taxonID	type	description	language	source
C3C7C641FE2D52C9AC85EECBA36E96AF.taxon	diagnosis	Diagnosis. Separation from other species of Aldinomyces based on ITS 2 (positions 225 – 244 taaagaagatttcttcttta; two mismatches allowed) and LSU D 2 (positions 709 – 728 gcggctggacagctgtgcaa; one mismatch allowed) as indicated in Fig. 50. Intraspecific variation up to 1.1 % in ITS 2 and up to 0.4 % in LSU. Interspecific distance at least 3.9 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C3C7C641FE2D52C9AC85EECBA36E96AF.taxon	description	Description. Other sequences: EUK 1205356 (type locality); EUK 1124394 (GSMc plot G 5912, temperate grassland soil in Rahinge, Estonia, 58.3804 ° N, 26.6289 ° E); EUK 0320467 (FunAqua sediment sample W 0315 s in Cottonwood Lake, ND, USA, 47.1, – 99.09); EUK 0529885 (GSMc plot G 2838 X; tundra soil in Kvaenangsfjellet, Norway, 69.8972 ° N, 21.5778 ° E); and GlobalFungi accessions fa 0 dd 51 fb 77 b 34 cf 5 b 5659 d 5 f 4 d 674 c 1 (forest soil in Yunnan, China, 27.12 ° N, 100.17 ° E); ea 737611 f 71505778 a 4409 d 28 eec 463 d (woodland soil in MT, USA, 45.3982, – 110.704); and 5363 d 0 a 2 c 4815 da 4 d 7 fb 36 f 7 f 94 d 6 c 6 e (forest soil in Austria, 47.36 ° N, 15.05 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C3C7C641FE2D52C9AC85EECBA36E96AF.taxon	etymology	Etymology. Aldino (Italian) refers to the type locality, and Tarquin (English) refers to Tarquin Netherway, who collected the material from the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C00F1B46B22C5C6FB3C1C68E3D941521.taxon	type_taxon	Type species. Aldinomyces tarquinii Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C00F1B46B22C5C6FB3C1C68E3D941521.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 139 – 158 gcggatttcgaaagatttct in type species; one mismatch allowed). Forms a monophyletic, least inclusive clade in Aldinomycetaceae, covering sequences EUK 1205365, EUK 1124394, and EUK 0529911 (Figs 1, 49).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FDED5FD6E64C55A6BD6C968396FA849E.taxon	type_taxon	Type genus. Aldinomyces Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FDED5FD6E64C55A6BD6C968396FA849E.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1686 – 1695 tagcgatagg in S. cerevisiae; no mismatch allowed) and ITS 2 (positions 125 – 137 gcaacatartaat in type species; one mismatch allowed). Forms a monophyletic, least inclusive clade in Aldinomycetales, covering sequences EUK 1205365, EUK 1124394, EUK 0529911, and EUK 0529884 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0B22B2F5324D5725B3E5210289CBDDEB.taxon	type_taxon	Type family. Aldinomycetaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0B22B2F5324D5725B3E5210289CBDDEB.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 732 – 746 gattcaggaccttca in S. cerevisiae; no mismatch allowed), SSU V 7 (positions 1346 – 1355 gttgttggtc in S. cerevisiae; no mismatch allowed), and LSU D 3 (positions 912 – 926 in the type species and 771 – 785 in S. cerevisiae: ggttttgagaaaaag; one mismatch allowed). Forms a monophyletic, least inclusive clade in Aldinomycetes, covering sequences EUK 0320466, EUK 0529888, EUK 1205365, EUK 1124394, EUK 0529884, EUK 1111390, EUK 0529911, and EUK 0320468 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
56AEDF1491A859789F6EA2E9A32479A3.taxon	type_taxon	Type order. Aldinomycetales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
56AEDF1491A859789F6EA2E9A32479A3.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 732 – 746 gattcaggaccttca in S. cerevisiae; no mismatch allowed), SSU V 7 (positions 1346 – 1355 gttgttggtc in S. cerevisiae; no mismatch allowed), and LSU D 3 (positions 912 – 926 in the type species and 771 – 785 in S. cerevisiae: ggttttgagaaaaag; one mismatch allowed). Forms a monophyletic, least inclusive clade in Aldinomycota, covering sequences EUK 0320466, EUK 0529888, EUK 1205365, EUK 1124394, EUK 0529884, EUK 1111390, EUK 0529911, and EUK 0320468 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
752126C0938B50C59E795234344031E8.taxon	type_taxon	Type class. Aldinomycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
752126C0938B50C59E795234344031E8.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 732 – 746 gattcaggaccttca in S. cerevisiae; no mismatch allowed), SSU V 7 (positions 1346 – 1355 gttgttggtc in S. cerevisiae; no mismatch allowed), and LSU D 3 (positions 912 – 926 in the type species and 771 – 785 in S. cerevisiae: ggttttgagaaaaag; one mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 0320466, EUK 0529888, EUK 1205365, EUK 1124394, EUK 0529884, EUK 1111390, EUK 0529911, and EUK 0320468 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FBFF4A6C9250540F9707D6CD24A575AC.taxon	type_taxon	Type genus. Algovorax Tedersoo & Y. Ding.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FBFF4A6C9250540F9707D6CD24A575AC.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S-ITS 2 (positions starting from 150 in type species and 154 in S. cerevisiae gtgaaacctcctcaa; one mismatch allowed) and from other groups of Monoblepharomycota in SSU V 7 (positions 1485 – 1494 in S. cerevisiae acgagtatat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Algovoracales, covering sequences MF 163176, OQ 702880, EF 024210, and OQ 687303 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6A8FB21D8A08546895031C486E4E738F.taxon	type_taxon	Type family. Algovoracaceae Tedersoo & Y. Ding.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6A8FB21D8A08546895031C486E4E738F.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S-ITS 2 (positions starting from 150 in type species and 154 in S. cerevisiae gtgaaacctcctcaa; one mismatch allowed) and from other groups of Monoblepharomycota in SSU V 7 (positions 1485 – 1494 in S. cerevisiae acgagtatat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Algovoracomycetes, covering sequences MF 163176, OQ 702880, EF 024210, and OQ 687303 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1EB976B9745450A9BFEAEF43903308C3.taxon	type_taxon	Type order. Algovoracales Tedersoo & Y. Ding.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1EB976B9745450A9BFEAEF43903308C3.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 5 (positions 696 – 714 in S. cerevisiae tctttctttctggggaacc or ycttttcttttggggaacc; no mismatch allowed). Forms a monophyletic, least inclusive clade in Monoblepharomycota, covering sequences MF 163176, OQ 702880, EF 024210, OQ 687303, OQ 687304, OQ 687305, OQ 687310, OQ 687311, EUK 1216850, DQ 244008, UDB 014650, EUK 1124454, EUK 1216854, EUK 1216849, and OQ 687309 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4BE03F6F92B359DE8BA5A5FA42E7BA5F.taxon	diagnosis	Diagnosis. Separation from species of Phlyctidium and Rhizophydium based on the lack of rhizoids, large sporangium (5 – 8 µm), and thin-walled zoospores. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 79 – 98 tgttttgcataaaaacagga; one mismatch allowed) as indicated in Fig. 15. Intraspecific variation up to 1.7 % in ITS 2. Interspecific distance at least 6.7 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4BE03F6F92B359DE8BA5A5FA42E7BA5F.taxon	description	Description. As in Fott (1967). Other sequence: EUK 0319835 (FunAqua sediment sample W 0265; Malinówka river in Krzesławicka, Poland, 49.9864 ° N, 20.0136 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4BE03F6F92B359DE8BA5A5FA42E7BA5F.taxon	etymology	Etymology. Algovorax is derived from the Latin words algos (algae) and vorare (to devour), referring to algae eaters following the parasitic habit characteristic of the type species.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C908FF2E296055D5A3D53E5EFE4A6530.taxon	type_taxon	Type species. Algovorax scenedesmi (Fott) Tedersoo & Y. Ding.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C908FF2E296055D5A3D53E5EFE4A6530.taxon	description	Description. Thallus monocentric, consisting of extramatrical, inoperculate, spherical to oval sporangium, and intramatrical spherical apophysis. Rhizoids absent. Zoospores spherical, thin-walled. Resting spores spherical, thick-walled, arising from the extramatrical sporangium. Parasitic on green algae.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C908FF2E296055D5A3D53E5EFE4A6530.taxon	diagnosis	Diagnosis. Distinguishable from other genera of Monoblepharomycota by an intramatrical spherical apophysis and inoperculate sporangium. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S-ITS 2 (positions starting from 150 in type species and 154 in S. cerevisiae gtgaaacctcctcaa; one mismatch allowed) and from other groups of Monoblepharomycota in SSU V 7 (positions 1485 – 1494 in S. cerevisiae acgagtatat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Algovoracaceae, covering sequences MF 163176, OQ 702880, EF 024210, and OQ 687303 (Figs 1, 14).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
59FF4FA3C5915D9BA37CB247CAD6B746.taxon	type_taxon	Type class. Aphelidiomycota Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T. W. May, M. Ryberg & Abarenkov.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
59FF4FA3C5915D9BA37CB247CAD6B746.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
66C30CDDAAE152098DF95B0129A7512F.taxon	type_taxon	Type class. Aphelidiomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T. W. May, M. Ryberg & Abarenkov.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
66C30CDDAAE152098DF95B0129A7512F.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2AC2D7276E8D5414B1D372E58E83161B.taxon	type_taxon	Type genus. Aquamastix Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2AC2D7276E8D5414B1D372E58E83161B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 4 (positions 966 – 975 gatcaagagc in S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Aquamastigales, covering sequences EUK 1124847 and EUK 0320721 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8B25E62215D950D7970A0CB34ADCCA7F.taxon	type_taxon	Type family. Aquamastigaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8B25E62215D950D7970A0CB34ADCCA7F.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 4 (positions 966 – 975 gatcaagagc in S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Aquamastigomycetes, covering sequences EUK 1102371, EUK 1107057, EUK 1124848, EUK 1138328, EUK 1102991, EUK 1124847, and EUK 0320721 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6D83079E1F655681AD408154F575CB3D.taxon	type_taxon	Type order. Aquamastigales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6D83079E1F655681AD408154F575CB3D.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 4 (positions 966 – 975 gatcaagagc in S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Neocallimastigomycota, covering sequences EUK 1102371, EUK 1107057, EUK 1124848, EUK 1138328, EUK 1102991, EUK 1124847, and EUK 0320721 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
67574F56DCA459EA8EB17CEB8D43DBA9.taxon	diagnosis	Diagnosis. Separation from other species of Aquamastix based on ITS 2 (positions 112 – 131 aatattaatatatttattaa; one mismatch allowed) and LSU (positions 471 – 490 aagacttataattaaaggac; one mismatch allowed) as indicated in Fig. 18. Intraspecific variation up to 1.2 % in ITS 2. Closest species differ by> 20 % in ITS 2 and> 10 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
67574F56DCA459EA8EB17CEB8D43DBA9.taxon	description	Description. Other sequences: EUK 0320721 (FunAqua sediment sample W 0987 s, Szczecin Lagoon, Poland, 53.74 ° N, 14.44 ° E) and GlobalFungi accession 19 c 3 de 17 f 55 f 5 bab 0644510 c 210733 d 4 (sediment in Hulun Lake, Inner Mongolia, China, 48.86 ° N, 117.4 ° E; two biological samples).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
67574F56DCA459EA8EB17CEB8D43DBA9.taxon	etymology	Etymology. Aqua (Latin) and mastix (Greek) refer to water and Neocallimastix, respectively, and Sandusky (Wyandot) refers to the cold waters and the part of Lake Erie where the type material originates.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DB2930C4813E5C6E9C1B8A033D0A419B.taxon	type_taxon	Type species. Aquamastix sanduskyensis Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DB2930C4813E5C6E9C1B8A033D0A419B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 120 – 139 ttcctctttg in type species and 118 – 127 in S. cerevisiae; no mismatch allowed), SSU V 9 (positions 1685 – 1699 agtaacttccccttg in S. cerevisiae; no mismatch allowed), and LSU D 1 (positions 125 – 139 in type species and 123 – 137 in S. cerevisiae gtgacggtttaactg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Aquamastigaceae, covering sequences EUK 1124847 and EUK 0320721 (Figs 1, 17).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A37740D8E1965B998E6602B8C1C7E3EA.taxon	type_taxon	Type genus. Aquieurochytrium Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A37740D8E1965B998E6602B8C1C7E3EA.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 1 (positions 171 – 185 in type species and S. cerevisiae ggcaagccgggcaaa; one mismatch allowed), SSU V 4 (positions 871 – 885 in S. cerevisiae atactttcattagtc; one mismatch allowed), and ITS 2 (positions 173 – 195 in type species taatgctgggcgtcagcctgctt OR taatgacgggcgtcagcctgctt; three mismatches allowed). Forms a monophyletic, least inclusive clade in Aquieurochytriales, covering sequences AB 971081, EUK 1100022, EUK 1102113, EUK 1123700, and EUK 1102276 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5D92AAD1D97C5062B56F2830AB425770.taxon	type_taxon	Type family. Aquieurochytriaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5D92AAD1D97C5062B56F2830AB425770.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 1 (positions 171 – 185 in type species and S. cerevisiae ggcaagccgggcaaa; one mismatch allowed), SSU V 4 (positions 871 – 885 in S. cerevisiae atactttcattagtc; one mismatch allowed), and ITS 2 (positions 173 – 195 in type species taatgctgggcgtcagcctgctt OR taatgacgggcgtcagcctgctt; three mismatches allowed). Forms a monophyletic, least inclusive clade in Aquieurochytriomycetes, covering sequences AB 971081, EUK 1100022, EUK 1102113, EUK 1123700, and EUK 1102276 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
419B76C464CC5F879FAEA1FFB4AAD13E.taxon	type_taxon	Type order. Aquieurochytriales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
419B76C464CC5F879FAEA1FFB4AAD13E.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 1 (positions 171 – 185 in type species and S. cerevisiae ggcaagccgggcaaa OR ggctgctcggacaaa; two mismatches allowed). Forms a monophyletic, least inclusive clade in Chytridiomycota, covering sequences EUK 1107407, AB 971081, EUK 1100022, EUK 1102113, EUK 1123700, and EUK 1102276 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
602728F4A52257E78C933EB92F79E768.taxon	diagnosis	Diagnosis. Separation from other species of Aquieurochytrium based on the ITS 2 (positions 297 – 316 gaaaggggatctgttttttt; one mismatch allowed) and LSU D 2 (positions 470 – 489 in type species and 450 – 469 in S. cerevisiae atgtcgagtccccgatcagt; no mismatch allowed) as indicated in Fig. 7. Intraspecific variation up to 1.1 % in ITS 2. Interspecific distance at least 3.4 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
602728F4A52257E78C933EB92F79E768.taxon	description	Description. Other sequences: EUK 0584914 (FunAqua sample W 0790 w; lake water in Beukenlaan, Netherlands, 52.000 ° N, 4.487 ° E); EUK 0584915 (FunAqua sample W 0038 w; water in Lake Luke Vanajärv, Estonia, 58.2438 ° N, 26.5751 ° E); EUK 0584916 (FunAqua sample W 0458 w; water in Lake Stübnitzsee, Germany, 53.1071 ° N, 13.1891 ° E); EUK 0584917 (FunAqua sample W 0624 w; water in Lake Vejlsø, Denmark, 56.1514 ° N, 9.5618 ° E); EUK 0584918 (FunAqua sample W 0454 w; water in Lake Kleiner Wentowsee, Germany, 53.4494 ° N, 13.1052 ° E); and EUK 0584919 (FunAqua sample W 0364 s; sediment in Lake Ototoa, New Zealand, – 36.5302, 174.2324).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
602728F4A52257E78C933EB92F79E768.taxon	etymology	Etymology. Aqua (Latin) and Europa (Greek) refer to the habitat in European waters; and lacuster (Latin) specifies the lake habitat.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CD469F26842156BD9CF05B38F3BBABCC.taxon	type_taxon	Type species. Aquieurochytrium lacustre Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CD469F26842156BD9CF05B38F3BBABCC.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 160 – 168 ccgcgacga; one mismatch allowed) and LSU D 2 (positions 618 – 637 in type species and 591 – 610 in S. cerevisiae tcgcagcgcaccgtaaggcg). Forms a monophyletic, least inclusive clade in Aquieurochytriaceae, covering sequences EUK 1102113 and EUK 1102276 (Figs 1, 6).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9DB42ADB8BE252F29DDF699CF5AD0277.taxon	type_taxon	Type genus. Ruderalia Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9DB42ADB8BE252F29DDF699CF5AD0277.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 209 – 222 in type species aacgatagtgaagt; two mismatches allowed). Forms a monophyletic, least inclusive clade in Ruderaliales, covering sequences EUK 1138161, EUK 1137899, EUK 0531800, and EUK 1124460 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
97A81F34F6225DFF985A49B505AE76A4.taxon	type_taxon	Type species. Borikenia urbinae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
97A81F34F6225DFF985A49B505AE76A4.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1684 – 1703 in S. cerevisiae tgcggtccacatgttggcaa; one mismatch allowed) and ITS 2 (positions 26 – 45 in type species ttggtggacttggtcgttca; two mismatches allowed). Forms a monophyletic, least inclusive clade in Borikeniaceae, covering sequences EUK 1189254, EUK 0530094, and EUK 0530090 (Figs 1, 51).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
41E2A976CEEE5A6DB75BCFAE71303BE2.taxon	diagnosis	Diagnosis. Separation from other species of Borikenia based on ITS 2 (positions 56 – 75 acgttgtgtacacacacgtg; one mismatch allowed) and LSU (positions 170 – 189 ctgatcttggttgttgggta; one mismatch allowed) as indicated in Fig. 52. Intraspecific variation up to 2.3 % in ITS 2 and up to 0.4 % in LSU. Interspecific distance at least 6.1 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
41E2A976CEEE5A6DB75BCFAE71303BE2.taxon	description	Description. Other sequences: EUK 1102667 (tropical rainforest soil in El Yunque, Puerto Rico, 18.29, – 65.78); EUK 0330861 (GSMc plot AV 207, tropical rainforest soil in Puerto Santander, Colombia, – 0.6161, – 72.401); EUK 0330863 (GSMc plot S 1227, Eucalyptus plantation soil in Ayapel, Colombia, 8.27, – 75.2); EUK 0330864 (GSMc plot S 1026, tropical rainforest soil in Matouta, Reunion, France, – 21.3522 ° N, 55.7059 ° E); EUK 0330862 (GSMc plot S 003, Uapaca tropical forest soil in Manangotry, Madagascar, – 24.745, 46.852); EUK 0330869 (GSMc plot S 048, tropical rainforest soil in El Yunque, Puerto Rico, 18.3167, – 65.8167 ° E); EUK 0330870 (GSMc plot JYK 035, Eucalyptus plantation soil in Rivercess, Liberia, 5.7282, – 9.629); and EUK 0330871 (GSMc plot S 1267, tropical rainforest soil in Khong Ngam, Thailand, 20.2433 ° N, 100.0981 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
41E2A976CEEE5A6DB75BCFAE71303BE2.taxon	etymology	Etymology.> Boriken (Taino) refers to Puerto Rico, where the type material originates, and Urbina (Spanish) refers to Hector Urbina, who was the first to collect material from this species (EUK 1102667).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C3FDB6E50AAA531EABFA738C8E522988.taxon	type_taxon	Type genus. Borikenia Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C3FDB6E50AAA531EABFA738C8E522988.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 3 (positions 956 – 967 in type species and 761 – 772 in S. cerevisiae tcaatttattga; no mismatch allowed) and 5.8 S (positions 130 – 142 in type species and 132 – 144 in S. cerevisiae aagagtatttctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Borikeniales, covering sequences EUK 1189254, EUK 0530094, EUK 0530090, EUK 1189255, and EUK 1189256 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
55DBFC6882BA52F689EC30B79A436A7C.taxon	type_taxon	Type family. Borikeniaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
55DBFC6882BA52F689EC30B79A436A7C.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 3 (positions 956 – 967 in type species and 761 – 772 in S. cerevisiae tcaatttattga OR ggaatttattcc; no mismatch allowed). Forms a monophyletic, least inclusive clade in Borikeniomycetes, covering sequences EUK 1105319, EUK 1189257, EUK 0530094, EUK 0530090, EUK 1189254, EUK 1189255, and EUK 1189256 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9A4688DE408655E5ADEE2A888180984B.taxon	type_taxon	Type order. Borikeniales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9A4688DE408655E5ADEE2A888180984B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 3 (positions 956 – 967 in type species and 761 – 772 in S. cerevisiae tcaatttattga OR ggaatttattcc; two mismatches allowed). Forms a monophyletic, least inclusive clade in Borikeniomycota, covering sequences EUK 1105319, EUK 1189257, EUK 0530094, EUK 0530090, EUK 1189254, EUK 1189255, and EUK 1189256 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95C666707222579A86001BA54F951155.taxon	type_taxon	Type class. Borikeniomycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95C666707222579A86001BA54F951155.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 3 (positions 956 – 967 in type species and 761 – 772 in S. cerevisiae tcaatttattga OR ggaatttattcc; two mismatches allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1105319, EUK 1189257, EUK 0530094, EUK 0530090, EUK 1189254, EUK 1189255, and EUK 1189256 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EFE68EAB30B55D6E99FBA142E79AA7E8.taxon	type_taxon	Type genus. Bryolpidium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EFE68EAB30B55D6E99FBA142E79AA7E8.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1706 – 1719 in S. cerevisiae gtcgagaagttatc; one mismatch allowed), ITS 2 (positions 102 – 113 in type species agngaacagcgg; one mismatch allowed), and LSU D 1 (positions 169 – 183 in type species and 167 – 181 in S. cerevisiae cgcggctgccgaagt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Bryolpidiales, covering sequences EUK 1124873 and EUK 1608195 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
40A362F482A05E778C70FD24395E2B23.taxon	type_taxon	Type family. Bryolpidiaceae Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
40A362F482A05E778C70FD24395E2B23.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 1 (positions 169 – 183 in type species and 167 – 181 in S. cerevisiae cgcggctgccgaagt or ggtcgcgaccgcggt; one mismatch allowed) and ITS 2 (positions 102 – 113 in type species agngaacagcgg or aggcacggcagt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Bryolpidiomycetes, covering sequences EUK 1124873 and EUK 1608195 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8B01EF6BB6805E1BB72FDF9EBA853814.taxon	type_taxon	Type order. Bryolpidiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8B01EF6BB6805E1BB72FDF9EBA853814.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 1 (positions 169 – 183 in the type species and 167 – 181 in S. cerevisiae cgcggctgccgaagt or ggtcgcgaccgcggt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Olpidiomycota, covering sequences EUK 1124873, EUK 1608195, EUK 1186288, and EUK 1186289 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DD346D1BA97F5C6BA0E980B5DEEC1FB6.taxon	diagnosis	Diagnosis. Separation from other species of Bryolpidium based on ITS 2 (positions 226 – 245 ctgaaaacaattcgagtgat; no mismatch allowed) and LSU (positions 465 – 494 gacggggctctcgctcgtga; no mismatch allowed) as indicated in Fig. 38. Intraspecific variation up to 4.4 % in ITS 2. Interspecific distance> 20 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DD346D1BA97F5C6BA0E980B5DEEC1FB6.taxon	description	Description. Other sequences: EUK 0530197 (GSMc plot G 6091, subtropical desert soil in Al Zita, Saudi Arabia, 28.9243 ° N, 35.4438 ° E); EUK 0530198 (urban park soil in Mildura, VIC, Australia, – 34.1854 ° N, 142.1696 ° E); EUK 0649726 (urban soil in Põlva, Estonia, 58.0666 ° N, 27.0939 ° E); and GlobalFungi records d 4847020 b 222 e 5 b 7830540 d 220 e 20499 (subtropical woodland soil in El Tepeyac, San Luis Potosi, Mexico, 57.7165 ° N, 27.0549 ° E); 32232805 aef 1 d 4510876 e 11 cba 753 f 7 d (temperate shrubland soil in Elche, Spain, 38.30, – 0.72); and 156 ae 55 aafdd 258247 d 24 e 567 a 23 cdf 3 (temperate forest soil in Ait Tamlil, Morocco, 31.56, – 6.99).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DD346D1BA97F5C6BA0E980B5DEEC1FB6.taxon	etymology	Etymology.> Bryum (Greek and Latin) refers to its common habitat amongst mosses, and mundanum (Latin) refers to cosmopolitan distribution.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CB3E75E16D0D54358DD557003FB3FF6A.taxon	type_taxon	Type species. Bryolpidium mundanum Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CB3E75E16D0D54358DD557003FB3FF6A.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1706 – 1719 in S. cerevisiae gtcgagaagttatc; one mismatch allowed), ITS 2 (positions 102 – 113 in type species agngaacagcgg; one mismatch allowed), and LSU D 1 (positions 169 – 183 in type species and 167 – 181 in S. cerevisiae cgcggctgccgaagt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Bryolpidiaceae, covering sequences EUK 1124873 and EUK 1608195 (Figs 1, 37).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6422C218627652E2A2B77C965D54FD6A.taxon	type_taxon	Type order. Calcarisporiellales Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T. W. May, M. Ryberg & Abarenkov.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6422C218627652E2A2B77C965D54FD6A.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
18D6DA70537B5ABDB8490BB0CEF094F9.taxon	type_taxon	Type class. Calcarisporiellomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T. W. May, M. Ryberg & Abarenkov.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
18D6DA70537B5ABDB8490BB0CEF094F9.taxon	description	Description. As in Tedersoo et al. (2018)	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2F9B1523A43A59DFB9D54A87FB1EA3D2.taxon	type_taxon	Type genus. Cantoromastix Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2F9B1523A43A59DFB9D54A87FB1EA3D2.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 117 – 136 in type species and S. cerevisiae ctttcgggtaayccccggga; one mismatch allowed) and ITS 2 (positions 156 – 170 in type species cgtaaccaaaaggct or cgtaccaratctttt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Cantoromastigales, covering sequences EUK 1124338, EUK 0136917, EUK 0017791, and EUK 0523855 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DB68778F7C60562596894A72C7977263.taxon	type_taxon	Type family. Cantoromastigaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DB68778F7C60562596894A72C7977263.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5 ’ end (positions – 2 – 8 in the type species and S. cerevisiae acgtggtctc or atatggtctc; no mismatch allowed). Forms a monophyletic, least inclusive clade in Cantoromastigomycetes, covering sequences EUK 1152054, EUK 1103194, EUK 1216883, EUK 1124338, EUK 1103697, EUK 1216882, EUK 1124339, EUK 1124340, KU 359437, EUK 1216885, EUK 1137900, and EUK 1216886 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
72FD6806E94356B586029927B9DA642F.taxon	type_taxon	Type order. Cantoromastigales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
72FD6806E94356B586029927B9DA642F.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 6 (positions 1759 – 1778 in type species and 1662 – 1681 in S. cerevisiae ggagacgtcgggdggagccc; no mismatch allowed). Forms a monophyletic, least inclusive clade in Neocallimastigomycota, covering sequences EUK 1107297, EUK 1201627, OQ 687232, OQ 687239, EUK 1103194, EUK 1188586, EUK 1152054, EUK 1103194, EUK 1216883, EUK 1124338, EUK 1103697, EUK 1216882, EUK 1124339, EUK 1124340, KU 359437, EUK 1216885, EUK 1137900, and EUK 1216886 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EF7FD2A032A558FD981137C6BD3A9DBB.taxon	diagnosis	Diagnosis. Separation from other species of Cantoromastix based on ITS 2 (positions 33 – 52 actcgtaaaccattagtttt; one mismatch allowed) and LSU D 2 (positions 679 – 698 ttactcggccatgttagtct; one mismatch allowed) as indicated in Fig. 20. Intraspecific variation up to 1.1 % in ITS 2. Interspecific distance> 20 % in ITS 2 and> 15 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EF7FD2A032A558FD981137C6BD3A9DBB.taxon	description	Description. Other sequences: EUK 0482535 (temperate fallow soil in Haava, Estonia, 58.4611 ° N, 26.7738 ° E); EUK 0330677 (Populus tremula forest soil in Vasula, Estonia, 58.4699 ° N, 26.7266 ° E); and EUK 0330675 (GSMc plot G 5923, Malus domestica cropland soil in Kalnabeites, Latvia, 57.1333 ° N, 24.8567 ° E); EUK 0330674 (GSMc plot G 5920, temperate grassland soil in Viinamärdi, Estonia, 58.2497 ° N, 26.5394 ° E); EUK 0330670 (temperate grassland soil in Kihnu, Estonia, 58.1467 ° N, 23.9852 ° E); EUK 0330673 (GSMc plot G 5930, Zea mays cropland soil in Saulkalne, Latvia, 56.8442 ° N, 24.4072 ° E); and EUK 0330671 (coppiced fallow soil in Lombi, Estonia, 58.4551 ° N, 26.7451 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EF7FD2A032A558FD981137C6BD3A9DBB.taxon	etymology	Etymology. Cantor (Latin) refers to singers, which reflects the origin of the type material at the song festival grounds, and mastix refers to Neocallimastix; and holos and arcticos (Greek) refer to the entire northern (holarctic) distribution of the type species.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C8743E36712B55F4968E3424896724DD.taxon	type_taxon	Type species. Cantoromastix holarctica Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C8743E36712B55F4968E3424896724DD.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 123 – 137 in type species ctagtcatctttaag; two mismatches allowed) and SSU 3 ’ end (positions 1796 – 1800 in S. cerevisiae and 5 bases of ITS: cattagctta; no mismatch allowed). Forms a monophyletic, least inclusive clade in Cantoromastigaceae, covering sequences EUK 1124338, EUK 0136917, and EUK 0017791 (Figs 1, 19).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95D6714BF69750D9A4C12A7B31B32C69.taxon	type_taxon	Type genus. Chthonolpidium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95D6714BF69750D9A4C12A7B31B32C69.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 142 – 154 in type species and 143 – 155 in S. cerevisiae tgttcgacaycc; one mismatch allowed) and LSU D 2 (positions 695 – 714 in type species and 604 – 623 in S. cerevisiae gactgcttgcaggctgcata; one mismatch allowed). Forms a monophyletic, least inclusive clade in Chthonolpidiales, covering sequences EUK 1124876, EUK 0534797, EUK 0534798, EUK 0534818, EUK 1191212, and EUK 1138033 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1C91CB5B5110501792CBDBA1E69B6915.taxon	type_taxon	Type family. Chthonolpidiaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1C91CB5B5110501792CBDBA1E69B6915.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 695 – 714 in type species and 604 – 623 in S. cerevisiae gactgcttgcaggctgcata; two mismatches allowed). Forms a monophyletic, least inclusive clade in Chthonolpidiomycetes, covering sequences EUK 1124876, EUK 0534797, EUK 0534798, EUK 0534818, EUK 1191212, and EUK 1138033 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
636BADF25750582182628E5AEE2541A5.taxon	type_taxon	Type order. Chthonolpidiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
636BADF25750582182628E5AEE2541A5.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in LSU D 2 (positions 695 – 714 in type species and 604 – 623 in S. cerevisiae gactgcttgcaggctgcata; three mismatches allowed). Forms a monophyletic, least inclusive clade in Olpidiomycota, covering sequences EUK 1124876, EUK 0534818, EUK 0534797, EUK 1191212, and EUK 1138033 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3303B4C9453152FF9F96525C1F11041B.taxon	diagnosis	Diagnosis. Separation from other species of Chthonolpidium based on ITS 2 (positions 245 – 264 cacttggctgaaaaggttt; one mismatch allowed) and LSU (positions 608 – 627 ccttctagccctacggtacg; no mismatch allowed) as indicated in Fig. 40. Intraspecific variation up to 4.8 % in ITS 2. Interspecific distance at least 10.3 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3303B4C9453152FF9F96525C1F11041B.taxon	description	Description. Other sequences: EUK 0534827 (type location); EUK 0534801 (temperate shrubland soil in Bliss, MI, USA); and EUK 0534800 (GSMc plot G 6167, subtropical shrubland soil in Al Hiwayb, Oman, 23.2140 ° N, 57.3337 ° E); and from GlobalFungi: 3949559 c 2 adbb 6 dd 485 ee 858 c 50 aa 75 b (shrubland soil in Morocco, 33.9766, – 3.3735 ° E); 170 f 6 c 5254 bf 08 a 8 d 3 be 2909670 b 3 d 85 (coniferous woodland soil in Utah, USA, 37.5819, – 109.91); 3 f 89 b 0 fffeb 9329 f 6 db 10351 b 45 fd 923 (woodland rhizosphere soil in Spain, 37.888, – 3.634); and 03 a 49631062976236 cecf 37723044 ad 8 (shrubland soil in Tunisia, 35.1678 ° N, 8.6738 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3303B4C9453152FF9F96525C1F11041B.taxon	etymology	Etymology.> Khthonios (Greek) refers to the common underground habitat, and enigma (Greek) means puzzling or mysterious.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C235C11C77EC5A90941A59E7D337C4F3.taxon	type_taxon	Type species. Chthonolpidium enigmatum Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C235C11C77EC5A90941A59E7D337C4F3.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 60 – 74 gggccaagctggtta; one mismatch allowed) and LSU D 2 (positions 672 – 686 in type species and 604 – 618 in S. cerevisiae ttgcagttgggcgcc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Chthonolpidiaceae, covering sequences EUK 1124876 and EUK 1138033 (Figs 1, 39).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
65AF78153662523F8BB39E6669C93901.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BA5B3EFB0C6350A999F16BEEFECAFD47.taxon	type_taxon	Type class. Chytridiomycetes Caval. - Sm.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BA5B3EFB0C6350A999F16BEEFECAFD47.taxon	description	Description. As in Doweld (2001).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0FEC9C809B775CE888DB4FA58FBB5613.taxon	diagnosis	Diagnosis. Separation from other species of Curlevskia based on ITS 2 (positions 187 – 206 acgcttytgtgacttcctcc; two mismatches allowed) and LSU D 2 (positions 493 – 512 caatgttcagcgcccctcgt; no mismatch allowed) as indicated in Fig. 30. Intraspecific variation up to 2.1 % in ITS 2 and 1.3 % in LSU. Interspecific distance at least 4.4 % in ITS 2 and 2.8 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0FEC9C809B775CE888DB4FA58FBB5613.taxon	description	Description. Other sequences: EUK 1630897 (GSMc plot G 4803, Ulmus-Alnus forest soil in Meegaste, Estonia, 58.0563 ° N, 26.3355 ° E); EUK 1603984 (GSMc plot G 5821, gravel quarry soil in Siimusti, Estonia, 58.7306 ° N, 26.3198 ° E); EUK 1602442 (GSMc plot G 4128, Quercus robur woodland soil in Ööriku, Estonia, 58.5831 ° N, 22.9322 ° E); and EUK 1700061 (GSMc plot IH. ME 05, Abies forest soil in Mestia, Georgia, 43.0209 ° N, 42.7325 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0FEC9C809B775CE888DB4FA58FBB5613.taxon	etymology	Etymology. Curlevski refers to Nathalie J. A. Curlevski, who was the first to collect material of this genus (GU 187865; Curlevski et al. 2014), and holarctica refers to its distribution.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1CE152A924965A72A72A43EC0B83B291.taxon	type_taxon	Type species. Curlevskia holarctica Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1CE152A924965A72A72A43EC0B83B291.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 88 – 97 in type species tcgcgaatcc; one mismatch allowed) and LSU D 2 (positions 565 – 574 in type species and 509 – 518 in S. cerevisiae atcgcgggaa; one mismatch allowed). Forms a monophyletic, least inclusive clade in Curlevskiaceae, covering sequences EUK 1603989, EUK 1603990, KF 849654, EUK 1103703, EUK 1603986, EUK 1602443, EUK 1603988, EUK 1124409, and EUK 1630897 (Figs 1, 29).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
266F379124AF5BDFB449CC127074AB0E.taxon	type_taxon	Type genus. Curlevskia Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
266F379124AF5BDFB449CC127074AB0E.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 88 – 97 tcgcgaatcc or tcgcaaaacg; one mismatch allowed) and LSU D 2 (positions 541 – 550 in type species and 486 – 495 in S. cerevisiae cacgcaggtc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Curlevskiales, covering sequences EUK 1700102, EUK 1603989, EUK 1603990, KF 849654, EUK 1103703, EUK 1603986, EUK 1602443, EUK 1603988, EUK 1124409, and EUK 1630897 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
639BAD8678745BE98628B740BFE547C6.taxon	type_taxon	Type family. Curlevskiaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
639BAD8678745BE98628B740BFE547C6.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 88 – 97 tcgcgaatcc or tcgcaaaacg; one mismatch allowed) and LSU D 2 (positions 541 – 550 in type species and 486 – 495 in S. cerevisiae cacgcaggtc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Curlevskiomycetes, covering sequences EUK 1700102, EUK 1603989, EUK 1603990, KF 849654, EUK 1103703, EUK 1603986, EUK 1602443, EUK 1603988, EUK 1124409, and EUK 1630897 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5FB813020D3D5E6BB94E2B2D09832961.taxon	type_taxon	Type order. Curlevskiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5FB813020D3D5E6BB94E2B2D09832961.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 2 (positions 549 – 559 in type species and 494 – 504 in S. cerevisiae tcagcgtcagc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Curlevskiomycota, covering sequences EUK 1103826, EUK 1103868, EUK 1700102, EUK 1603989, EUK 1603990, KF 849654, EUK 1103703, EUK 1603986, EUK 1602443, EUK 1603988, EUK 1124409, and EUK 1630897 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8AA271D4AAE952EFB2B4E3DDF66B55D9.taxon	type_taxon	Type class. Curlevskiomycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8AA271D4AAE952EFB2B4E3DDF66B55D9.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in the LSU 5 ’ end (positions 52 – 73 in the type species and S. cerevisiae ccgaggaaaagaaactaacaag or tggaggaaaagaaaaaaacatt; no mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1124408, EUK 1103576, MG 664460, EUK 1700038, EUK 1700047, EUK 1124407, EUK 1631674, EUK 1103826, EUK 1103868, EUK 1700102, EUK 1603989, EUK 1603990, KF 849654, EUK 1103703, EUK 1603986, EUK 1602443, EUK 1603988, EUK 1124409, and EUK 1630897 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
37CDD1AD61AB5A4E8C9714769B597B86.taxon	type_taxon	Type genus. Dobrisimastix Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
37CDD1AD61AB5A4E8C9714769B597B86.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in the ITS 2 region (positions 2 – 18 in type species taaaatrtcacaaccac; three mismatches allowed), SSU V 4 (positions 700 – 719 in S. cerevisiae ctggtgaatcatcgtgctct; one mismatch allowed), and LSU D 1 (positions 124 – 138 in the type species and 122 – 136 in S. cerevisiae tgggtaggttacctg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Dobrisimastigales, covering sequences EUK 1189296, EUK 1138904, EUK 0534669, EUK 0534670, and EUK 0534680 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1A475CDED4F6562984983968787F95B5.taxon	type_taxon	Type family. Dobrisimastigaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
1A475CDED4F6562984983968787F95B5.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 700 – 719 in S. cerevisiae ctggtgaatcatcgtgctct; one mismatch allowed) and LSU D 1 (positions 124 – 138 in the type species and 122 – 136 in S. cerevisiae tgggtaggttacctg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Dobrisimastigomycetes, covering sequences EUK 1189296, EUK 1138904, EUK 0534669, EUK 0534670, and EUK 0534680 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
AAF9817A5E4059C99E5D2E966D2EDE8E.taxon	type_taxon	Type order. Dobrisimastigales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
AAF9817A5E4059C99E5D2E966D2EDE8E.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 1 (positions 124 – 138 in the type species and 122 – 136 in S. cerevisiae tgggtaggttacctg; three mismatches allowed). Forms a monophyletic, least inclusive clade in Neocallimastigomycota, covering sequences EUK 1189296, EUK 1138904, EUK 0534669, EUK 0534670, and EUK 0534680 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5851DC3496435533AA1BB3D9852890C2.taxon	type_taxon	Type species. Dobrisimastix vlkii Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5851DC3496435533AA1BB3D9852890C2.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 2 – 18 in type species taaaatrtcacaaccac; one mismatch allowed), SSU V 4 (positions 700 – 719 in S. cerevisiae ctggtgaatcatcgtgctct; no mismatch allowed), LSU D 1 (positions 124 – 138 in the type species and 122 – 136 in S. cerevisiae tgggtaggttacctg; one mismatch allowed), and LSU D 2 (positions 507 – 526 in type species and 452 – 471 in S. cerevisiae tgtataagaggcttcgcttg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Dobrisimastigaceae, covering sequences EUK 1189296, EUK 1138904, EUK 0534669, EUK 0534670, and EUK 0534680 (Figs 1, 21).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F78E705D5FAE5C63B2DDBA25C603BD81.taxon	diagnosis	Diagnosis. Separation from other species of Dobrisimastix based on ITS 2 (positions 85 – 104 tgcctggttgtctaactata; one mismatch allowed) and LSU D 2 (positions 477 – 496 ttaattcttcgaccgcaagg; one mismatch allowed) as indicated in Fig. 22. Intraspecific variation up to 1.5 % in ITS 2. Interspecific distance at least 8.4 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F78E705D5FAE5C63B2DDBA25C603BD81.taxon	description	Description. Other sequences: EUK 0332259 (GSMc plot S 947, boreal forest soil in Malyi Tigirek, Altai, Russian Federation, 51.1247 ° N, 83.0368 ° E); EUK 0332261 (GSMc plot S 149, temperate Pinus forest soil in Stanislaus, CA, USA, 37.8138, – 119.8926); EUK 0332264 (GSMc plot IH. ME 29, temperate Fagus orientalis forest soil in Mestia, Georgia, 42.9764 ° N, 42.5429 ° E); EUK 0332267 (GSMc plot G 2629, temperate mixed forest soil in Nigula, Estonia, 58.0458 ° N, 24.7119 ° E); EUK 0534684 (GSMc plot S 431, Arctic tundra soil in Toolik Lake, AK, USA, 68.622, – 149.5977); EUK 0584891 (FunAqua sediment sample W 0220 s, Triefenbach, Germany, 49.2812 ° N, 8.1135 ° E); and EUK 0584892 (FunAqua sediment sample W 0525 s, Novaki, Croatia, 45.6573 ° N, 15.6345 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F78E705D5FAE5C63B2DDBA25C603BD81.taxon	etymology	Etymology. Dobříš (Czech) and mastix (Greek) refer to the type locality and Neocallimastix, respectively, and Vlk (Czech) refers to Lukáš Vlk, who collected the type material.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FF8CD3CAC7EB570BB1830EEB07F80A1A.taxon	type_taxon	Type genus. Edaphochytrium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FF8CD3CAC7EB570BB1830EEB07F80A1A.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5 ’ end (positions 15 – 24 in the type species and S. cerevisiae tagtggacta or tgatggacta; one mismatch allowed). Forms a monophyletic, least inclusive clade in Edaphochytriales, covering sequences EUK 1008462, EUK 1200051, EUK 1123749, EUK 1630709, EUK 1123746, EUK 1200763, and EUK 1123748 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
31016709247B5E029EDC2234BA66CA96.taxon	type_taxon	Type family. Edaphochytriaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
31016709247B5E029EDC2234BA66CA96.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 45 – 64 catagtgaaatgtgataact in type species and S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Edaphochytriomycetes, covering sequences EUK 1671450, EUK 1671451, EUK 1008462, EUK 1200051, EUK 1101631, EUK 1101779, EUK 1123746, EUK 1200763, and EUK 1123748 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
803D1E66296251A0AA026341AEFC1E66.taxon	type_taxon	Type order. Edaphochytriales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
803D1E66296251A0AA026341AEFC1E66.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 45 – 64 in type species and S. cerevisiae catagtgaaatgtgataact or catggtgaaatgtgacaatt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Chytridiomycota, covering sequences EUK 1104126, EUK 1107474, EUK 1671450, EUK 1671451, EUK 1008462, EUK 1200051, EUK 1101631, EUK 1101779, EUK 1200763, and EUK 1123748 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
743523EEA3FD58E79B138FA3F9184DF6.taxon	type_taxon	Type species. Edaphochytrium valuojaense Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
743523EEA3FD58E79B138FA3F9184DF6.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 114 – 133 in type species and S. cerevisiae cagtctcttaaggagataat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Edaphochytriaceae, covering sequences EUK 1200051, EUK 1630709, EUK 1200763, and EUK 1123748 (Figs 1, 8).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3CBD05F11A47576B918D75F77FA7B862.taxon	diagnosis	Diagnosis. Separation from other species of Edaphochytrium based on ITS 2 (positions 99 – 118 tttctataatatttttgaca; one mismatch allowed) and LSU (positions 614 – 633 tgagatatttctgatttttg; one mismatch allowed) as indicated in Fig. 9. Intraspecific variation up to 2.1 % in ITS 2 and up to 0.6 % in LSU. Interspecific distance at least 11.8 % in ITS 2 and 6.9 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3CBD05F11A47576B918D75F77FA7B862.taxon	description	Description. Other sequences: EUK 0133766 (GSMc plot G 3522, temperate deciduous forest soil in Pidula, Estonia, 58.4211 ° N, 22.1522 ° E); EUK 0474798 (Populus × wettsteinii plantation soil in Nõgiaru, Estonia, 58.3262 ° N, 26.5545 ° E); and OU 941982 (grassland soil in Kungsängen, Sweden, 59.837 ° N, 17.661 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3CBD05F11A47576B918D75F77FA7B862.taxon	etymology	Etymology. Edaphos (Greek) refers to ground, and Valuoja (Estonian) refers to the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
789DCE7E9D365502A642775FB1F8B674.taxon	type_taxon	Type phylum. None.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
789DCE7E9D365502A642775FB1F8B674.taxon	description	Description. As in Moore (1980).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B148F9AE45E45E2A927A7BCA24039FC6.taxon	type_taxon	Type genus. Gelotisporidium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B148F9AE45E45E2A927A7BCA24039FC6.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 613 – 627 in type species and 692 – 696 in S. cerevisiae cccttgggcgcaaag; one mismatch allowed). Forms a monophyletic, least inclusive clade in Gelotisporidiales, covering sequences EUK 1138568, EUK 1138757, EUK 1100418, EUK 1138778, EUK 1202629, EUK 1201985, EUK 1104844, EUK 1101158, and EUK 1123671 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4EC045F186215E86A344A54F79E587B4.taxon	type_taxon	Type family. Gelotisporidiaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4EC045F186215E86A344A54F79E587B4.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1541 – 1550 in S. cerevisiae ggatcagtca; no mismatch allowed) and LSU D 1 (positions 302 – 311 in the type species and 305 – 314 in S. cerevisiae cgcgccatct; one mismatch allowed). Forms a monophyletic, least inclusive clade in Gelotisporidiomycetes, covering sequences EUK 1138731, EUK 1138718, EUK 1138568, EUK 1138757, EUK 1100925, EUK 1105586, EUK 1105726, EUK 1105789, EUK 1101184, EUK 1202629, EUK 1201985, EUK 1104844, and EUK 1123671 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
844911DB3E0F52D7945DEB9DC375F6EB.taxon	type_taxon	Type order. Gelotisporidiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
844911DB3E0F52D7945DEB9DC375F6EB.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1541 – 1550 in S. cerevisiae ggatcagtca; no mismatch allowed) and LSU D 1 (positions 302 – 311 in type species and 305 – 314 in S. cerevisiae cgcgccatct; one mismatch allowed). Forms a monophyletic, least inclusive clade in Rozellomycota, covering sequences EUK 1138731, EUK 1138718, EUK 1138568, EUK 1138757, EUK 1100925, EUK 1105586, EUK 1105726, EUK 1105789, EUK 1101184, EUK 1202629, EUK 1201985, EUK 1104844, and EUK 1123671 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B8D340E73DEB58D2980EF95EDC245443.taxon	diagnosis	Diagnosis. Separation from other species of Gelotisporidiaceae based on ITS 2 (positions 111 – 130 ggcaagcccaaccgggagta; one mismatch allowed) and LSU (positions 481 – 500 gagttgtgtcacatatagca; one mismatch allowed) as indicated in Fig. 44. Intraspecific variation up to 4.7 % in ITS 2 and up to 1.0 % in LSU. Interspecific distance> 15 % in ITS 2 and> 10 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B8D340E73DEB58D2980EF95EDC245443.taxon	description	Description. Other sequences: EUK 1101158 (coniferous forest soil in Hofors, Sweden, 60.49 ° N, 16.3 ° E); EUK 0325837 (GSMc plot S 1124, mixed forest soil in Zavodoukovskiy, Tyumen Oblast, Russian Federation, 56.5299 ° N, 66.5028 ° E); EUK 0473501 (GSMc plot IHPR 02, Betula pubescens tundra soil in Stora Sjöfallet, Sweden, 67.6367 ° N, 17.8216 ° E); EUK 0325836 (Betula pubescens tundra soil at Lake Sobach’ye, Krasnoyarsk Krai, Russian Federation, 69.0033 ° N, 90.9875 ° E); and EUK 0325821 (GSMc plot S 1081, Araucaria araucana forest soil in Nahuelbuta, Chile, – 37.7897, – 73.0034 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B8D340E73DEB58D2980EF95EDC245443.taxon	etymology	Etymology. Gelot (Sámi) refers to the type locality at Gelotjávri (Kelottijärvi), and boreale (Latin) refers to the mainly boreal habitat of the species.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FCE220F032B354699805B7C9B8AD4896.taxon	type_taxon	Type species. Gelotisporidium boreale Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
FCE220F032B354699805B7C9B8AD4896.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1699 – 1718 in S. cerevisiae acccgtctttcgttg; one mismatch allowed) and 5.8 S-ITS 2 (positions starting from 151 in type species and 153 in S. cerevisiae agaattgaaa; one mismatch allowed). Forms a monophyletic, least inclusive clade in Gelotisporidiaceae, covering sequences EUK 1201985 and EUK 1101158 (Figs 1, 43).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
17801D1CD81554A2A913A227945F743E.taxon	type_taxon	Type class. Kickxellomycetes Tedersoo, Sánchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T. W. May, M. Ryberg & Abarenkov.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
17801D1CD81554A2A913A227945F743E.taxon	description	Description. As in Tedersoo et al. 2018.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
235650E026F052209505EBB267468D4A.taxon	diagnosis	Diagnosis. Separation from other species of Maerjamyces based on ITS 2 (positions 43 – 75 atacctgtttgagtaccatattcttttcccttt; one mismatch allowed) and LSU D 1 (positions 233 – 252 ttgcactcgtgggttatgta; one mismatch allowed) as indicated in Fig. 34. Intraspecific variation up to 5.4 % in ITS 2 and up to 1.6 % in LSU. Closest species differ by at least 7.4 % in ITS 2 and 5.0 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
235650E026F052209505EBB267468D4A.taxon	description	Description. Other sequences: EUK 1138156 (GSMc plot G 5235, Larix decidua plantation soil in Rõõmu, Estonia, 58.3835 ° N, 26.7742 ° E); EUK 1138160 (GSMc plot G 5803, urban park soil in Toomemägi, Estonia, 58.3786 ° N, 26.7185 ° E); EUK 1138157 (GSMc plot G 5283, Quercus robur plantation in Rahinge, Estonia, 58.3845 ° N, 26.5943 ° E); MT 277862 (book paper in Turin, Italy); OU 939288 (Kungsängen, Sweden, 59.837 ° N, 17.661 ° E); (GSMc plot G 4800, Ulmus laevis forest soil in Tuhkja, Estonia, 58.4159 ° N, 25.2326 ° E); and EUK 1138159 (urban soil in Tartu, Estonia, 58.3913 ° N, 26.6965 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
235650E026F052209505EBB267468D4A.taxon	etymology	Etymology. Maerjamyces refers to the type locality in Märja (Estonian), and mykos (Greek) stands for a fungus; Jumpponen (Finnish) refers to Ari Jumpponen, who was the first to collect material of this species (FJ 780627; Jumpponen et al. 2010).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2A849F7A1A0A57279854362E7BD51733.taxon	type_taxon	Type species. Maerjamyces jumpponenii Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2A849F7A1A0A57279854362E7BD51733.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 77 – 86 in type species and 78 – 87 in S. cerevisiae agagtacgtg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Maerjamycetaceae, covering sequences EUK 1138158, EUK 0484301, and EUK 0484311 (Figs 1, 33).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2432328FB50A56228BD7D02B31126714.taxon	type_taxon	Type genus. Maerjamyces Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2432328FB50A56228BD7D02B31126714.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 77 – 86 in type species and 78 – 87 in S. cerevisiae agagtacgtg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Maerjamycetales, covering sequences EUK 1138158, EUK 0484301, and EUK 0484311 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9773BE4E95645CC98079EE6582AC5F4B.taxon	type_taxon	Type family. Maerjamycetaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9773BE4E95645CC98079EE6582AC5F4B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1684 – 1690 in S. cerevisiae gatgcat; no mismatch allowed) and LSU D 1 (positions 114 – 122 in type species and 115 – 123 in S. cerevisiae cactttctg; no mismatch allowed). Forms a monophyletic, least inclusive clade in Maerjamycetes, covering sequences EUK 1200032, EUK 1217336, EUK 1009005, EUK 0484311, EUK 0484301, and EUK 1138158 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D8D9CB6817E6564FB7D0E91E00A1A572.taxon	type_taxon	Type order. Maerjamycetales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D8D9CB6817E6564FB7D0E91E00A1A572.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1684 – 1690 in S. cerevisiae gatgcat; no mismatch allowed) and LSU D 1 (positions 114 – 122 in type species and 115 – 123 in S. cerevisiae cactttctg; no mismatch allowed). Forms a monophyletic, least inclusive clade in Mortierellomycota, covering sequences EUK 1200032, EUK 1217336, EUK 1009005, EUK 0484311, EUK 0484301, and EUK 1138158 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0B9123C4B23D5826B6981673656AA253.taxon	diagnosis	Diagnosis. Separation from other species of Mirabilomyces based on ITS 2 (positions 68 – 87 cttcggttwtaaaacaaggt; two mismatches allowed) and LSU (positions 534 – 553 ctacgctgtggttgcgcttt; one mismatch allowed) as indicated in Fig. 54. Intraspecific variation up to 11.4 % in ITS 2 due to length polymorphism in multiple mono- and dinucleotide repeats and up to 1.0 % in LSU. Interspecific distance> 20 % in ITS 2 and at least 6.0 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0B9123C4B23D5826B6981673656AA253.taxon	description	Description. Other sequences: EUK 0483305 (temperate broadleaf forest soil in Arcais, France, 46.3038, – 0.6844 ° E); EUK 1124377 (GSMc plot G 5235, Larix decidua plantation soil in Rõõmu, Estonia, 58.3835 ° N, 26.7742 ° E); EUK 1216948 (GSMc plot G 4777, flooded grassland soil in Suur-Pakri Härs-hämani, Estonia, 59.3310 ° N, 23.9272 ° E); EUK 1216949 (GSMc plot G 4679, Salix triandra wetland soil in Prangli Rivimaa, Estonia, 59.6150 ° N, 24.9871 ° E); OU 939561 (grassland soil in Kungsängen, Sweden, 59.837 ° N, 17.661 ° E); EUK 1202060 (GSMc plot G 4747, Prunus-Rhamnus-Euonymus forest soil in Tsirgumäe, Estonia, 57.5942 ° N, 26.3241 ° E); EUK 1216950 (GSMc plot G 4742, Fraxinus-Ulmus forest soil in Lüütre, Estonia, 58.1444 ° N, 25.2628 ° E); and EUK 1216946 (GSMc plot S 1366, temperate grassland soil in Innhavet, Norway, 67.9676 ° N, 15.9277 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0B9123C4B23D5826B6981673656AA253.taxon	etymology	Etymology. Mirabilis (Latin) refers to the remarkable and astonishing finding of a large, unrecognized fungal lineage, and Abruka (Estonian) refers to the type locality of the species.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B3FE3A94FDA15C71A66A46D7D33132EC.taxon	type_taxon	Type species. Mirabilomyces abrukanus Tedersoo & R. H. Nilsson.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B3FE3A94FDA15C71A66A46D7D33132EC.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 205 – 214 cttgattagt in type species; one mismatch allowed). Forms a monophyletic, least inclusive clade in Mirabilomycetaceae, covering sequences EUK 1200676 and EUK 1124366 (Figs 1, 53).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
486765941E595DEAA5C8315E1C70D389.taxon	type_taxon	Type genus. Mirabilomyces Tedersoo & R. H. Nilsson.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
486765941E595DEAA5C8315E1C70D389.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in the LSU 5 ’ end (positions 1 – 11 in the type species and S. cerevisiae tcattctcaac; one mismatch allowed) and ITS 2 (positions 196 – 208 in type species aacaatgacttga; one mismatch allowed). Forms a monophyletic, least inclusive clade in Mirabilomycetales, covering sequences EUK 1200676, EUK 1201256, EUK 1203033, EUK 1201246, EUK 1201583, EUK 1201728, and EUK 1124366 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DF6AA243BEE95033B233F98BE284745B.taxon	type_taxon	Type family. Mirabilomycetaceae Tedersoo & R. H. Nilsson.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DF6AA243BEE95033B233F98BE284745B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5 ’ end (positions 1 – 11 in the type species and S. cerevisiae tcattctcaac or cggatctcaaa; one mismatch allowed). Forms a monophyletic, least inclusive clade in Mirabilomycetes, covering sequences EUK 1100742, EUK 1204083, EUK 1200757, EUK 1201873, EUK 1211619, EUK 1200676, EUK 1201256, EUK 1203033, EUK 1201246, and EUK 1201583 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
47A202E2D65550928B857154A032CC2A.taxon	type_taxon	Type order. Mirabilomycetales Tedersoo & R. H. Nilsson.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
47A202E2D65550928B857154A032CC2A.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 102 – 109 cttgaaat in the type species; one mismatch allowed). Forms a monophyletic, least inclusive clade in Mirabilomycota, covering sequences EUK 1100742, EUK 1204083, EUK 1200757, EUK 1201873, EUK 1211619, EUK 1200676, EUK 1201256, EUK 1203033, EUK 1201246, EUK 1201583, EUK 1107008, EUK 1110728, EUK 1109741, EUK 1201657, EUK 1109988, EUK 1108787, and EUK 1115028 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E44AF40A0E805AEC979E81E0192B894D.taxon	type_taxon	Type class. Mirabilomycetes Tedersoo & R. H. Nilsson.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E44AF40A0E805AEC979E81E0192B894D.taxon	diagnosis	Diagnosis. Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1101676, EUK 1107181, EUK 1103059, EUK 1100023, EUK 1103883, EUK 1102753, EUK 1100742, EUK 1204083, EUK 1200757, EUK 1201873, EUK 1211619, EUK 1200676, EUK 1201256, EUK 1203033, EUK 1201246, EUK 1201583, EUK 1107008, EUK 1110728, EUK 1109741, EUK 1201657, EUK 1109988, EUK 1108787, and EUK 1115028 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3122B554E8DA561AAC345C45238AC500.taxon	type_taxon	Type class. Monoblepharidomycetes J. H. Schaffner.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3122B554E8DA561AAC345C45238AC500.taxon	description	Description. As in Schaffner (1909).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D5EDF9C6663F522A9FBC3CDDFCEFCC7A.taxon	type_taxon	Type order. Mortierellales Caval. - Sm.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D5EDF9C6663F522A9FBC3CDDFCEFCC7A.taxon	description	Description. As in Doweld (2014).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
442E84212AC559A5BD6DA20D463FBDC3.taxon	type_taxon	Type class. Mortierellomycetes Doweld.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
442E84212AC559A5BD6DA20D463FBDC3.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9AAB646EF816588A9ADF4674F62EE7BE.taxon	type_taxon	Type phylum. Mucoromycota Doweld.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
9AAB646EF816588A9ADF4674F62EE7BE.taxon	description	Description. As in Tedersoo et al. (2018)	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F1A3F5560B4B522CB7E6A5F769B5FBDE.taxon	diagnosis	Diagnosis. Separation from other species of Mycosocceria based on ITS 2 (positions 338 – 357 agaactttgttctttttaac; one mismatch allowed) and LSU D 2 (positions 466 – 485 aatctggtcccggtggatgg; one mismatch allowed) as indicated in Fig. 32. Intraspecific variation up to 2.3 % in ITS 2 and 0.2 % in LSU. Interspecific distance at least 10.2 % in ITS 2 and 10.3 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F1A3F5560B4B522CB7E6A5F769B5FBDE.taxon	description	Description. Other sequences: EUK 1603994 (GSMc plot G 5816, Trifolium pratense cropland soil in Hermani, Estonia, 58.8071 ° N, 25.7564 ° E); EUK 1008618 (GSMc plot G 4599, Ulmus laevis forest soil in Liutsepa, Estonia, 58.0791 ° N, 26.0094 ° E); EUK 0331576 (GSMc plot G 5820, Acer - Fraxinus - Ulmus woodland soil in Pajusi, Estonia, 58.7052 ° N, 25.9389 ° E); EUK 0331575 (GSMc plot G 5422, Pinus strobus woodland soil in Tartu, Estonia, 58.3909 ° N, 26.6973 ° E); EUK 0331574 (GSMc plot G 5765 y, grassland soil in Rebaste, Estonia, 58.41 ° N, 25.93 ° E); EUK 0331573 (urban park soil in Slovenia); and EUK 0331572 (GSMc plot S 950, forest tundra soil in Mt. Mayak, Altai kray, Russian Federation, 51.0474 ° N, 82.9718 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F1A3F5560B4B522CB7E6A5F769B5FBDE.taxon	etymology	Etymology. Soccer (English) refers to the football field where the type specimen was collected; and estonica (Latin) refers to Estonia, where this species’ type and most additional materials originate.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
79531C1D48A15E80BC7764DCEEF403F5.taxon	type_taxon	Type species. Mycosocceria estonica Tedersoo, Bahram & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
79531C1D48A15E80BC7764DCEEF403F5.taxon	diagnosis	Diagnosis. Distinguishable from other species of fungi based on diagnostic nucleotide signatures in 5.8 S (positions 120 – 129 in type species and S. cerevisiae cccggtaggc). Forms a monophyletic, least inclusive clade in Mycosocceriaceae, covering sequences EUK 1008618, EUK 0531631, and EUK 1124462 (Figs 1, 31).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0082E1A2F1215F05BEC5A7528320CB27.taxon	type_taxon	Type genus. Mycosocceria Tedersoo, Bahram & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0082E1A2F1215F05BEC5A7528320CB27.taxon	diagnosis	Diagnosis. Distinguishable from other species of Mortierellomycetes based on diagnostic nucleotide signatures in SSU V 9 (positions 1654 – 1663 in S. cerevisiae gattgaacgg; no mismatch allowed), 5.8 S (positions 90 – 99 in type species and S. cerevisiae tcatcaaatc; no mismatch allowed), and LSU D 2 (positions 573 – 592 in type species and 521 – 540 in S. cerevisiae aagttggaggaatgtggctc; two mismatches allowed). Forms a monophyletic, least inclusive clade in Mycosocceriales, covering sequences EUK 0531595, EUK 1102426, EUK 1202279, EUK 0531631, EUK 1008618, and EUK 1124462 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
80E2C6E722AC5D498845C73A33A25A0E.taxon	type_taxon	Type family. Mycosocceriaceae Tedersoo, Bahram & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
80E2C6E722AC5D498845C73A33A25A0E.taxon	diagnosis	Diagnosis. Distinguishable from other species of Mortierellomycota based on diagnostic nucleotide signature in SSU V 9 (positions 1654 – 1663 in S. cerevisiae gattgaacgg; no mismatch allowed) and from all fungi in LSU D 2 (positions 573 – 92 in the type species and 521 – 540 in S. cerevisiae aagttggaggaatgtggctc; two mismatches allowed). Forms a monophyletic, least inclusive clade in Mortierellomycetes, covering sequences EUK 0531595, EUK 1102426, EUK 1202279, EUK 0531631, EUK 1008618, and EUK 1124462 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BD51D3A42BE2537A82E078190BC23410.taxon	diagnosis	Diagnosis. Separation from other species of Nematovomyces based on ITS 2 (positions 491 – 510 aaaaccctttttcccccaca; one mismatch allowed) and LSU (positions 601 – 620 tgttcttggtactgagttta; one mismatch allowed) as indicated in Fig. 56. Intraspecific variation up to 2.8 % in ITS 2 and up to 0.3 % in LSU. Interspecific distance at least 8.0 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BD51D3A42BE2537A82E078190BC23410.taxon	description	Description. Other sequences: EUK 1200775 (GSMc plot S 1183, mixed forest soil in Aldino, Italy, 46.4072 ° N, 11.4964 ° E); EUK 1217250 (GSMc plot G 4679, Salix triandra swamp soil in Prangli Rivimaa, Estonia, 59.6151 ° N, 24.9871 ° E); EUK 0330847 (GSMc plot S 141, Carpinus- Quercus - Alnus forest soil in Shirgah, Iran, 36.2122 ° N, 52.8243 ° E); EUK 0483680 (GSMc plot G 4196, mixed forest soil in Kahvena, Estonia, 58.2799 ° N, 25.2316 ° E); EUK 1217249 (GSMc plot G 4800, Ulmus-Alnus temperate forest soil in Tuhkja, Estonia, 58.4159 ° N, 25.2327 ° E); EUK 033840 (GSMc plot S 939, tropical rainforest soil in Parotania, Bolivia, – 17.5815, – 66.3443 ° E); and EUK 0330843 (GSMc plot G 4030, Quercus-Arbutus forest soil in Ain Boumahdi, Morocco, 34.0096, – 4.2858, 24.9871 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BD51D3A42BE2537A82E078190BC23410.taxon	etymology	Etymology. Soinaste (Estonian) refers to the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED79EBD649275F33A6260BE13B115E36.taxon	type_taxon	Type species. Nematovomyces vermicola (G. L. Barron & Szuarto) Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED79EBD649275F33A6260BE13B115E36.taxon	diagnosis	Diagnosis. Separated from the vascular plant-associated species and algae-associated species of Olpidium s. stricto based on reticulate to spiky ornamentation in resting spores instead of star-like shapes. Nematovomyces spp. infect nematodes, rotifers, and their eggs. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 729 – 743 aaccgggtgtggcct in S. cerevisiae; no mismatch allowed) and ITS 2 (positions 68 – 77 aacaatgtct in N. soinasteënsis; one mismatch allowed) and LSU D 2 (positions 679 – 698 in N. soinasteënsis and 595 – 614 in S. cerevisiae: gttgtctttgttattttcca; one mismatch allowed). Forms a monophyletic, least inclusive clade in Nematovomycetaceae, covering sequences OQ 702947, GQ 330624, OQ 702883, JN 054659, JN 054675, OQ 702805, EUK 1100016, EUK 1107129, EUK 1102228, EUK 1204135, EUK 1124398, EUK 1124395, EUK 1124396, and EUK 1124397 (Figs 1, 55).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED79EBD649275F33A6260BE13B115E36.taxon	description	Description. Sporangia in the host cell singly or in rows, spherical or pyriform, 15 – 35 µm diam., with a smooth wall and curved exit tube. Zoospores spherical, 2.5 – 3.5 µm diam, with one posterior flagellum. Flagellum arched near the insertion point to the zoospore body. Thallus produces a single evacuation tube that leaves a narrow exit tube. Resting spores spherical or oblong, with surface ornamented by delicate reticular pattern or linear or branched spines, arranged in chains outside the animal cuticle or in culture. Infects nematodes, rotifers, and their eggs internally.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
989CDAF51A785DB9B37CC5543904D488.taxon	diagnosis	Diagnosis. Separated from other species of Nematovomyces by echinulate resting spores and parasitism exclusively on nematode eggs.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
989CDAF51A785DB9B37CC5543904D488.taxon	description	Description. As in Barron and Szuarto (1986).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
989CDAF51A785DB9B37CC5543904D488.taxon	etymology	Etymology. Nematoda and ovum (Latin) refer to roundworms and their eggs, respectively, and describe the specific association with nematode eggs, indicating parasitic relationship with these structures.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3AC66B436BE454D7849B0E398F1BAB74.taxon	type_taxon	Type genus. Nematovomyces Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3AC66B436BE454D7849B0E398F1BAB74.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 68 – 77 aacaatgtct or atcaatggtt in N. soinasteënsis; one mismatch allowed). Forms a monophyletic, least inclusive clade in Nematovomycetales, covering sequences AB 971072, EUK 1106088, OQ 702947, GQ 330624, OQ 702883, JN 054659, JN 054675, OQ 702805, EUK 1100016, EUK 1107129, EUK 1102228, EUK 1204135, EUK 1124398, EUK 1124395, EUK 1124396, and EUK 1124397 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
986895C5F6DA59F7B958013848DF120C.taxon	type_taxon	Type family. Nematovomycetaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
986895C5F6DA59F7B958013848DF120C.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 127 – 146 in N. soinasteënsis and S. cerevisiae atccggyaggtatacctatt or gcctgcaggtatacctattt or acgtgcaagtatacctattt; one mismatch allowed) and in LSU D 3 (positions 905 – 919 in N. soinasteënsis and 748 – 762 in S. cerevisiae: acccgatcctagctc; two mismatches allowed). Forms a monophyletic, least inclusive clade in Nematovomycetes, covering sequences EUK 1137920, EUK 1124402, EUK 1124400, AB 971072, EUK 1106088, OQ 702947, GQ 330624, OQ 702883, JN 054659, JN 054675, OQ 702805, EUK 1100016, EUK 1107129, EUK 1102228, EUK 1204135, EUK 1124398, EUK 1124395, EUK 1124396, EUK 1200775, and EUK 1124397 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3E348D7CA3D057D6B52935DB51798B69.taxon	type_taxon	Type order. Nematovomycetales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3E348D7CA3D057D6B52935DB51798B69.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 127 – 146 in N. soinasteënsis and S. cerevisiae: atccggyaggtatacctatt or gcctgcaggtatacctattt or acgtgcaagtatacctattt or atccaaagagtatacttgtt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Nematovomycota, covering sequences EUK 1217270, EUK 1137920, EUK 1124402, EUK 1124400, AB 971072, EUK 1106088, OQ 702947, GQ 330624, OQ 702883, JN 054659, JN 054675, OQ 702805, EUK 1100016, EUK 1107129, EUK 1102228, EUK 1204135, EUK 1124398, EUK 1124395, EUK 1124396, EUK 1200775, and EUK 1124397 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
31C68CB955F65C2F86CEAC8A33D3AA3D.taxon	type_taxon	Type class. Nematovomycetes Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
31C68CB955F65C2F86CEAC8A33D3AA3D.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5´end (positions 5 – 14 in the type species and S. cerevisiae cctgaawtta; one mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1124405, EUK 1137897, EUK 1138000, EUK 1105583, EUK 1217236, EU 162639, AB 971078, OL 869110, EUK 1217234, EUK 1137920, EUK 1124400, AB 971072, EUK 1106088, OQ 702947, GQ 330624, OQ 702883, JN 054659, JN 054675, OQ 702805, EUK 1100016, EUK 1217270, and EUK 1124397 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0EFCBE20C5D45127852B555747970FA4.taxon	type_taxon	Type class. Neocallimastigomycetes M. J. Powell.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0EFCBE20C5D45127852B555747970FA4.taxon	description	Description. As in Hibbett et al. (2007).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BF2354CFBCC25551B4657944C8CE358D.taxon	type_taxon	Type phylum. Olpidiomycota Doweld.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BF2354CFBCC25551B4657944C8CE358D.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F3574A173B0A5E59AC01304343BE78C7.taxon	type_taxon	Type class. Olpidiomycetes Doweld.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F3574A173B0A5E59AC01304343BE78C7.taxon	description	Description. As in Doweld (2013).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
07FBEC084A265224B840BD08BB850074.taxon	type_taxon	Type genus. Palomastix Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
07FBEC084A265224B840BD08BB850074.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1664 – 1678 in S. cerevisiae tttagtgaggactcg; no mismatch allowed) and 5.8 S (positions 116 – 135 in type species and S. cerevisiae gcctcccggtattccaggag or gcttcatggtattccgtga; one mismatch allowed). Forms a monophyletic, least inclusive clade in Palomastigales, covering sequences EUK 1124846, EUK 1123686, EUK 0320705, and EUK 0320700 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8E313381A6E258B994F917D790EFCD1F.taxon	type_taxon	Type family. Palomastigaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8E313381A6E258B994F917D790EFCD1F.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1664 – 1678 in S. cerevisiae tttagtgaggactcg; no mismatch allowed) and 5.8 S (positions 116 – 135 in type species and S. cerevisiae gcctcccggtattccaggag or gcttcatggtattccgtga; one mismatch allowed). Forms a monophyletic, least inclusive clade in Palomastigomycetes, covering sequences EUK 1124846, EUK 1123686, EUK 0320705, and EUK 0320700 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BF5B17104026529F9F06727027061BDD.taxon	type_taxon	Type order. Palomastigales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BF5B17104026529F9F06727027061BDD.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 8 (positions 1664 – 1678 in S. cerevisiae tttagtgaggactcg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Neocallimastigomycota, covering sequences EUK 1124846, EUK 1123686, EUK 0320705, and EUK 0320700 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
7151F516666954FCA9E5A571EF62631F.taxon	diagnosis	Diagnosis. Separation from other species of Palomastix based on ITS 2 (positions 225 – 244 ctaaaagtcgggtttgattt; one mismatch allowed) and ITS 1 (positions 464 – 483 cagcaggtcttgactgactt; one mismatch allowed) as indicated in Fig. 24. Intraspecific variation up to 1.4 % in ITS 2. Interspecific distance at least 9.2 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
7151F516666954FCA9E5A571EF62631F.taxon	description	Description. Other sequences: EUK 0320710 and EUK 0574068 (type locality); EUK 0320711, EUK 0320713 (FunAqua lake sediment sample in Viitna Pikkjärv, Estonia, 59.4469 ° N, 26.0107 ° E); and EUK 0320712 (FunAqua lake sediment sample W 0992 s in Svartkulpen, Norway, 59.9741 ° N, 10.7373 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
7151F516666954FCA9E5A571EF62631F.taxon	etymology	Etymology.> Palo (Estonian) refers to the type locality, Lake Palojärv; lacustris refers to habitat in lakes.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
880F0DA574395ACD9D697205386B0EC4.taxon	type_taxon	Type species. Palomastix lacustris Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
880F0DA574395ACD9D697205386B0EC4.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in 5.8 S (positions 116 – 135 in type species and S. cerevisiae gcctcccggtattccaggag; one mismatch allowed), SSU V 9 (positions 1691 – 1710 in S. cerevisiae tgttgggctcacgccctcct; one mismatch allowed), and ITS 2 (positions 129 – 148 in type species tctcaagttaagtgattggt; two mismatches allowed). Forms a monophyletic, least inclusive clade in Palomastigaceae, covering sequences EUK 1123686, EUK 0320705, and EUK 0320700 (Figs 1, 23).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3D7B89BF1BA55054AC0D06CEF12A5232.taxon	diagnosis	Diagnosis. Separation from other species of Pantelleria based on ITS 2 (positions 131 – 155 tttacatctttttctaaacttaatc; one mismatch allowed) and LSU D 2 (positions 722 – 741 aagagtgatggtgatcaagt; one mismatch allowed) as indicated in Fig. 3. Intraspecific variation up to 3.8 % in ITS 2 and up to 1.5 % in LSU. Interspecific distance at least 10.1 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3D7B89BF1BA55054AC0D06CEF12A5232.taxon	description	Description. Other sequences: OW 839340 and OW 840352 (unspecified soil in Tianshan Mountains, Uyghur Autonomous Region, China); EUK 1138064 (GSMc plot G 4627, mixed forest soil in Tudusoo, Estonia, 59.11368 ° N, 26.75944 ° E); EUK 1120811 (GSMc plot S 281, Quercus robur alley soil in Tartu, Estonia, 58.379 ° N, 26.706 ° E); EUK 0348125 (urban park soil in Niort, France, 46.325, – 0.4672 ° E); MH 625427 (microcosm soil in New Zealand); and MF 484888 (unspecified soil in Great Britain).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3D7B89BF1BA55054AC0D06CEF12A5232.taxon	etymology	Etymology.> Pantelleria (Maltese) refers to the type locality, and Saitta (Sicilian) refers to Alessandro Saitta, who collected material from the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0C93F2C096145E579A4C29093B795F7D.taxon	type_taxon	Type species. Pantelleria saittana Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0C93F2C096145E579A4C29093B795F7D.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in SSU V 9 (positions 1671 – 1684 gaamctcggatcgtt in S. cerevisiae; one mismatch allowed), ITS 2 (positions 99 – 113 tcatttacttttaag in type species; one mismatch allowed), and 5.8 S (positions 119 – 133 in type species and 120 – 134 in S. cerevisiae ataggtattcctrtg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Pantelleriaceae, covering sequences EUK 1200019 and EUK 1137930 (Figs 1, 2).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E1A8C972094256BFAF1B0DCA2BFA8931.taxon	type_taxon	Type genus. Pantelleria Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E1A8C972094256BFAF1B0DCA2BFA8931.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in SSU V 9 (positions 1671 – 1684 in S. cerevisiae gaamctcggatcgtt; one mismatch allowed), and 5.8 S (positions 119 – 133 in type species and 120 – 134 in S. cerevisiae ataggtattcctrtg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Pantelleriales, covering sequences EUK 1200019 and EUK 1137930 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5920EF54D087557FBA2E65AA00F2FDEA.taxon	type_taxon	Type family. Pantelleriaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
5920EF54D087557FBA2E65AA00F2FDEA.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in SSU V 7 (positions 1389 – 1408 in S. cerevisiae ctatcgacgtwtagtcgatg; no mismatch allowed). Forms a monophyletic, least inclusive clade in Pantelleriomycetes, covering sequences EUK 1200019, EUK 1137930, EUK 1120813, and EUK 1700280 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
199821CC2B6558E3A970E6760E830C88.taxon	type_taxon	Type order. Pantelleriales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
199821CC2B6558E3A970E6760E830C88.taxon	diagnosis	Diagnosis. Distinguishable from other Aphelidiomycota based on diagnostic nucleotide signature in LSU 5 ’ end (positions 42 – 51 in type species and S. cerevisiae tatcattaag; no mismatch allowed). Forms a monophyletic, least inclusive clade in Aphelidiomycota, covering sequences EUK 1203048, EUK 1205018, EUK 1200019, EUK 1137930, EUK 1120815, EUK 1103262, GU 568135, EUK 1200059, EUK 1138870, EUK 1138872, EUK 1120814, EUK 1102231, and EUK 1201826 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
710F5E4C87B45F4297A651672DF63862.taxon	diagnosis	Diagnosis. Separation from other species of Parakickxella based on ITS 2 (positions 98 – 117 cgtgaacatatggtgccccc; one mismatch allowed) and LSU D 2 (positions 444 – 463 in the type species and 436 – 455 in S. cerevisiae cgccgcgctgtttgtgcgcg; one mismatch allowed) as indicated in Fig. 48. Intraspecific difference up to 1.4 % in ITS 2 and up to 0.5 % in LSU. Interspecific distance at least 15.0 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
710F5E4C87B45F4297A651672DF63862.taxon	description	Description. Other sequences: EUK 1107422 (tropical rainforest soil in El Yunque, Puerto Rico, 18.29, – 65.78); EUK 0147219 (GSMc plot G 5034, tropical rainforest soil in Los Pinos, Puerto Rico, 18.1268, – 66.0724 ° E); and GlobalFungi accession dfbf 3 c 41964 a 835260 bb 3 a 9 afcdaf 69 a (tropical rainforest soil in Luquillo, Puerto Rico, 18.3, – 65.8).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
710F5E4C87B45F4297A651672DF63862.taxon	etymology	Etymology. Parakickxella (Latin) refers to a taxon distant from Kickxella, and Boriken (Taino) refers to the native name of Puerto Rico, where the type material and other specimens originate.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F3551AD13938564285D33636089EAA44.taxon	type_taxon	Type species. Parakickxella borikenica Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F3551AD13938564285D33636089EAA44.taxon	diagnosis	Diagnosis. Separation from other fungi based on diagnostic nucleotide signatures in SSU V 3 (positions 677 – 691 in S. cerevisiae gttccgcccggtctc; one mismatch allowed) and LSU D 2 (positions 697 – 711 in the type species and 687 – 701 in S. cerevisiae cgacacgtcatggtg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Parakickxellaceae, covering sequences EUK 1189325, EUK 1189316, and EUK 1189321 (Figs 1, 47).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4729255D8B905E8AA5F35D7EFD865685.taxon	type_taxon	Type genus. Parakickxella Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4729255D8B905E8AA5F35D7EFD865685.taxon	diagnosis	Diagnosis. Separation from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 117 – 126 in the type species and 120 – 129 in S. cerevisiae tggattactc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Parakickxellales, covering sequences EUK 1700155, EUK 1189325, EUK 1189316, and EUK 1189321 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E71D5B4E964C5D1E9D0A2B8934C66393.taxon	type_taxon	Type family. Parakickxellaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E71D5B4E964C5D1E9D0A2B8934C66393.taxon	diagnosis	Diagnosis. Separation from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1476 – 1495 in S. cerevisiae ccaagkcaacgagtytacaa; one mismatch allowed) and LSU D 1 (positions 184 – 198 in type species and 180 – 194 in S. cerevisiae ggtataatttgcctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Parakickxellomycetes, covering sequences EUK 1189320, EUK 1189314, EUK 1100806, EUK 1189309, UDB 014747, EUK 1107625, EUK 1105293, EUK 1189310, EUK 1700155, EUK 1189325, EUK 1189316, and EUK 1189321 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CBF6F269FC8856AE908EFC1807888A25.taxon	type_taxon	Type order. Parakickxellales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
CBF6F269FC8856AE908EFC1807888A25.taxon	diagnosis	Diagnosis. Separation from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1476 – 1495 in S. cerevisiae ccaagkcaacgagtytacaa; two mismatches allowed) and LSU D 1 (positions 184 – 198 in type species and 180 – 194 in S. cerevisiae ggtataatttgcctg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Kickxellomycota, covering sequences EUK 1189320, EUK 1189314, EUK 1100806, EUK 1189309, UDB 014747, EUK 1107625, EUK 1105293, EUK 1189310, EUK 1700155, EUK 1189325, EUK 1189316, and EUK 1189321 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
29EC75DDEBF85828A95E588A79B3F4CD.taxon	diagnosis	Diagnosis. Separation from other species of Paraspizellomyces based on ITS 2 (positions 220 – 239 tttatgaattartgattgta; no mismatch allowed) and LSU (positions 562 – 581 ccgagtgttatagcctgagg; no mismatch allowed) as indicated in Fig. 5. Intraspecific variation up to 3.7 % in ITS 2. Interspecific distance at least 3.7 % in ITS 2; i. e., there is no clear barcode gap in ITS 2. In ITS 1, the maximum intraspecific difference 2.4 %, and the minimum interspecific distance 3.3 %.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
29EC75DDEBF85828A95E588A79B3F4CD.taxon	description	Description. Other sequences: EF 619657 (soil in Pinus taeda plantation, NC, USA); EUK 0327288 (GSMc plot G 6004, tropical rainforest soil in Cascada Julieta in Panama, 9.2274, – 79.4312); EUK 0327292 (GSMc plot G 4982, subtropical rainforest soil in Weiloi, Meghalaya, India, 25.3570 ° N, 91.6060 ° E); EUK 0327297 (GSMc plot S 372, subtropical forest soil in Menglun, Yunnan, China, 21.572 ° N, 101.57 ° E); EUK 0327298 (GSMc plot S 013, tropical woodland soil in Isalo, Madagascar, – 22.5339, 45.3703); EUK 0327299 (GSMc plot S 765, tropical rainforest soil in Mbomole, Tanzania, – 5.0946, 38.6292); and EUK 0519411 (GSMc plot S 1190, tropical rainforest soil in La Palma, Costa Rica, 10.5046, – 84.6949).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
29EC75DDEBF85828A95E588A79B3F4CD.taxon	etymology	Etymology. Para (Greek) and Spizellomyces (Latin) refer to phylogenetic relatedness to Spizellomycetales, and Parrent (English) refers to Jeri Lynn Parrent, who was the first to collect material from this species (EF 619657; Parrent and Vilgalys 2007).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
75DAB7E4C1B75619BFF38377DCF725F0.taxon	type_taxon	Type species. Paraspizellomyces parrentiae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
75DAB7E4C1B75619BFF38377DCF725F0.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 1 (positions 228 – 237 in type species and 227 – 236 in S. cerevisiae taacgaccca; one mismatch allowed) and SSU V 9 (positions 1695 – 1709 in S. cerevisiae aatttcggttgctgg or agtttcggccgctgg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Paraspizellomycetaceae, covering sequences EUK 1187441, EUK 1187447, UDB 014658, and EUK 1100628 (Figs 1, 4).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2C2903E2456552488C52A534C8F54406.taxon	type_taxon	Type genus. Paraspizellomyces Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2C2903E2456552488C52A534C8F54406.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 2 (positions 578 – 592 in type species and 562 – 576 in S. cerevisiae aaggtcatgcttt; one mismatch allowed) and ITS 2 (positions 134 – 148 in type species aatggttcccaagtg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Paraspizellomycetales, covering sequences EUK 1187441, EUK 1187447, UDB 014658, EUK 1100628, EUK 1123745, and EUK 1139262 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A5C16A3D11ED577CA5825644C11E5AAD.taxon	type_taxon	Type family. Paraspizellomycetaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A5C16A3D11ED577CA5825644C11E5AAD.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 1 (positions 128 – 142 in type species and 123 – 137 in S. cerevisiae cggttcgccggtgcg or gggttcttacctatg or gggttccacctatgc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Spizellomycetes, covering sequences EUK 1138322, EUK 1152022, EUK 1187448, EUK 1202178, EUK 1187441, EUK 1187447, UDB 014658, EUK 1100628, EUK 1123745, and EUK 1139262 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
52332B84FF125789A6A676801B58DE74.taxon	type_taxon	Type phylum. Rozellomycota Doweld.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
52332B84FF125789A6A676801B58DE74.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EDEC19532BAA5A76B1C8DD6A00BFE92F.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A73530CCE5215BD5874CFF06EA9B533E.taxon	diagnosis	Diagnosis. Separation from other species of Ruderalia based on ITS 2 (positions 154 – 176 ggaggcttgaaattgagaaaaag; one mismatch allowed) and LSU D 2 (positions 583 – 607 cctcgggaatgtgatccgcctttac; one mismatch allowed) as indicated in Fig. 36. Intraspecific variation up to 9.8 % (including up to 7 % in the type locality) in ITS 2 due to multiple microsatellite-like repeats and long homopolymers and up to 1.5 % in LSU. Interspecific distance> 20 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A73530CCE5215BD5874CFF06EA9B533E.taxon	description	Description. Other sequences: EUK 1137942, EUK 1137899 and EUK 1124460 (type locality); EUK 0018130 (GSMc plot S 150, Quercus woodland soil in Jamestown, CA, USA, 37.8489, – 120.581); EUK 0531861 (temperate grassland soil in Murrietta, CA, USA, 33.5319, – 117.2492 ° E), EUK 0015594 (GSMc plot EO 077, Cistus shrubland soil in Essouira, Morocco, 31.5136, – 9.6542 ° E); EUK 0531686 (GSMc plot G 6066, subtropical Vachellia desert soil in Al Mudawih, Saudi Arabia, 25.8411 ° N, 39.2793 ° E); EUK 0531846 (GSMc plot G 6106, cropland soil in Betas, Iraqi Kurdistan, 37.0588 ° N, 42.7622 ° E); EUK 0531869 (GSMc plot G 5748, subtropical forest soil in Cebollati, Uruguay, – 33.8292, – 54.7672 ° E); and EUK 0531757 (subtropical shrubland soil in Los Panguiles, Chile, – 33.3822, – 70.9636 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A73530CCE5215BD5874CFF06EA9B533E.taxon	etymology	Etymology. Rudus (Latin) refers to a common habitat in early successional land, and cosmopolites (Greek) refers to the global distribution.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2FD5110D0131591B9339C1E8BDB57A89.taxon	type_taxon	Type species. Ruderalia cosmopolita Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2FD5110D0131591B9339C1E8BDB57A89.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 209 – 222 aacgatagtgaagt; two mismatches allowed). Forms a monophyletic, least inclusive clade in Ruderaliaceae, covering sequences EUK 1138161, EUK 1137899, EUK 0531800, and EUK 1124460 (Figs 1, 35).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
40AEC11BE10C559B9B015AD535B19F17.taxon	type_taxon	Type family. Ruderaliaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
40AEC11BE10C559B9B015AD535B19F17.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 631 – 640 in the type species and 564 – 573 in S. cerevisiae acggatacgg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Ruderaliomycetes, covering sequences EUK 1138161, EUK 1137899, EUK 0531800, EUK 1124460, EUK 1103744, EUK 1103025, EUK 1103555, EUK 1103599 and EUK 1203462 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D98BD96618B45B318C9F3AC3F9B47D2D.taxon	type_taxon	Type order. Ruderaliales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D98BD96618B45B318C9F3AC3F9B47D2D.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 631 – 640 in the type species and 564 – 573 in S. cerevisiae acggatacgg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Mortierellomycota, covering sequences EUK 1138161, EUK 1137899, EUK 1124460, EUK 0531800, EUK 1103744, EUK 1103025, EUK 1103555, EUK 1103599, EUK 1203462, and EUK 1700231 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
17B16EC9CD0F57F7BBB44B77104B0317.taxon	type_taxon	Type genus. Savannolpidium Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
17B16EC9CD0F57F7BBB44B77104B0317.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in the ITS 2 - LSU interface (LSU positions – 2 – 18 in type species and S. cerevisiae aagtgatctgaaatcagaca; one mismatch allowed) and ITS 2 (positions 137 – 151 in type species gcgtactccttgtcc; two mismatches allowed) and SSU V 8 (positions 1589 – 1608 in S. cerevisiae atgattcatcagatcatgct; one mismatch allowed). Forms a monophyletic, least inclusive clade in Savannolpidiales, covering sequences EUK 1191209, EUK 1191210, EUK 0534704, EUK 1124874, EUK 1701673, EUK 1701672, and EUK 1124875 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8308B450488C50FEB24AE534FED886B0.taxon	type_taxon	Type family. Savannolpidiaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8308B450488C50FEB24AE534FED886B0.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in the ITS 2 - LSU interface (LSU positions – 2 – 18 in type species and S. cerevisiae aagtgatctgaaatcagaca; two mismatches allowed) and SSU V 8 (positions 1589 – 1608 in S. cerevisiae atgattcatcagatcatgct; two mismatches allowed). Forms a monophyletic, least inclusive clade in Savannolpidiomycetes, covering sequences EUK 1191209, EUK 1191210, EUK 0534704, EUK 1124874, EUK 1701673, EUK 1701672, and EUK 1124875 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E3DB8A3577075D3B9FA7A468A5225D7B.taxon	type_taxon	Type order. Savannolpidiales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E3DB8A3577075D3B9FA7A468A5225D7B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 8 (positions 1546 – 1565 in S. cerevisiae gagcattgcaactattgctc; one mismatch allowed) and LSU D 2 (positions 525 – 534 in the type species and 472 – 481 in S. cerevisiae gtgcactttt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Olpidiomycota, covering sequences EUK 1191172, EUK 1191209, EUK 1191210, EUK 0534704, EUK 1701673, EUK 1701672, EUK 1124874, and EUK 1124875 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
532E9F06099A53DABA96D10380A0B08B.taxon	diagnosis	Diagnosis. Separation from other species of Savannolpidium based on ITS 2 (positions 16 – 40 agatctcatcttctttagagttggc; no mismatch allowed) and LSU (positions 468 – 492 tataaagggaggctagtgtgagcgc; no mismatch allowed) as indicated in Fig. 42. Intraspecific variation up to 1.6 % in ITS 2 and 0.9 % in LSU. Interspecific distance at least 2.8 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
532E9F06099A53DABA96D10380A0B08B.taxon	description	Description. Other sequences: EUK 1138191 (type locality); EUK 1217298 (GSMc plot G 4777, flooded grassland soil Suur-Pakri Härs-hämani, Estonia, 59.3310 ° N, 23.9272 ° E); EUK 0332294 (GSMc plot G 5899, dry Juniperus shrubland soil in Virtsu, Estonia, 58.5775 ° N, 23.5547 ° E); EUK 0332296 (flooded grassland soil in Haanja, Estonia, 57.7165 ° N, 27.0549 ° E); EUK 0534754 (GSMc plot G 6107, Quercus woodland soil in Armishte, Iraqi Kurdistan, 37.0468 ° N, 42.8049 ° E); EUK 0584862 (FunAqua river sediment sample W 1356 s, Spey, Scotland, 57.0552, – 4.1276 ° E); EUK 0584863 (FunAqua saltwater sediment sample W 0938 s, Laguna di Orbetello, Italy, 42.4296 ° N, 11.1988 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
532E9F06099A53DABA96D10380A0B08B.taxon	etymology	Etymology. Savanna (Taino) refers to treeless grasslands, and Raadi (Estonian) refers to the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A04943B3E22E52C78C861297A3BE7DCA.taxon	type_taxon	Type species. Savannolpidium raadiense Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A04943B3E22E52C78C861297A3BE7DCA.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 121 – 135 in type species grtagtaaaagtagc; one mismatch allowed), SSU V 9 (positions 1684 – 1693 in S. cerevisiae ccttttttyg; one mismatch allowed), and LSU D 2 (positions 719 – 728 in type species and 604 – 613 in S. cerevisiae caaaaggatt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Savannolpidiaceae, covering sequences EUK 1124874 and EU 1191210 (Figs 1, 41).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
055F6432250F57D3BED325D94E8A684C.taxon	type_taxon	Type genus. Sedimentomastix Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
055F6432250F57D3BED325D94E8A684C.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in SSU V 9 (positions 1672 – 1686 in S. cerevisiae ggcctccggattgat; no mismatch allowed) and LSU D 2 (positions 524 – 538 in type species and 460 – 474 in S. cerevisiae ttggtattttgggtg; three mismatches allowed). Forms a monophyletic, least inclusive clade in Sedimentomastigales, covering sequences EUK 0319782, EUK 0574067, EUK 0574066, EUK 1152060, and EUK 1191154 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
250D47DE755950B083F1EAC510C5E3BE.taxon	type_taxon	Type family. Sedimentomastigaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
250D47DE755950B083F1EAC510C5E3BE.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1672 – 1686 in S. cerevisiae ggcctccggattgat; no mismatch allowed) and LSU D 2 (positions 524 – 538 in the type species and 460 – 474 in S. cerevisiae ttggtattttgggtg; three mismatches allowed). Forms a monophyletic, least inclusive clade in Sedimentomastigomycetes, covering sequences EUK 0319782, EUK 0574067, EUK 0574066, EUK 1152060, and EUK 1191154 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8156049A2BED5A05B833C8AC1677D7F6.taxon	type_taxon	Type order. Sedimentomastigales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
8156049A2BED5A05B833C8AC1677D7F6.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 9 (positions 1672 – 1686 in S. cerevisiae ggcctccggattgat; no mismatch allowed). Forms a monophyletic, least inclusive clade in Neocallimastigomycota, covering sequences EUK 0319782, EUK 0574067, EUK 0574066, EUK 1152060, and EUK 1191154 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EC39DD3CA5E45017BB474111732DEEF5.taxon	type_taxon	Type species. Sedimentomastix tueriensis Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
EC39DD3CA5E45017BB474111732DEEF5.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1672 – 1686 in S. cerevisiae ggcctccggattgat; no mismatch allowed) and LSU D 2 (positions 524 – 538 in the type species and 460 – 474 in S. cerevisiae ttggtattttgggtg; two mismatches allowed). Forms a monophyletic, least inclusive clade in Sedimentomastigaceae, covering sequences EUK 0319782, EUK 0574067, EUK 0574066, EUK 1152060, and EUK 1191154 (Figs 1, 25).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
81A33060ABF4555C81C3ACA7380CDD24.taxon	diagnosis	Diagnosis. Separation from other species of Sedimentomastix based on ITS 2 (positions 15 – 34 ctaaaagtcgggtttgattt; one mismatch allowed) and LSU D 2 (positions 572 – 591 gaaacaatggataaagggca; one mismatch allowed) as indicated in Fig. 26. Intraspecific variation up to 1.7 % in ITS 2. Interspecific distance> 15 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
81A33060ABF4555C81C3ACA7380CDD24.taxon	description	Description. Other sequences: EUK 0302143 (Populus tremula forest soil in Soinaste, Estonia, 58.3408 ° N, 26.6864 ° E); EUK 0584897 (FunAqua stream water sample in Kangilleq, Greenland, 60.8571, – 46.4233); EUK 0584898 (FunAqua river water sample W 0597 w in Aveleda, Portugal, 41.8919, – 6.6972); and EUK 0584899 (FunAqua lake sediment sample W 0307 s in Goldwin, ND, USA, 47.0996, – 99.0916).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
81A33060ABF4555C81C3ACA7380CDD24.taxon	etymology	Etymology. Sedimentomastix refers to sedimentum (the Latin term for sediment) and Neocallimastix; the epithet refers to Türi (Estonian), the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6DD4E2A76002570CB923F64DED31CFC5.taxon	type_taxon	Type genus. Solivorax Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
6DD4E2A76002570CB923F64DED31CFC5.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions in type species gctaagttta; two mismatches allowed) and LSU D 2 (positions 694 – 703 in type species and 596 – 605 in S. cerevisiae ctaacgtgct; one mismatch allowed). Forms a monophyletic, least inclusive clade in Solivoracales, covering sequences OQ 687305, OQ 687310, OQ 687311, EUK 1216850, DQ 244008, UDB 014650, EUK 1124454, EUK 1216854, EUK 1216849, and OQ 687309 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
7CB6B20299DA52B3BA5AC25BE06DF806.taxon	type_taxon	Type family. Solivoracaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
7CB6B20299DA52B3BA5AC25BE06DF806.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 694 – 703 in type species and 596 – 605 in S. cerevisiae ctaacgtgct or cctttgtgct; one mismatch allowed). Forms a monophyletic, least inclusive clade in Algovoracomycetes, covering sequences OQ 687305, OQ 687310, OQ 687311, EUK 1216850, DQ 244008, UDB 014650, EUK 1124454, EUK 1216854, EUK 1216849, and OQ 687309 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
24CB805A51775C519CD4DB95B1B27B14.taxon	diagnosis	Diagnosis. Separation from other species of Solivorax based on ITS 2 (positions 273 – 292 gtctgaccgaaatatctgaa; one mismatch allowed) and LSU D 2 (positions 672 – 691 gcctgctatgctagcgccgc; one mismatch allowed) as indicated in Fig. 16. Intraspecific variation up to 2.4 % in ITS 2. Interspecific distance at least 8.2 % in ITS 2 and 6.2 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
24CB805A51775C519CD4DB95B1B27B14.taxon	description	Description. Other sequences: EUK 0484226 (GSMc plot S 1278, Eucalyptus plantation soil near Durban, South Africa, – 29.7502, 30.7419); EUK 0331319 (GSMc plot G 5027, subtropical swamp forest soil in Deweyville, LO, USA, 30.3076 ° N, 93.7304 ° E); EUK 0331320 (GSMc plot S 915, tropical dry forest soil in Rancho Calimayor, Mexico, 16.5461, – 93.8828); EUK 0331312 (GSMc plot G 5687, tropical garden soil in Kakamega, Kenya, 0.2866 ° N, 34.7673 ° E); EUK 0331316 (GSMc plot JYK 054, Eucalyptus plantation soil in Bome, Liberia, 6.5245, – 10.8400); EUK 0331318 (GSMc plot S 1267, tropical rainforest soil in Khong Ngam, Thailand, 19.6691 ° N, 99.8199 ° E); and EUK 0331314 (GSMc plot G 5068, tropical rainforest soil in Quixada, Brazil, 4.8876, – 39.0461).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
24CB805A51775C519CD4DB95B1B27B14.taxon	etymology	Etymology. Solum and vorax (Latin) refer to soil devouring, and pantropicos (Greek) refers to widespread distribution across tropical regions.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4BAB13E32D145F278C95563FCC10068D.taxon	type_taxon	Type species. Solivorax pantropicus Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4BAB13E32D145F278C95563FCC10068D.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in ITS 2 (positions 85 – 102 in type species gtaccgctaagtttaagc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Solivoracaceae, covering sequences UDB 014650, EUK 1124454, EUK 1216854, EUK 1216849, and OQ 687309 (Figs 1, 14).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
023AAF1BB47D52E7BB9E82FD194D8C37.taxon	type_taxon	Type order. Spizellomycetales D. J. S. Barr.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
023AAF1BB47D52E7BB9E82FD194D8C37.taxon	description	Description. As in Tedersoo et al. (2018).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
352963399E315BD4A4A5F8B5BE91F994.taxon	type_taxon	Type genus. Sumavosporidium Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
352963399E315BD4A4A5F8B5BE91F994.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 124 – 138 in the type species and 125 – 139 in S. cerevisiae gcaatcygcaggcat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Sumavosporidiales, covering sequences UDB 029033, UDB 029043, UDB 029027, UDB 028954, EUK 1104656, EUK 1106151, and EUK 1104875 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3A792B941CDB5E6EA9CBC500633FB3B4.taxon	type_taxon	Type family. Sumavosporidiaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3A792B941CDB5E6EA9CBC500633FB3B4.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 6 (positions 1157 – 1176 in S. cerevisiae acaccaaaagtggattttgc or acaccaagagtggagcatgc or acacaagaagtggagcctgc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Sumavosporidiomycetes, covering sequences EUK 1206927, EUK 1202246, EUK 1200658, UDB 029033, EUK 1105717, EUK 1107386, EUK 1106576, EUK 1101061, and EUK 1101529 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
38C0201E8A1C582AAB5FD2F35ED529AA.taxon	type_taxon	Type order. Sumavosporidiales Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
38C0201E8A1C582AAB5FD2F35ED529AA.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in SSU V 6 (positions 1153 – 1167 gagcacaccaaragt or gacgacacaagaagt in S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Rozellomycota, covering sequences EUK 1206927, EUK 1202246, EUK 1200658, UDB 029033, EUK 1105717, EUK 1107386, EUK 1106576, EUK 1101061, and EUK 1101529 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2E70C63A1BB053A5B3FC3A332E8A0A5A.taxon	diagnosis	Diagnosis. Separation from other species of Sumavosporidium based on ITS 2 (positions 7 – 31 gaatgaagatgtgatcgaactgtgc; one mismatch allowed) and LSU (positions 465 – 484 caactagttggccttcaggt; one mismatch allowed) as indicated in Fig. 46. Intraspecific variation up to 2.0 % in ITS 2 and up to 0.2 % in LSU. Interspecific distance at least 12.0 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2E70C63A1BB053A5B3FC3A332E8A0A5A.taxon	description	Description. Other sequences: UDB 014611 and EUK 0482169 (type locality); UDB 029027, UDB 029028 and UDB 014612 (GSMc plot S 121, Fagus sylvatica forest soil in Taunus, Germany, 50.1413 ° N, 8.2677 ° E); EUK 0482019 and EUK 0520242 (GSMc plot S 426, Fagus sylvatica forest soil in Kistrupvang, Denmark, 56.0264 ° N, 12.3364 ° E); HQ 022097 (mixed forest soil in Bartlett Experimental Forest, NH, USA, 44.06, – 71.30); EUK 0522425 and EUK 0522433 and (GSMc plot G 4145, mixed deciduous forest soil in Promised Land, PA, USA, 41.30491, – 75.2021 ° E); EUK 0523233 (GSMc plot S 878, Alnus alnobetula tundra soil in Anadyr, Chukotka, Russia, 64.7219 ° N, 177.4238 ° E); EUK 0034322 (GSMc plot G 4713, Tsuga mertensiana forest soil in Crater Lake, OR, USA, 42.9786, – 122.13); and EUK 0521849 (GSMc plot S 892, forest tundra soil in Arman, Magadan, Russia, 59.6972 ° N, 150.4118 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2E70C63A1BB053A5B3FC3A332E8A0A5A.taxon	etymology	Etymology. Šumava (Czech) refers to the type locality, and sylva (Latin) refers to the forest habitat.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
006E10ABBF53574BB8D6111E43179016.taxon	type_taxon	Type species. Sumavosporidium sylvestre Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
006E10ABBF53574BB8D6111E43179016.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in 5.8 S (positions 124 – 138 in the type species and 125 – 139 in S. cerevisiae gcaatcygcaggcat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Sumavosporidiaceae, covering sequences UDB 029033, UDB 029043, UDB 029027, UDB 028954, and EUK 1104656 (Figs 1, 45).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
628A60FCB5055A1D867B4E2A35FE1DD4.taxon	diagnosis	Diagnosis. Separation from other species of Tartumyces based on ITS 2 (positions 310 – 329 ggggggtataaaaactcgtt; one mismatch allowed) and LSU D 2 (positions 518 – 539 tattcgccggataatggtac; no mismatch allowed) as indicated in Fig. 62. Intraspecific variation up to 2.8 % in ITS 2. Interspecific distance at least 4.5 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
628A60FCB5055A1D867B4E2A35FE1DD4.taxon	description	Description. Other sequences: EUK 1703744 (GSMc plot G 4372, mixed forest soil in Kiisli, Estonia, 58.6955 ° N, 26.9128 ° E); EUK 1703739 (GSMc plot G 3569, Quercus robur park soil in Äksi, Estonia, 58.5290 ° N, 26.6385 ° E); and EUK 1703737 (GSMc plot G 3413, Salix caprea forest soil in Väägvere, Estonia, 58.4389 ° N, 26.8976 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
628A60FCB5055A1D867B4E2A35FE1DD4.taxon	etymology	Etymology.> Tartu (Estonian) refers to the city and county in Estonia, where the type material and most other specimens were collected. The epithet refers to Kensuke Seto, the first to obtain coarse single-cell photographs of species belonging to this phylum (Seto et al. 2023).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
00ECABB66B0C5FF0A9629C819B8D84CF.taxon	type_taxon	Type species. Tartumyces setoi Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
00ECABB66B0C5FF0A9629C819B8D84CF.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 289 – 300 in type species gggtttgcaaac, one mismatch allowed) and LSU D 4 (positions 624 – 633 in type species and 601 – 610 in S. cerevisiae gaatttattc, one mismatch allowed). Forms a monophyletic, least inclusive clade in Tartumycetaceae, covering sequences EUK 1123648 and EUK 1138300 (Figs 1, 61).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E8A6F21799655ACE93B423E828402B65.taxon	type_taxon	Type genus. Tartumyces Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E8A6F21799655ACE93B423E828402B65.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 173 – 187 in type species ggaaagcgtagtagg, two mismatches allowed) and LSU D 4 (positions 1439 – 1453 in type species and 1404 – 1418 in S. cerevisiae gatgccgcgtcgaac, one mismatch allowed). Forms a monophyletic, least inclusive clade in Tartumycetales, covering sequences EUK 1123648, EUK 1138300, EUK 1186160, EUK 1186168, and EUK 1186172 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
082BFA7199D450DEAA6CAF8F0A69374A.taxon	type_taxon	Type family. Tartumycetaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
082BFA7199D450DEAA6CAF8F0A69374A.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 120 – 129 in type species gaaccaaagg, one mismatch allowed) and LSU D 1 (positions 164 – 178 in type species and 161 – 175 in S. cerevisiae gatgcctgtgggagc, one mismatch allowed). Forms a monophyletic, least inclusive clade in Tartumycetes, covering sequences EUK 1186157, EUK 1200073, EUK 1186162, EUK 1123648, EUK 1138300, OQ 702816, ON 754309, UDB 028835, and HQ 191300 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0E6A3AF0B186574EAFC3CB2585907523.taxon	type_taxon	Type order. Tartumycetales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
0E6A3AF0B186574EAFC3CB2585907523.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 4 (positions 1439 – 1453 in type species and 1404 – 1418 in S. cerevisiae gatgccgcgtcgaac, one mismatch allowed). Forms a monophyletic, least inclusive clade in Tartumycota, covering sequences EUK 1186157, EUK 1200073, EUK 1186162, EUK 1123648, EUK 1138300, OQ 702816, ON 754309, UDB 028835 and HQ 191300 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D0475B58916D5516A0EAA122F9DA3B69.taxon	type_taxon	Type class. Tartumycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
D0475B58916D5516A0EAA122F9DA3B69.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 3 (positions 1009 – 1023 in the type species and 969 – 983 in S. cerevisiae ggaacttgtacagtt, no mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences OQ 702815, EUK 1186161, OQ 687331, EUK 1186165, EUK 1186157, EUK 1200073, EUK 1186162, EUK 1123648, EUK 1138300, OQ 702816, ON 754309, UDB 028835, and HQ 191300 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
92F02C2B343D50158C6102449E3D9EF9.taxon	type_taxon	Type species. Terrincola waldropii Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
92F02C2B343D50158C6102449E3D9EF9.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in ITS 2 (positions 23 – 32 ggccgtacgg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Terrincolaceae, covering sequences MW 791967, EUK 0473585, and EUK 1138132 (Figs 1, 27).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
75FB219161D0507C8807A426187E95D5.taxon	diagnosis	Diagnosis. Separation from other species of Terrincola based on diagnostic nucleotide signatures in ITS 2 (positions 359 – 383 atagatgggacccggtcgaggatca; one mismatch allowed) and LSU D 2 (positions 569 – 588 agtcctctatttgtacaatg; one mismatch allowed) as indicated in Fig. 28. Intraspecific variation up to 1.2 % in ITS 2 and up to 0.5 % in LSU sequences. Interspecific distance at least 4.9 % in ITS 2 and 3.9 % in LSU.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
75FB219161D0507C8807A426187E95D5.taxon	description	Description. Other sequences: EUK 1138131 (GSMc plot G 5295, Pinus mugo plantation soil in Märja, Estonia, 58.3592 ° N, 26.6443 ° E); EUK 0326024 (GSMc plot S 1087, tundra soil in Zackenberg, Greenland, 74.4682, – 20.6142); EUK 0326022 (GSMc plot G 5038, tropical dry forest soil in West End, British Virgin Islands, 18.3907, – 64.7073); EUK 0326084 (GSMc plot S 639, subtropical forest soil in Platbos, South Africa, – 34.5676, 19.4461); EUK 0326080 (GSMc plot S 618, montane desert soil in Tanglang La, India, 33.5051 ° N, 77.7655 ° E); EUK 0326071 (GSMc plot G 5769, temperate grassland soil in Rõõmu, Estonia, 58.3877 ° N, 26.7770 ° E); and DQ 421306 (temperate grassland soil in Cedar Creek, MN, USA, 45.40, – 93.19).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
75FB219161D0507C8807A426187E95D5.taxon	etymology	Etymology. Terra and incola (Latin) refer to the soil habitat, and Waldrop (English) refers to the last name of Mark P. Waldrop, who was the first to collect material of this species and order (DQ 421306; Waldrop et al. 2006).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E4BE1CA3D6EC5C598491277CCC6A32FF.taxon	type_taxon	Type genus. Terrincola Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E4BE1CA3D6EC5C598491277CCC6A32FF.taxon	diagnosis	Diagnosis. Distinguishable from other members of Calcarisporiellomycota based on diagnostic nucleotide signature in ITS 2 (positions 285 – 292 aaaatrtt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Terrincolales, covering sequences MW 791967, EUK 1138132, EUK 1123677, EUK 0332618, EUK 1123675, and EUK 1123676 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BCA3DB630FC95F7D9479CA4F0F8FB536.taxon	type_taxon	Type family. Terrincolaceae Tedersoo & Esmaeilzadeh-Salestani.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BCA3DB630FC95F7D9479CA4F0F8FB536.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in 5.8 S (positions 6 – 26 in type species and S. cerevisiae ttcaacaatggatccctcg; no mismatch allowed), LSU D 1 (positions 4 – 23 in type species and S. cerevisiae tcctcaaatcaagcaagagt; no mismatch allowed), LSU D 2 (positions 255 – 264 in type species and 244 – 253 in S. cerevisiae ttggtagtgg; one mismatch allowed), and SSU V 3 (positions 647 – 651 ggcttg in S. cerevisiae; no mismatch allowed). Forms a monophyletic, least inclusive clade in Calcarisporiellomycetes, covering sequences MW 791967, EUK 1138132, EUK 1123677, EUK 0332618, EUK 1123675, EUK 1604147, EUK 1604155, and EUK 1123676 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
29B6676850255C88ACB2924703FD0A6C.taxon	type_taxon	Type genus. Tibetochytrium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
29B6676850255C88ACB2924703FD0A6C.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 722 – 736 in S. cerevisiae gtgggttagggatcc; one mismatch allowed), 5.8 S (positions 120 – 134 in type species and S. cerevisiae gctggtattccggcg; one mismatch allowed), and LSU D 2 (positions 505 – 619 in type species and 600 – 614 in S. cerevisiae ggcttagctggatac; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tibetochytriales, covering sequences EUK 1186747, EUK 1123755, and EUK 1186750 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B1584609431355B984D9CCE38CE198A1.taxon	type_taxon	Type family. Tibetochytriaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B1584609431355B984D9CCE38CE198A1.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 722 – 736 in S. cerevisiae gtgggttagggatcc or gtgggttagggagct; one mismatch allowed), 5.8 S (positions 120 – 134 in type species and S. cerevisiae gctggtattccggcg or tttggtatcccgaag; one mismatch allowed), and LSU D 2 (positions 505 – 619 in type species and 600 – 614 in S. cerevisiae ggcttagctggatac or agcttttgcagggat; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tibetochytriomycetes, covering sequences EUK 1186747, EUK 1123755, EUK 1186750, EUK 1102822, and EUK 1186746 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
88EC7C3A54845D7992EDEFCFDC2FA99E.taxon	type_taxon	Type order. Tibetochytriales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
88EC7C3A54845D7992EDEFCFDC2FA99E.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 722 – 736 in S. cerevisiae gtgggttagggatcc or gtgggttagggagct; one mismatch allowed), 5.8 S (positions 120 – 134 in type species and S. cerevisiae gctggtattccggcg or tttggtatcccgaag; one mismatch allowed), and LSU D 2 (positions 505 – 619 in type species and 600 – 614 in S. cerevisiae ggcttagctggatac or agcttttgcagggat; two mismatches allowed). Forms a monophyletic, least inclusive clade in Chytridiomycota, covering sequences EUK 1186747, EUK 1123755, EUK 1186750, EUK 1102822, and EUK 1186746 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4499B23FFEAB5A70AE9ABE273C68F07B.taxon	diagnosis	Diagnosis. Separation from other species of Tibetochytrium based on ITS 2 (positions 85 – 104 ttggctatatctcgctttga; one mismatch allowed) and LSU (positions 656 – 675 ctgattgtcagtggagccat; no mismatch allowed) as indicated in Fig. 11. Intraspecific variation up to 3.8 % in ITS 2 and up to 1.0 % in LSU. Interspecific distance> 20 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4499B23FFEAB5A70AE9ABE273C68F07B.taxon	description	Description. Other sequences: EUK 1186749 (GSMc plot S 949; boreal coniferous forest soil in Mt. Mayak, Altai, Russian Federation, 51.0443 ° N, 82.9694 ° E); EUK 1186750 (GSMc plot S 958, temperate broadleaf forest soil in Měšice, Czechia, 50.2006 ° N, 14.5284 ° E); EUK 1186752 (GSMc plot S 966; temperate broadleaf forest soil in Orlík nad Vltavou, Czechia, 49.5002 ° N, 14.1742 ° E); OW 841378 (unspecified soil in Tianshan Mountains, Uyghuria, China); MW 215915 (rhizosphere soil in Lithuania); EF 434111 (boreal forest soil in Bonanza Creek, AK, USA); EUK 0519405 (GSMc plot S 1406, grassland soil in Chuy, Kyrgyzstan, 42.5502 ° N, 74.5121 ° E); GU 311731 (grassland soil in KS, USA); OX 032019 (Festuca brevipila roots in temperate grassland, Mallnow, Germany).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4499B23FFEAB5A70AE9ABE273C68F07B.taxon	etymology	Etymology. Tibet (Tibetan) refers to the region of the type habitat, and Taylor (English) refers to the last name of D. Lee Taylor, who was the first to collect material of this species (EF 434111; Taylor et al. 2007).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2DE7214597C355DD8A88082191CDECB6.taxon	type_taxon	Type species. Tibetochytrium taylorii Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
2DE7214597C355DD8A88082191CDECB6.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in ITS 2 (positions 100 – 119 in type species ttgacagacttacgcgtctt; two mismatches allowed), LSU D 2 (positions 552 – 571 in type species and 547 – 566 in S. cerevisiae aaagtgttatagcttttcat; two mismatches allowed), and SSU V 9 (positions 1700 – 1714 in S. cerevisiae caacgaaaatagatt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tibetochytriaceae, covering sequences EUK 1123755 and EUK 1186750 (Figs 1, 10).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E98E6ABF02375E69BFD6009170375C7B.taxon	type_taxon	Type genus. Tropicochytrium Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
E98E6ABF02375E69BFD6009170375C7B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 841 – 850 in S. cerevisiae tccgggracc; no mismatch allowed) and LSU D 2 (positions 598 – 607 in the type species and 592 – 601 in S. cerevisiae agcagcgctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tropicochytriales, covering sequences EUK 1102342, EUK 1186762, EUK 1100009, EUK 1186758, EUK 0519487, EUK 1186756, and EUK 1102527 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
862DE849425A57518C0E005AB0B227E3.taxon	type_taxon	Type family. Tropicochytriaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
862DE849425A57518C0E005AB0B227E3.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 841 – 850 in S. cerevisiae tccgggracc; no mismatch allowed) and LSU D 2 (positions 598 – 607 in the type species and 592 – 601 in S. cerevisiae agcagcgctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tropicochytriomycetes, covering sequences EUK 1102342, EUK 1186762, EUK 1100009, EUK 1186758, EUK 0519487, EUK 1186756, and EUK 1102527 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C9A27E99923A52A89A23890C55A5F4AF.taxon	type_taxon	Type order. Tropicochytriales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C9A27E99923A52A89A23890C55A5F4AF.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 4 (positions 841 – 850 in S. cerevisiae tccgggracc; no mismatch allowed) and LSU D 2 (positions 598 – 607 in the type species and 592 – 601 in S. cerevisiae agcagcgctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Chytridiomycota, covering sequences EUK 1102342, EUK 1186762, EUK 1100009, EUK 1186758, EUK 0519487, EUK 1186756, and EUK 1102527 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B5DD887FEC7E5FD6849069B66C9792E2.taxon	type_taxon	Type species. Tropicochytrium toronegroense Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
B5DD887FEC7E5FD6849069B66C9792E2.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signature in ITS 2 (positions 108 – 127 in type species ctcgtggtccgcaaggcttt; one mismatch allowed) and LSU D 2 (positions 557 – 577 in type species and 549 – 569 in S. cerevisiae agtttatagcctccggtcctg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Tropicochytriaceae, covering sequences EUK 1186758, EUK 0519487, and EUK 1186756 (Figs 1, 12).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BC7695C5B463568B95622A35EEAE9C2D.taxon	diagnosis	Diagnosis. Separation from other species of Tropicochytrium based on ITS 2 (positions 182 – 201 gggggcctcgtctccccttt; one mismatch allowed) and LSU D 2 (positions 536 – 555 gaccccgccctcacgggtgg; no mismatch allowed) as indicated in Fig. 13. Intraspecific variation up to 2.1 % in ITS 2. Interspecific distance at least 3.1 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BC7695C5B463568B95622A35EEAE9C2D.taxon	description	Description. Other sequences: EUK 0649723 (GSMc plot MX 35, Pinus chiapensis - dominated tropical forest, Mecacalvo, Veracruz, Mexico, 19.7760, – 97.1016); EUK 0649724 (GSMc plot S 914, tropical forest in Lagos de Monte Bello, Chiapas, Mexico, 16.1004, – 91.6871); EUK 0649725 (GSMc plot G 5037, tropical forest in Maricao, Puerto Rico, 18.1450, – 66.9669); EUK 0137040, EUK 0474865 and EUK 0519433 (all GSMc plot S 381, tropical forest soil in Col Palmarena, Costa Rica, 10.2211, – 84.5992); and EUK 0474859 and EUK 0519530 (both GSMc plot JYK 042, tropical forest soil in Barclayville, Liberia, 4.6777 ° N, 8.1230 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
BC7695C5B463568B95622A35EEAE9C2D.taxon	etymology	Etymology. Tropica (Greek) refers to the tropics, where the genus mainly occurs; Toro Negro (Spanish) refers to the type locality.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3AE386D8257B5AF8A7DA8D20FE40BC6C.taxon	diagnosis	Diagnosis. Separation from other species of Viljandia based on ITS 2 (positions 73 – 92 ggattgcatggactgccgtc; one mismatch allowed) and LSU (positions 594 – 613 gcaaagctaccgtgtccaga; one mismatch allowed) as indicated in Fig. 58. Intraspecific variation up to 7.5 % in ITS 2 and up to 2.2 % in LSU. Interspecific distance> 20 % in ITS 2.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3AE386D8257B5AF8A7DA8D20FE40BC6C.taxon	description	Description. Other sequences: EUK 1632579 (GSMc plot G 4506, woodland soil in Terikeste Hiiepärna, Estonia, 58.2972 ° N, 27.0681 ° E); LC 204214 (Picea crassifolia temperate forest soil in Inner Mongolia, China, 38.77 ° N, 105.89 ° E); EUK 1124345 (GSMc plot G 5901, Aesculus hippocastanum alley soil in Tartu, Estonia, 58.3676 ° N, 26.7255 ° E); EUK 1105441 (boreal coniferous forest soil near Hofors, Sweden, 60.49 ° N, 16.3 ° E); EUK 1216896 (GSMc plot G 4796, Acer platanoides forest soil in Alavere, Estonia, 58.7562 ° N, 26.5109 ° E); KF 296788 (tundra soil in Prince Patrick Island, Canada, 76.23, – 119.3); and MK 536720 (soil crust in Victoria Land, Antarctica).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
3AE386D8257B5AF8A7DA8D20FE40BC6C.taxon	etymology	Etymology. Viljandi (Estonian) refers to the type locality, and globus (Latin) refers to the globe, reflecting the cosmopolitan distribution.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
325473CB7AED52A8A79A36AA5398D88B.taxon	type_taxon	Type species. Viljandia globalis Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
325473CB7AED52A8A79A36AA5398D88B.taxon	diagnosis	Diagnosis. Distinguishable from other species of Viljandiaceae based on diagnostic nucleotide signatures in SSU V 9 (positions 1695 – 1709 in S. cerevisiae ggcttccggcagcca; one mismatch allowed) and 5.8 S (positions 126 – 135 in the type species and 126 – 134 in S. cerevisiae cactctaagg; one mismatch allowed). Forms a monophyletic, least inclusive clade in Viljandiaceae, covering sequences EUK 1105441, EUK 1124341, and EUK 1124345 (Figs 1, 57).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
45A0007CDA43544ABFB49B0D76769315.taxon	type_taxon	Type genus. Viljandia Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
45A0007CDA43544ABFB49B0D76769315.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in SSU V 9 (positions 1695 – 1709 in S. cerevisiae gccagcaatggcagc; one mismatch allowed). Forms a monophyletic, least inclusive clade in Viljandiales, covering sequences EUK 1105441, EUK 1124341, EUK 1124345, and EUK 0524033 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
96478BBAFDF2500A94425145D82A76FC.taxon	type_taxon	Type family. Viljandiaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
96478BBAFDF2500A94425145D82A76FC.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 464 – 478 in type species and 490 – 504 in S. cerevisiae ctggccaacatcagt, one mismatch allowed). Forms a monophyletic, least inclusive clade in Viljandiomycetes, covering sequences EUK 1699905, EUK 1104555, EUK 1124343, EUK 1124346, EUK 1104962, EUK 1124344, EUK 1202387, EUK 1100361, EUK 1201679, EUK 1105441, EUK 1124341, and EUK 1124345 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DC6147AF12A751DAB9358A4435A43014.taxon	type_taxon	Type order. Viljandiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
DC6147AF12A751DAB9358A4435A43014.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in LSU D 2 (positions 464 – 478 in type species and 490 – 504 in S. cerevisiae ctggccaacatcagt, one mismatch allowed). Forms a monophyletic, least inclusive clade in Viljandiomycota, covering sequences EUK 1699905, EUK 1104555, EUK 1124343, EUK 1124346, EUK 1104962, EUK 1124344, EUK 1202387, EUK 1100361, EUK 1201679, EUK 1105441, EUK 1124341, and EUK 1124345 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
300045A0CCFA5CA59F9B6FBA33BF5541.taxon	type_taxon	Type class. Viljandiomycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
300045A0CCFA5CA59F9B6FBA33BF5541.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 2 (positions 464 – 478 in type species and 490 – 504 in S. cerevisiae ctggccaacatcagt; one mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1699905, EUK 1104555, EUK 1124343, EUK 1124346, EUK 1104962, EUK 1124344, EUK 1202387, EUK 1100361, EUK 1201679, EUK 1105441, EUK 1124341, and EUK 1124345 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED3F60193F2E59F3B8707B96AAA54785.taxon	diagnosis	Diagnosis. Separation from other species of Waitukubulimyces based on ITS 1 (positions 59 – 78 actgtgaaattgctctggta; one mismatch allowed) and LSU (positions 470 – 489 tttttgtttgatgagtagag; one mismatch allowed) as indicated in Fig. 60. Intraspecific variation up to 5.3 % in ITS 1. Interspecific distance> 20 % in ITS 1.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED3F60193F2E59F3B8707B96AAA54785.taxon	description	Description. Other sequences: MK 718926 and MK 718947 (both: barren soil in CO, USA); GlobalFungi records 3 c 686302 c 6 bfd 00 ff 4 db 5 b 414 d 28 c 645 (woodland soil in Marina, CA, USA, 36.6849, – 121.7780 ° E); 4 d 070 bd 97 b 6 d 68091 a 85749 c 15 e 6 c 744 (forest soil in Soria, Spain, 41.8694, – 2.87528 ° E); 61 c 43 b 4 d 22 e 1 a 0 efa 38 d 2 dba 3311970 e (cropland soil in Hangle, Uyghuria, China, 46.1886 ° N, 83.3294 ° E); 90 b 3386 fc 8 d 47 acc 87 e 18126 f 1 c 5 e 50 b (near-glacier soil in Arikaree, CO, USA); and abb 8924 cf 48 b 17 d 892 b 88816 e 96 f 0 ff 0 (grassland soil in Yahelong Gongma, Tibet, 38.21 ° N, 98.16 ° E).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
ED3F60193F2E59F3B8707B96AAA54785.taxon	etymology	Etymology.> Waitukubuli> (Igneri) refers to the country of Dominica, where the type material was collected, and Clifton refers to Clifton P. Bueno de Mesquita, who collected the first material of this species (MK 718926 and MK 718947; Bueno de Mesquita et al. 2020).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
AECA181964F2519BA850C302EA439ED4.taxon	type_taxon	Type species. Waitukubulimyces cliftonii Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
AECA181964F2519BA850C302EA439ED4.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU 5 ’ end (positions 52 – 66 in type species and S. cerevisiae tggaggaaaagaaaa, no mismatch allowed), LSU D 2 (positions 246 – 262 in type species and 240 – 256 in S. cerevisiae tgtgttcrctctgtgat; one mismatch allowed), and ITS 2 (positions 129 – 138 in type species tgggtcactt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Waitukubulimycetaceae, covering sequences EUK 1120710, EUK 1173015, EUK 1186290, EUK 1186291, and EUK 1186292 (Figs 1, 59).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4D06A5841D525AA292C715D77204B61B.taxon	type_taxon	Type genus. Waitukubulimyces Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
4D06A5841D525AA292C715D77204B61B.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU 5 ’ end (positions 52 – 66 in type species and S. cerevisiae tggaggaaaagaaaa, no mismatch allowed) and LSU D 2 (positions 246 – 262 in type species and 240 – 256 in S. cerevisiae tgtgttcrctctgtgat; one mismatch allowed) and ITS 2 (positions 129 – 138 in type species tgggtcactt; one mismatch allowed). Forms a monophyletic, least inclusive clade in Waitukubulimycetales, covering sequences EUK 1120710, EUK 1173015, EUK 1186290, EUK 1186291, and EUK 1186292 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
761C2DF9552959EBB171F2092D1BCEC5.taxon	type_taxon	Type family. Waitukubulimycetaceae Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
761C2DF9552959EBB171F2092D1BCEC5.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU 5 ’ end (positions 52 – 66 in type species and S. cerevisiae tggaggaaaagaaaa, no mismatch allowed) and LSU D 2 (positions 246 – 262 in type species and 240 – 256 in S. cerevisiae tgtgttcrctctgtgat; two mismatches allowed). Forms a monophyletic, least inclusive clade in Waitukubulimycetes, covering sequences EUK 1120710, EUK 1173015, EUK 1186290, EUK 1186291, and EUK 1186292.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C07D24BE95A351658B782A71BAFC7686.taxon	type_taxon	Type order. Viljandiales Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
C07D24BE95A351658B782A71BAFC7686.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5 ’ end (positions 52 – 66 in the type species and S. cerevisiae tggaggaaaagaaaa, no mismatch allowed). Forms a monophyletic, least inclusive clade in Waitukubulimycota, covering sequences EUK 1120710, EUK 1173015, EUK 1186290, EUK 1186291, and EUK 1186292 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95333BF59D23564A84851B74ED5E7593.taxon	type_taxon	Type phylum. Tartumycota Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
95333BF59D23564A84851B74ED5E7593.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on diagnostic nucleotide signatures in LSU D 3 (positions 1009 – 1023 in type species and 969 – 983 in S. cerevisiae ggaacttgtacagtt, no mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences OQ 702815, EUK 1186161, OQ 687331, EUK 1186165, EUK 1186157, EUK 1200073, EUK 1186162, EUK 1123648, EUK 1138300, OQ 702816, ON 754309, UDB 028835, and HQ 191300 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A5ABFD10E4C25B09BE9E7F45767127DB.taxon	type_taxon	Type class. Waitukubulimycetes Tedersoo.	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
A5ABFD10E4C25B09BE9E7F45767127DB.taxon	diagnosis	Diagnosis. Distinguishable from other fungi based on a diagnostic nucleotide signature in the LSU 5 ’ end (positions 52 – 66 in the type species and S. cerevisiae tggaggaaaagaaaa, no mismatch allowed). Forms a monophyletic, least inclusive clade in fungi, covering sequences EUK 1120710, EUK 1173015, EUK 1186290, EUK 1186291, and EUK 1186292 (Fig. 1).	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F7FBB75A57BB5A809C793A066ED67037.taxon	type_taxon	Type phylum. Zoopagomycota Gryganskyi, M. E. Smith, Spatafora & Stajich, Mycologia 108 (5): 1035 (2016) [816300].	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
F7FBB75A57BB5A809C793A066ED67037.taxon	description	Description. As in Tedersoo et al. (2018)	en	Tedersoo, Leho, Hosseyni Moghadam, Mahdieh S., Panksep, Kristel, Prins, Victoria, Anslan, Sten, Mikryukov, Vladimir, Bahram, Mohammad, Abarenkov, Kessy, Kõljalg, Urmas, Esmaeilzadeh-Salestani, Keyvan, Pawłowska, Julia, Wurzbacher, Christian, Ding, Yi, Alkahtani, Saad Hussin, Nilsson, R. Henrik (2025): Thirty novel fungal lineages: formal description based on environmental samples and DNA. MycoKeys 124: 1-121, DOI: 10.3897/mycokeys.124.161674
