.10. Ac absent. LR 1 1.55–1.71. Tarsomere l of mid leg with sensilla chaetica distally. Anal point slender triangular, with six lateral setae on each side, 8–11 spinulae evenly placed at 2/3 length of anal point. Superior volsella sub-rectangular with ventral, small patch of microtrichia. Digitus bent downwards. Stem of median volsella long, with six setiform lamellae and one distal, spatulate lamella. The larva of the new species can be distinguished from known species of the  T. norvegicus species group by the following combination of characters: AHR 0.15–0.21. LOR 1.92–2.63. Mandible with one ventral apical tooth, three ventral inner teeth, four dorsal teeth. Mentum with 11 teeth; median tooth with small lateral notches. Premandible with five teeth in addition to lateral spine; premandibular brush well-developed.</p><p>Description. Adult male (n = 13).</p><p>Total length 3.68–4.08, 3.84 mm; wing length 2.25–2.93, 2.46 mm; total length/wing length 1.39–1.66, 1.56.</p><p>Colouration (Figs 2B; 3A–B). Head and legs brown. Thorax and abdomen with pale brown ground colour. Ventral part of parapsidal suture, postnotum, preepisternum and median anepisternum II dark brown, scutum and epimeron II light brown. Tergite I–V with horizontal dark brown bands at the posterior quarter. Tergite VI with horizontal light brown band at the posterior seventh.</p><p>Head (Fig. 3A). AR 0.86–1.10, 0.94. Antenna with 13 flagellomeres, ultimate flagellomere 500–619, 564 μm long. Frontal tubercles 25–38, 31 μm long, 8–13, 10 μm wide at base. Temporal setae 8–12, 10. Clypeus with 10–15, 13 setae. Tentorium 145–176, 156 μm long, 33–50, 39 μm wide. Palpomere lengths (μm): 35–40, 39; 38–63, 51; 102–140, 122; 118–145, 129; 200–237, 218.</p><p>Thorax (Fig. 3B). Ac absent; Dc 7–10, 8; Pa 1; Scts 5–8, 6. Haltere with 4–5, 4 setae.</p><p>Wing (Fig. 3C). VR 1.08–1.18, 1.11. Brachiolum with l seta, R with 8–21, 15 setae, R 1 with 6–15, 10 setae, R 4+5 with 7–28, 16 setae, M 1+2 with 3–57, 28 setae, M 3+4 with 1–11, 5 setae, false vein with 0–3 setae (usually 0), Cu with 0–6, 1 seta, Cu 1 with 0–3, 1 seta, r 4+5 with 79–115, 96 setae, m 1+2 with 14–117, 60 setae, m 3+4 with 0–17, 3 setae, remaining veins and cells bare.</p><p>Legs. Fore leg bearing single tibial spur 26–43, 34 μm long. Combs of mid tibia 11–21, 16 μm wide with 25–33, 30 μm long spur, and 10–20, 15 μm wide with 23–30, 27 μm long spur; combs of hind tibia 15–28, 22 μm wide with 28–38, 33 μm long spur, 17–27, 22 μm wide with 23–35, 31 μm long spur. Tarsomere l of mid leg with 4–7, 5 sensilla chaetica distally. Lengths and proportions of legs as in Table 1.</p><p>Hypopygium (Figs 4–5). Tergite IX 100–125, 113 μm long. Anal tergite bands V-shaped, well separated, ending at mid-length of tergite. Anal point (Fig. 5) 48–67, 56 μm long, slender triangular, tapering near the apex, with six lateral setae on each side, 8–11, 9 spinulae evenly placed at 2/3 length of anal point. Transverse sternapodeme 62–91, 78 μm long, with oral projections. Phallapodeme 90–125, 101 μm long. Gonocoxite 152–177, 165 μm long. Gonostylus 140–173, 159 μm long. Superior volsella (Fig. 4D) sub-rectangular, with concave inner margin, bearing seven dorsal setae and two ventral setae, ventral surface with small patch of microtrichia at posterior margin. Digitus long and bent downwards, curved, with swollen blunt apex, extending beyond the inner margin of superior volsella. Stem of median volsella 20–25, 23 μm long, with 6 setiform lamellae and one distal spatulate lamella (Fig. 4E). Inferior volsella weakly curved, 100–128, 111 μm long, with 11–23, 19 strong setae. HR 0.94–1.18, 1.04; HV 2.36–2.66, 2.51.</p><p>Adult female (n = 1)</p><p>Total length 4.29 mm. Wing length 2.98 mm. Total length/wing length 1.44.</p><p>Colouration. As in male, the colour of scutum, parapsidal suture and median anepisternum II lighter. Tergite I–II light brown with horizontal dark brown bands at the posterior quarter.</p><p>Head (Fig. 6A). Antenna (Fig. 6B) with four flagellomeres, flagellomere length (in μm): 117, 75, 79, 214. AR 0.79. Frontal tubercles 14 μm long, 18 μm wide at base. Temporal setae 8. Clypeus with 17 setae. Tentorium 141 μm long, 26 μm wide. Palpomere lengths (in μm): 261, 144, 134, 62, 42.</p><p>Thorax (Fig. 6C). Ac absent; Dc 15; Pa 1; Scts 3. Halteres with 1 seta.</p><p>Wing (Fig. 6D). VR 1.28. Brachiolum with one seta, R with 11 setae, R 1 with 32 setae, R 4+5 with 64 setae, M 1+2 with 38 setae, M 3+4 with 41 setae, Cu with 20 setae, Cu 1 with 24 setae, false vein with 50 setae, An with 17 setae, remaining veins bare. Cell r 4+5 with 180 setae, m 1+2 with 88 setae, m 3+4 with 150 setae, cu + an with 61 setae, remaining cells bare. Anal lobe strongly reduced.</p><p>Legs. Foreleg bearing single tibial spur 38 μm long. Combs of mid tibia 13 μm wide with 34 μm long spur, and 14 μm wide with 36 μm long spur; combs of hind tibia 16 μm wide with 42 μm long spur, 20 μm wide with 38 μm long spur. Tarsomere l of mid leg with 28 sensilla chaetica distally. Lengths and proportions of legs as in Table 2.</p><p>Genitalia (Fig. 7). Tergite IX semicircular with 22 setae; sternite VIII with 36 setae; gonapophysis VIII single lobe with long posteromedially directed microtrichia; gonocoxapodeme 124 μm wide, slightly curved; coxosternapodeme well-developed with distinct anterior and posterior lobes. Notum including rami 134 μm long, notum alone 36 μm long. Seminal capsules semi-circular, 63 μm long, 47 μm wide. Postgenital plate triangular. Cercus 96 μm long, with 29 setae.</p><p>Pupa. Unknown.</p><p>Larva (n = 4)</p><p>Total length 5.5–7.2, 6.5 mm.</p><p>Head Capsule (Fig. 8). Length 346–396, 376 μm, width 341–370, 355 μm, length/width 0.99–1.16, 1.06; mental and mandibular teeth dark brown. Antennal pedestal 54–78, 68 μm long (Fig. 9A). AHR 0.15–0.21, 0.18; AAR 0.53–0.74, 0.64. Antenna shorter than head length, 5-segmented, only first and second antennal segments well sclerotized. Antennal segment length (in μm): 101–106, 104; 20–25, 23; 9–13, 12; 4–7, 6; 3–5, 4. AR 2.06–2.81, 2.34. Antennal blade 14 μm long. LO stems + stylus 42–51, 47 μm long; LOR 1.92–2.63, 2.19. Mandible (Fig. 9 B-C) 149–153, 151 μm long, 43–50, 47 μm wide, 1 ventral apical tooth, 3 ventral inner teeth, 4 dorsal teeth and one robust dorsal plate; seta subdentalis thick and curved, 44–55, 50 μm long, reaching beyond apex of dorsal tooth; seta interna well–developed with 4 main branches. Mentum (Fig. 9 D-E) 98–115, 107 μm wide, with 11 teeth; median tooth with small lateral notches or absent; ventromental plate 122–128, 126 μm wide, 22–28, 26 μm long, MVR 0.80–0.91, 0.85. Premandible (Fig. 10 A-B) 78–81, 80 μm long, with 5 teeth in addition to lateral spine; premandibular brush well-developed. Labrum (Fig. 10C) with S I pectinate and chaetae plumose; S II long, simple; clypeal seta S3 long, simple. Labral lamella pectinate. Pecten epipharyngis consisting of three plumose lobes. Maxilla (Fig. 10D) with two long and one short lacinial chaetae, palp normally developed, postoccipital margin darkly pigmented. Postmentum 156–165, 162 μm long.</p><p>Abdomen. Procercus 40–54, 47 μm high, with 8 anal setae, 551–569, 560 μm long. Anal tubules two pairs,156– 180, 168 μm and 108–110, 109 μm long.</p><p>Remarks. The new species is closely related to  Tanytarsus sinuatus in the shape of hypopygium, but can be separated from the latter species by the following characters of adult males: LR 1 1.55–1.71, frontal tubercles longer, tarsomere l of mid leg with sensilla chaetica distally, superior volsella more rectangular, digitus curved and without a protuberance at mid length, median volsella has one distal spatulate lamella, and anal point has more spines in the new species. The larvae of the new species exhibit a  Tanytarsus glabrescens - type mandible, characterized by the presence of 2+2 dorsal teeth (Eggermont 2008), similar to the mandible morphology observed in the larvae of  T. gracilentus and  T. lapponicus (Stur &amp; Ekrem 2011) . However, the dorsal plate is smaller compared to those of these latter two species.</p><p>Ecological notes. The larvae of the new species were found in glacier lakes of Qinghai-Tibet Plateau.</p><p>Distribution. China (Xizang).</p></div>	https://treatment.plazi.org/id/815C87EAFFF15966FF1A0B4FEB32FD10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zhang, Zhi-Chao;Rong, Yue-Xiao;Nyima, Tenzin;Zhang, Rui-Lei;Lin, Xiao-Long	Zhang, Zhi-Chao, Rong, Yue-Xiao, Nyima, Tenzin, Zhang, Rui-Lei, Lin, Xiao-Long (2025): Tanytarsus dawai sp. nov. and larval morphology of T. sinuatus within the T. norvegicus species group (Diptera: Chironomidae). Zootaxa 5633 (2): 316-335, DOI: 10.11646/zootaxa.5633.2.5, URL: https://doi.org/10.11646/zootaxa.5633.2.5
815C87EAFFFD5979FF1A0FC1EDA2FCAC.text	815C87EAFFFD5979FF1A0FC1EDA2FCAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus sinuatus Goetghebuer 1936	<div><p>Tanytarsus sinuatus Goetghebuer, 1936</p><p>(Figs 11–14)</p><p>Tanytarsus sinuatus Goetghebuer, 1936 in Thienemann 1936: 185.</p><p>Tanytarsus alpicola Goetghebuer, 1950: 5 .</p><p>Tanytarsus sinuatus Reiss &amp; Fittkau 1971: 134 .</p><p>Material examined.  Holotype: male (RBINS: 18.073), Germany, Bavaria, 1935, leg. et det. M. Goetghebuer.  3 males (SHOU &amp; BOLD sample ID: XL676, XL678, XL679), Italy, Trentino-Alto Adige/Sudtirol, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Trento</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Peio</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Stilfser Joch National Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Glacier Lake below the Lago Marmotta</a>, 46.434723°N, 10.677778°E, 2633 m, a.s.l., sweep net, 8.VII.2017, leg. X.L. Lin.   3 larvae (SHOU &amp; BOLD sample ID: XL55, XL438, SHOU sample ID: XL56), China, Xizang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Chamdo City</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Ranwu Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Ruanwu Lake</a>, 29.50581°N, 96.76048°E, 3963 m a.s.l., 10.VII.2014, leg. X.L. Lin;   3 larvae (SHOU &amp; BOLD sample ID: XL3586, XL3587, XL3593), China, Qinghai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Bande Lake</a>, 34.2161°N, 92.0769°E, 4600 m a.s.l., 12.VI.2020, leg. Jia-Ping Chen;   2 larvae (SHOU &amp; BOLD sample ID: XL2722, XL2723), China, Yunnan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Nu jiang of the Lisu Autonomous Prefecture</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Lanping Bai</a> and Pumi Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Erdaobaihe Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Glacial Lake</a>, 26.5897222°N, 99.0213899°E, 3789 m a.s.l., 24.V.2018, leg. X.L. Lin;   1 larva (SHOU &amp; BOLD sample ID: XL3843), China, Xizang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Naqu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Nima Wetland</a>, 31.7128°N, 92.0582°E, 4628 m a.s.l., 5.IX.2020, leg. Yu Peng;   1 larva (SHOU &amp; BOLD sample ID: XL3828), China, Xizang, Nyingchi City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.742&amp;materialsCitation.latitude=29.7617" title="Search Plazi for locations around (long 94.742/lat 29.7617)">Bayi</a>, 29.7617°N, 94.742°E, 3332 m a.s.l., 28.IIX.2020, leg. Yu Peng.</p><p>Diagnostic characters. The adult male of  Tanytarsus sinuatus can be distinguished from known species of the  T. norvegicus species group by the following combination of characters: Frontal tubercles (Fig. 11B) cylindrical. Thorax (Fig. 11C) pale brown with dark brown on scutum and parapsidal suture, light brown on median anepisternum II, preepisternum and epimeron II. Ac absent. LR 1 1.48–1.50. Anal point (Fig. 11D) slender triangular, tapering near the apex, with 9–13 spinulae in single row between large parallel crests. Superior volsella (Fig. 11E) roughly slender elliptical in outline but extended into a robust tooth on the anal median side. Digitus (Fig. 11F) is thickened distally, significantly surpassing superior volsella and bears a small, well-discernible protuberance or lobe at mid length. Stem of median volsella (Fig. 11F) clublike thickened distally, with several lamellae of the broad awl-type. Inferior volsella (Fig. 11F) distal part round head-like.</p><p>Larva (n = 5)</p><p>Total length 5.4–6.8, 6.1 mm.</p><p>Head Capsule (Fig. 12B). Length 357–423, 385 μm, width 322–378, 350 μm, length/width 1.05–1.14, 1.10; mental and mandibular teeth dark brown. Antennal pedestal (Fig. 13 A-B) 53–59, 57 μm long.AHR 0.13–0.17, 0.15; AAR 0.48–0.62, 0.53. Antenna shorter than head length, 5-segmented. Antennal segment length (in μm): 93–117, 108; 28–36, 32; 8–13, 11; 6–8, 7; 3–6, 5. AR 1.74–2.34, 1.97. Antennal blade 25–37, 31 μm long. LO stems + stylus 32–43, 36 μm long; LOR 1.44–1.70, 1.57. Style 6–9, 8 μm long. Mandible (Fig. 13 C-D) 135–156, 148 μm long, 54–69, 62 μm wide, one ventral apical tooth, three ventral inner teeth, 4 dorsal teeth and 1 robust dorsal plate; seta subdentalis thick and curved, 51–59, 56 μm long, reaching beyond apex of dorsal tooth; seta interna well-developed with four main branches. Mentum (Fig. 13 E-F) 106–122, 111 μm wide, with 11 teeth; median tooth lingulate with small lateral notches; ventromental plate 117–132, 125 μm wide, 26–32, 29 μm long, MVR 0.85–0.93, 0.89. Premandible (Fig. 14 A-B) 76–88, 83 μm long, with five teeth in addition to lateral spine; premandibular brush well-developed. Labrum (Fig. 14C) with S I pectinate and chaetae plumose; S II long, simple; clypeal seta S3 long, simple. Labral lamella pectinate. Pecten epipharyngis consisting of three plumose lobes. Maxilla (Fig. 14D) with two long and one short lacinial chaetae, palp well-developed, postoccipital margin darkly pigmented. Postmentum 154–171, 163 μm long.</p><p>Abdomen. Procercus 48–52, 51 μm high, with eight anal setae, 530–568, 550 μm long. Anal tubules two pairs,130–195, 162 μm and 107–132, 114 μm long.</p><p>Remarks: The larvae of  Tanytarsus sinuatus and  T. dawai are very similar, but can be distinguished by following morphological characters:  T. dawai has a higher LOR (1.92–2.63), whereas LOR = 1.44–1.70 in  T. sinuatus . In addition, the median tooth on the mentum exhibits a rounded shape in  T. dawai, whereas it bears lateral protrusions in  T. sinuatus .</p><p>Distribution. Austria, China (Qinghai, Xizang, Yunnan), Germany, Italy and Norway.</p></div>	https://treatment.plazi.org/id/815C87EAFFFD5979FF1A0FC1EDA2FCAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zhang, Zhi-Chao;Rong, Yue-Xiao;Nyima, Tenzin;Zhang, Rui-Lei;Lin, Xiao-Long	Zhang, Zhi-Chao, Rong, Yue-Xiao, Nyima, Tenzin, Zhang, Rui-Lei, Lin, Xiao-Long (2025): Tanytarsus dawai sp. nov. and larval morphology of T. sinuatus within the T. norvegicus species group (Diptera: Chironomidae). Zootaxa 5633 (2): 316-335, DOI: 10.11646/zootaxa.5633.2.5, URL: https://doi.org/10.11646/zootaxa.5633.2.5
