identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
444E414F87A25C0BACB03BA9F379AF08.text	444E414F87A25C0BACB03BA9F379AF08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clausilia plicata var. elongata Roffiaen 1868	<div><p>Clausilia plicata var. elongata Roffiaen, 1868</p><p>Fig. 3 A</p><p>Clausilia plicata var. elongata Roffiaen, 1868: 75, pl. 1, fig. 4.</p><p>Type material.</p><p>RBINS I.G.9154 / MT.1771: 3 possible syntypes, ex de Cort coll.</p><p>Type locality.</p><p>Switzerland:  Andeer .</p><p>Taxonomic status.</p><p>Junior synonym of  Laciniaria plicata plicata (Draparnaud, 1801) .</p><p>Discussion.</p><p>According to the specimens’ labels, they were collected at the type locality Andeer and stem from the collection of Hugo de Cort. Furthermore, one of the labels states “ types ”, but there is no mention of Roffiaen or his collection. Thus, the present specimens are here considered as possible syntypes.</p><p>Laciniaria plicata plicata is widely distributed in Central Europe, including Switzerland (Turner et al. 1998; Nordsieck 2008). The elongated spire of Roffiaen’s variety is well within the known variation in shell shape and size known in L. p.  plicata (Nordsieck 2008) .</p></div>	https://treatment.plazi.org/id/444E414F87A25C0BACB03BA9F379AF08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
2EEF7D401C855896AB5DAC575189742F.text	2EEF7D401C855896AB5DAC575189742F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clausilia plicatula var. albinos Roffiaen 1868	<div><p>Clausilia plicatula var. albinos Roffiaen, 1868</p><p>Clausilia plicatula var. albinos Roffiaen, 1868: 76.</p><p>Type material.</p><p>Single specimen, at present not located.</p><p>Type locality.</p><p>Switzerland: Iseltwald.</p><p>Taxonomic status.</p><p>Junior synonym of  Macrogastra plicatula plicatula (Draparnaud, 1801) .</p><p>Discussion.</p><p>No specimens belonging to this taxon were found. The original description mentioned a single shell, whitish and transparent (hence, “ albinos ”), as belonging to this variety, while commenting that the nominate form is common and very variable (Roffiaen 1868: 76). Thus, this variety based on an albino specimen (or otherwise presenting partial or total loss of pigments) can be considered synonymous with the nominate form.  Macrogastra plicatula is widespread in Europe, including many records in the area encompassing Iseltwald representing M. p.  plicatula (Turner et al. 1998; Nordsieck 2006).</p></div>	https://treatment.plazi.org/id/2EEF7D401C855896AB5DAC575189742F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
3FB197936A495DC19DBB8DC6D08CE0FB.text	3FB197936A495DC19DBB8DC6D08CE0FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clausilia weyersi Roffiaen 1868	<div><p>Clausilia weyersi Roffiaen, 1868</p><p>Fig. 3 B, Suppl. material 1</p><p>Clausilia weyersi Roffiaen, 1868: 75, pl. 1, fig. 3.</p><p>Type material.</p><p>RBINS I.G.9154 / MT.1773: 2 syntypes, ex Roffiaen coll., ex de Cort coll. RBINS I.G.9154 / HIST.2486: 29 possible syntypes, ex de Cort coll.</p><p>Type locality.</p><p>Switzerland: Weesen. “  Wesen ” (Roffiaen 1868: 75).</p><p>Taxonomic status.</p><p>Junior synonym of  Laciniaria plicata plicata (Draparnaud, 1801) .</p><p>Discussion.</p><p>Lot RBINS I.G.9154 / MT.1773 comes from the type locality, belonged to Roffiaen’s collection, and bears the writing “ types ”; its specimens are considered syntypes. Lot RBINS I.G.9154 / HIST.2486, however, while coming from the type locality, has no indication to have belonged to Roffiaen’s collection or of being types, though they belonged to de Cort’s collection. Therefore, the later specimens are considered possible syntypes.</p><p>Clausilia weyersi bears close resemblance to typical specimens of  Laciniaria plicata plicata, both regarding shell shape, size and apertural barriers (Nordsieck 2008; see Suppl. material 1 for CT scan snapshots of internal structures of shell). It can thus, be considered a junior synonym of the latter.</p></div>	https://treatment.plazi.org/id/3FB197936A495DC19DBB8DC6D08CE0FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
3F582E7AF3FC584AB386086C250D300F.text	3F582E7AF3FC584AB386086C250D300F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helix arbustorum var. icterica Roffiaen 1868	<div><p>Helix arbustorum var. icterica Roffiaen, 1868</p><p>Fig. 4 A</p><p>Helix arbustorum var. icterica Roffiaen, 1868: 70.</p><p>Type material.</p><p>RBINS I.G.7065 / MT.3421: 1 syntype from  Weesen, ex Colbeau coll. (nr. 7065)  .</p><p>Type locality.</p><p>Switzerland: Tamina, Weesen, Iseltwald,  Meiringen,  Sarnen . “ Gorge de la Tamina, Wesen, Iseltwald, Meyringen, Sarnen ” (Roffiaen 1868: 70).</p><p>Taxonomic status.</p><p>Junior synonym of  Arianta arbustorum (Linnaeus, 1758) .</p><p>Discussion.</p><p>The present specimen stems from the collection of Jules Colbeau, is from one of the localities that constitute the taxon’s type locality as mentioned by Roffiaen (1868) and the label bears the writing “ type ”. Thus, it can be considered a syntype.</p><p>This variety was described to denote specimens lacking the typical colour pattern of nominate  A. arbustorum, consisting instead of entirely pale-yellow shells. Roffiaen also mentioned that such specimens were “ not common but not rare either ” (Roffiaen 1868: 70). This type of colouration is often seen in some members of this species across its range, consistent with natural interspecific phenotypic variation and sometimes is more common at some localities than others (e. g., Grime and Blythe 1969; Parkin 1971; Arter 1990; Cameron and Riley 2008; von Proschwitz et al. 2023). Thus, A.  A. icterica is here considered synonymous with  A. arbustorum . The variability in shell shape, size and colour of  A. arbustorum has led to many forms, varieties, and subspecies to be described in the past centuries, which have almost always been considered junior synonyms (e. g., Taylor 1914).</p></div>	https://treatment.plazi.org/id/3F582E7AF3FC584AB386086C250D300F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
3FF1EC14161B55779931BC58DDB06A95.text	3FF1EC14161B55779931BC58DDB06A95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helix arbustorum var. marmorata Roffiaen 1868	<div><p>Helix arbustorum var. marmorata Roffiaen, 1868</p><p>Helix arbustorum var. marmorata Roffiaen, 1868: 70.</p><p>Type material.</p><p>Not located.</p><p>Type locality.</p><p>Switzerland:  Weesen . “ Se trouve en diverses localités, n’est pas rare à Wesen ” (Roffiaen, 1868: 70)  .</p><p>Taxonomic status.</p><p>Junior synonym of  Arianta arbustorum (Linnaeus, 1758) .</p><p>Discussion.</p><p>This variety was described to denote specimens lacking the dark spiral bands seen on the shells of typical  Arianta arbustorum . As for the case of the yellow shells of the  variety icterica above, this is part of the common colour polymorphism known in  A. arbustorum (Cook and King 1966; Grime and Blythe 1969; Burla and Stahel 1983; Burla 1984; von Proschwitz et al. 2023). Thus, even without having access to type material, Roffiaen’s (1868) description, along with current knowledge on  A. arbustorum, is sufficient to surmise that A.  A. marmorata is synonymous with  A. arbustorum .</p></div>	https://treatment.plazi.org/id/3FF1EC14161B55779931BC58DDB06A95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
9EBC9B42B9375C5B982EE560972BB696.text	9EBC9B42B9375C5B982EE560972BB696.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helix arbustorum var. trochoidalis Roffiaen 1868	<div><p>Helix arbustorum var. trochoidalis Roffiaen, 1868</p><p>Fig. 4 B</p><p>Helix arbustorum var. trochoidalis Roffiaen, 1868: 69, pl. 1, fig. 2.</p><p>Type material.</p><p>RBINS I.G.9154 / MT.4172 and I.G.9154 / MT.4171: respectively, 7 and 6 possible syntypes, ex de Cort coll.</p><p>Type locality.</p><p>Switzerland:  Andeer .</p><p>Taxonomic status.</p><p>Junior synonym of  Arianta arbustorum (Linnaeus, 1758) .</p><p>Discussion.</p><p>The two available specimen lots were collected at the type locality Andeer and stem from the collection of Hugo de Cort. There is no indication that they came from Roffiaen’s collection, but the specimens are a good conchological match to Roffiaen’s brief description and figures (Roffiaen 1868: pl. 1, fig. 2 a, b). Thus, we interpret the present specimens as possible syntypes.</p><p>This variety was described to denote specimens with an elongated spire, which gives the shell a more trochoid shape. As mentioned above for shell colouration, intraspecific variation in shell shape (including spire height) is common across members of  A. arbustorum (e. g., Germain 1930; Baminger 1997). Thus,  Helix arbustorum trochoidalis is here considered synonymous with  A. arbustorum .</p></div>	https://treatment.plazi.org/id/9EBC9B42B9375C5B982EE560972BB696	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
1B0E640C54CB57CEB3D9E3320CEA699E.text	1B0E640C54CB57CEB3D9E3320CEA699E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helix ruderata var. viridana Roffiaen 1868	<div><p>Helix ruderata var. viridana Roffiaen, 1868</p><p>Helix ruderata var. viridana Roffiaen, 1868: 68.</p><p>Type material.</p><p>Not located.</p><p>Type locality.</p><p>Switzerland: Handeck ( Handegg). “ La Handeck ” (Roffiaen 1868: 68)  .</p><p>Taxonomic status.</p><p>Junior synonym of  Discus ruderatus (Hartmann, 1821) .</p><p>Discussion.</p><p>This variety was described to allocate specimens for which the single conchological character noted by Roffiaen (1868) was the “ pale greenish colour ”. The shell of  Discus ruderatus is known to typically to be light brown to reddish brown (Welter-Schultes 2012), though paler yellowish and greenish brown specimens are also common (Kerney et al. 1979; Wiese 2014). Furthermore, the type locality of this variety is within the known distribution of  D. ruderatus in Switzerland (Turner et al. 1998), and Roffiaen (1868) himself mentioned “ typical ”  D. ruderatus occurring in the same place. Thus, even without access to type material, it can be surmised that  Helix ruderata viridana is very likely a junior synonym of  D. ruderatus .</p></div>	https://treatment.plazi.org/id/1B0E640C54CB57CEB3D9E3320CEA699E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
0F5EF3EE443154AE9BEED6112F2E0C8B.text	0F5EF3EE443154AE9BEED6112F2E0C8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lymnaea palustris var. fallaciosa Roffiaen 1868	<div><p>Lymnaea palustris var. fallaciosa Roffiaen, 1868</p><p>Fig. 2 A</p><p>Limnæa palustris var. fallaciosa Roffiaen, 1868: 78, pl. 1, fig. 6.</p><p>Type material.</p><p>RBINS I.G.7065 / MT.4174: 3 syntypes, ex Colbeau coll. RBINS I.G.28242 / HIST.180: 6 possible syntypes .  RBINS I.G.7065 / HIST.181: 1 possible syntype .</p><p>Type locality.</p><p>Switzerland: Brunnen,  along road to Schwyz . “ […] Brunnen, dans un fossé de la plaine, longeant la route de Schwyz ” (Roffiaen 1868: 79).</p><p>Taxonomic status.</p><p>Junior synonym of  Stagnicola palustris (O. F. Müller, 1774) (cf. Servain 1882; Hubendick 1951; Vinarski 2024).</p><p>Discussion.</p><p>Lot RBINS I.G.7065 / MT.4174 stems from the collection of Jules Colbeau, comes from the type locality (Brunnen), and the specimens are a good match to the published illustration (Roffiaen 1868: pl. 1, fig. 6); thus, the three specimens it contains are here considered syntypes.</p><p>Lot RBINS I.G.28242 / HIST.180 stems from the type locality but bear no other information regarding their former owners. The six specimens match morphologically the syntypes in the previous lot and, thus, are considered here as possible syntypes. Similarly, lot RBINS I.G.7065 / HIST.181 has 1 specimen from Colbeau’s collection whose location is given as “ Schwyz ” on the label, together with an indication that this specimen belonged to the study of Roffiaen. While the type locality of the present taxon could be generously interpreted as containing Schwyz, we deem it safer to consider the present specimen as a possible syntype.</p><p>Though conchologically, this variety corresponds well to a widespread  Stagnicola palustris, there is a possibility that Roffiaen’s shells could belong to  S. fuscus (C. Pfeiffer, 1821) instead. The latter can only be distinguished with certainty from  S. palustris by anatomical and / or genetic investigations (Welter-Schultes 2012; Glöer 2019). The range of  S. fuscus covers much of central and western Europe, including Switzerland (Welter-Schultes 2012).</p></div>	https://treatment.plazi.org/id/0F5EF3EE443154AE9BEED6112F2E0C8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
A1C3020837EF5CBD9C708A8B0D4CBB55.text	A1C3020837EF5CBD9C708A8B0D4CBB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lymnaea palustris var. pellucida Roffiaen 1868	<div><p>Lymnaea palustris var. pellucida Roffiaen, 1868</p><p>Fig. 2 B</p><p>Limnæa palustris var. pellucida Roffiaen, 1868: 79, pl. 1, fig. 7.</p><p>Type material.</p><p>RBINS I.G.7065 / MT.4175: 1 holotype, ex Colbeau coll.</p><p>Type locality.</p><p>Switzerland: Altdorf,  in a ditch on the plains towards Seedorf . “ […] dans un fossé de la plaine d’Altorf, vers Seedorf (M. Colbeau) ” (Roffiaen 1868: 79).</p><p>Taxonomic status.</p><p>Junior synonym of  Stagnicola palustris (O. F. Müller, 1774) (cf. Servain 1882; Hubendick 1951; Vinarski 2024).</p><p>Discussion.</p><p>Specimen RBINS I.G.7065 / MT.4175 stems from the collection of Jules Colbeau, comes from the type locality (Altdorf, spelled “ Altorf ” on the label as in the original publication) and is a good match to the published illustration (Roffiaen 1868: pl. 1, fig. 7). Considering the original description was written using a single individual, the present specimen can thus be considered the holotype.</p><p>A possibility that the holotype could belong to  S. fuscus cannot be rejected (see the discussion of  L. palustris var. fallaciosa above).</p></div>	https://treatment.plazi.org/id/A1C3020837EF5CBD9C708A8B0D4CBB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
0B12AA70BC9C54588FF93A760016471D.text	0B12AA70BC9C54588FF93A760016471D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lymnaea peregra var. pulchella Roffiaen 1868	<div><p>Lymnaea peregra var. pulchella Roffiaen, 1868</p><p>Fig. 2 C</p><p>Limnæa peregra var. pulchella Roffiaen, 1868: 77, pl. 1, fig. 8.</p><p>Type material.</p><p>RBINS I.G.7065 / MT.4173: 3 syntypes, ex Colbeau coll.</p><p>Type locality.</p><p>Switzerland: Altdorf,  in a ditch on the plains towards Seedorf . “ […] dans un fossé de la plaine d’Altorf vers Seedorf (M. Colbeau) ” (Roffiaen 1868: 77).</p><p>Taxonomic status.</p><p>Junior synonym of  Peregriana peregra (O. F. Müller, 1774) (cf. Hubendick 1951; Vinarski 2024).</p><p>Discussion.</p><p>The present lot stems from the collection of Jules Colbeau, it comes from the type locality (Altdorf, spelled “ Altorf ” on label as in the original publication), and the specimens are a good match to the published illustration (Roffiaen 1868: pl. 1, fig. 8); the three specimens it contains are here considered syntypes.</p></div>	https://treatment.plazi.org/id/0B12AA70BC9C54588FF93A760016471D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
D8A904670E075E7495108EE167D09A7C.text	D8A904670E075E7495108EE167D09A7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lymnaea stagnalis var. productissima Roffiaen 1868	<div><p>Lymnaea stagnalis var. productissima Roffiaen, 1868</p><p>Limnæa stagnalis var. productissima Roffiaen, 1868: 78: pl. 1, fig. 5.</p><p>Type material.</p><p>Not located.</p><p>Type locality.</p><p>Switzerland: Magadino,  Lago Maggiore . “ Lac Majeur à Magadino ” (Roffiaen 1868: 78)  .</p><p>Taxonomic status.</p><p>Junior synonym of  Lymnaea stagnalis (Linnaeus, 1758) sensu lato (cf. Hubendick 1951; Vinarski 2024).</p><p>Discussion.</p><p>There is only one lot (RBINS I.G.7065 / HIST.372, containing 7 shells) in the collection identified as Roffiaen’s taxon, collected in the type locality (“ Lac Majeur Magadino ” on the label). There is unfortunately no further collection data for this specimen lot, so it cannot be ascertained whether these shells came from Roffiaen. In any event, while the specimens do represent  Lymnaea stagnalis, they are only about half the size of Roffiaen’s illustrated adult specimen (Roffiaen 1868: pl. 1, fig. 5). Consequently, the shell morphology of these specimens does not correspond with Roffiaen’s diagnosis (“ Coguille très-allongée et efilée à dernier tour moins ventru que chez le type ”, i. e., very elongated and tapered shell with body whorl less rounded than in the type; Roffiaen 1868: 78). Thus, these specimens are likely not types of  Lymnaea stagnalis productissima . Being collected from a large lake, the shells most probably belong to a “ lacustrine ” morph of  L. stagnalis sensu lato, which is characterized by reduced body size and shortened spire (see Geyer 1929; Hubendick 1951).</p></div>	https://treatment.plazi.org/id/D8A904670E075E7495108EE167D09A7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
1B63B09542E951EBABC08905F39D01F5.text	1B63B09542E951EBABC08905F39D01F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lymnaea truncatula var. subangulata Roffiaen 1868	<div><p>Lymnaea truncatula var. subangulata Roffiaen, 1868</p><p>Limnæa truncatula var. subangulata Roffiaen, 1868: 78, pl. 1, fig. 9.</p><p>Type material.</p><p>Not located.</p><p>Type locality.</p><p>Switzerland: Altdorf,  in a ditch on the plains towards Seedorf . “ […] prés d’Altorf, dans un fossé de la plaine vers Seedorf (M. Colbeau) ” (Roffiaen 1868: 78)  .</p><p>Taxonomic status.</p><p>Junior synonym of  Galba truncatula (O. F. Müller, 1774) (cf. Hubendick 1951; Vinarski 2024).</p><p>Discussion.</p><p>No specimens belonging to this taxon were found. The original description mentions a few specimens and describe them as being “ contorted and misshapen ” (“ contourné et difforme ”; Roffiaen 1868: 78) and the accompanying illustrations confirm that description. Likely, these specimens are individuals of  Galba truncatula that experienced some problems during growth (e. g., shell breakage and consequent scar and irregular shell growth).</p><p>A species that has been named  Lymnaea (Galba) subangulata in the Russian literature (Kruglov and Starobogatov 1993; Kruglov 2005; Vinarski and Kantor 2016) differs from Roffiaen’s  Limnæa truncatula var. subangulata . The shell of the former is not “ contorted and misshapen ” and represents, possibly, an intraspecific morph of  G. truncatula characterized by a rounded shell with small spire and somewhat inflated body whorl.</p></div>	https://treatment.plazi.org/id/1B63B09542E951EBABC08905F39D01F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
D72C5B6CA24A5B279D70E291162211DA.text	D72C5B6CA24A5B279D70E291162211DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paludina contecta var. emiliana Roffiaen 1868	<div><p>Paludina contecta var. emiliana Roffiaen, 1868</p><p>Fig. 1 A</p><p>Paludina contecta var. emiliana Roffiaen, 1868: 80.</p><p>Type material.</p><p>RBINS I.G.34957 / MT.2029: 1 holotype, ex Roffiaen coll.</p><p>Type locality.</p><p>Italy: Arona, Lago Maggiore,  Italian shore of the lake . “ […] Lac Majeur, à Arona, sur la rive italienne ” (Roffiaen 1868: 80).</p><p>Taxonomic status.</p><p>Junior synonym of  Viviparus contectus (Millet, 1813) .</p><p>Discussion.</p><p>According to the specimen’s label, it stems from the collection of Roffiaen, it is from the type locality and the word “ type ” is written on it in the same handwriting. Considering that the description of Roffiaen (1868) was based on a single shell, the present specimen can be considered the holotype.</p><p>This taxon was distinguished by Roffiaen (1868) as lacking coloured spiral bands and by the first whorls being purplish. Both colour variations are observed in  Viviparus contectus, a species that is known from the region, and the shell shape of the present specimen (Fig. 1 A) is likewise consistent with that species, including the “ hammered ” markings on the teleoconch (Welter-Schultes 2012; Rowson et al. 2021). Nevertheless, one key diagnostic feature of  V. contectus, the sharply pointed tip of the spire (Rowson et al. 2021), is not observed in the present specimen. Still,  V. contectus is known for conchological variation across its distribution (Welter-Schultes 2012) and, thus,  Paludina contecta emiliana is here considered its junior synonym.</p></div>	https://treatment.plazi.org/id/D72C5B6CA24A5B279D70E291162211DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
5BC24B03AB655322946A4E1501D8C77C.text	5BC24B03AB655322946A4E1501D8C77C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valvata colbeaui Roffiaen 1868	<div><p>Valvata colbeaui Roffiaen, 1868</p><p>Fig. 1 B</p><p>Valvata colbeaui Roffiaen, 1868: 81, pl. 1, fig. 1.</p><p>Type material.</p><p>RBINS I.G.9154 / MT.2041: 2 syntypes, ex Roffiaen coll., ex de Cort coll.</p><p>Type locality.</p><p>Switzerland: Iseltwalt,  Lake Brienz (Brienzersee). “ Lac de Brientz à Iseltwalt ” (Roffiaen 1868: 81).</p><p>Taxonomic status.</p><p>Junior synonym of  Valvata antiqua Morris, 1838 (see below).</p><p>Discussion.</p><p>According to the label, the specimens stem from the collection of Roffiaen, they are from the type locality (Iseltwalt) and are labelled “ types ”. Thus, they can be considered syntypes. It is also worthwhile to note that the lot first became part of de Cort’s collection before being acquired by the RBINS.</p><p>Roffiaen’s shells correspond reasonably well to  Valvata antiqua Morris, 1838, a morphospecies for which genetic data remains unavailable and, consequently, a variety of opinions on its taxonomic status and rank exists among malacologists. It is sometimes regarded as merely an infraspecific morph of  V. piscinalis thought to develop in large lakes, including the Alpine lakes (Falniowski 1989; Piechocki and Wawrzyniak-Wydrowska 2016). Falniowski (1989: 73) considered this morph  Valvata piscinalis f.  antiqua colbeaui, an endemic to Lake Geneva. Glöer (2002) ranked it as a subspecies of  V. piscinalis, but this solution is untenable since the ranges of  V. piscinalis piscinalis and  V. piscinalis antiqua broadly overlap. The latest publications tend to accept  V. antiqua as a species, distinct from  V. piscinalis (Vinarski and Kantor 2016; Glöer 2019), although a DNA assessment of this insufficiently studied snail is urgently needed.</p><p>Valvata piscinalis (O. F. Müller, 1774) is a widespread species in Europe, and while it more usually displays a wide shell with an ample body whorl and aperture, more trochoid forms with comparatively smaller apertures are also known (Turner et al. 1998; Glöer 2002, 2019; Vinarski et al. 2013). The present specimens at first seem to fit within the range of conchological variation displayed by that species, though they are smaller at the same number of whorls. The largest specimen available (Fig. 1 B), which is the most similar to the published illustration (Roffiaen 1868: pl. 1, fig. 1), has a more strongly trochoid shell, with narrower and less bulging whorls.</p></div>	https://treatment.plazi.org/id/5BC24B03AB655322946A4E1501D8C77C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo Brincalepe;d'Udekem d'Acoz, Cedric;Vinarski, Maxim V.;Samyn, Yves;Tomotani, Barbara Mizumo	Salvador, Rodrigo Brincalepe, d'Udekem d'Acoz, Cedric, Vinarski, Maxim V., Samyn, Yves, Tomotani, Barbara Mizumo (2025): François Roffiaen's terrestrial and freshwater gastropod types in the collection of the Royal Belgian Institute of Natural Sciences. ZooKeys 1239: 103-121, DOI: 10.3897/zookeys.1239.150840
