taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322112/files/figure.png	https://doi.org/10.5281/zenodo.17322112	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322114/files/figure.png	https://doi.org/10.5281/zenodo.17322114	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322118/files/figure.png	https://doi.org/10.5281/zenodo.17322118	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322124/files/figure.png	https://doi.org/10.5281/zenodo.17322124	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322126/files/figure.png	https://doi.org/10.5281/zenodo.17322126	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322128/files/figure.png	https://doi.org/10.5281/zenodo.17322128	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322130/files/figure.png	https://doi.org/10.5281/zenodo.17322130	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322138/files/figure.png	https://doi.org/10.5281/zenodo.17322138	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322140/files/figure.png	https://doi.org/10.5281/zenodo.17322140	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF89FF82968585DB2EE754C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322147/files/figure.png	https://doi.org/10.5281/zenodo.17322147	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322112/files/figure.png	https://doi.org/10.5281/zenodo.17322112	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322114/files/figure.png	https://doi.org/10.5281/zenodo.17322114	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322118/files/figure.png	https://doi.org/10.5281/zenodo.17322118	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322126/files/figure.png	https://doi.org/10.5281/zenodo.17322126	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322130/files/figure.png	https://doi.org/10.5281/zenodo.17322130	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322138/files/figure.png	https://doi.org/10.5281/zenodo.17322138	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322140/files/figure.png	https://doi.org/10.5281/zenodo.17322140	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322147/files/figure.png	https://doi.org/10.5281/zenodo.17322147	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF88FF86968587B72F8D52BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322128/files/figure.png	https://doi.org/10.5281/zenodo.17322128	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322112/files/figure.png	https://doi.org/10.5281/zenodo.17322112	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322118/files/figure.png	https://doi.org/10.5281/zenodo.17322118	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322124/files/figure.png	https://doi.org/10.5281/zenodo.17322124	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322126/files/figure.png	https://doi.org/10.5281/zenodo.17322126	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322128/files/figure.png	https://doi.org/10.5281/zenodo.17322128	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322130/files/figure.png	https://doi.org/10.5281/zenodo.17322130	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322138/files/figure.png	https://doi.org/10.5281/zenodo.17322138	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322140/files/figure.png	https://doi.org/10.5281/zenodo.17322140	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF8DFF88968586DE2E2F54C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322147/files/figure.png	https://doi.org/10.5281/zenodo.17322147	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322112/files/figure.png	https://doi.org/10.5281/zenodo.17322112	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322114/files/figure.png	https://doi.org/10.5281/zenodo.17322114	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	FIGURE 2. Type species and diagnostic characters of Walteriella A. W. particularis (Pic, 1923), an authoritatively identified specimen by S. Kazantsev (housed in NHMB); B–C. pronotum; D–E. elytra; F–G. claws of forelegs. W. taiwana (Kasantsev, 1999): B, D, F; W. sanguinea brevemarginata (Wittmer, 1983): C, E, G. Scale bar = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322118/files/figure.png	https://doi.org/10.5281/zenodo.17322118	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322124/files/figure.png	https://doi.org/10.5281/zenodo.17322124	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	FIGURE 4. Diagnostic differences of W. brunnea (Wittmer, 1955) and W. sanguinea brevemarginata (Wittmer, 1983). A–D. head, A–B: W. brunnea, C–D: W. sanguinea brevemarginata, male: A, C, female: B, D; E–F. right antenna of male, E: W. brunnea, F: W. sanguinea brevemarginata.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322126/files/figure.png	https://doi.org/10.5281/zenodo.17322126	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322130/files/figure.png	https://doi.org/10.5281/zenodo.17322130	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322138/files/figure.png	https://doi.org/10.5281/zenodo.17322138	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322140/files/figure.png	https://doi.org/10.5281/zenodo.17322140	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF82FF8C968582622EFB51DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322147/files/figure.png	https://doi.org/10.5281/zenodo.17322147	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322112/files/figure.png	https://doi.org/10.5281/zenodo.17322112	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	FIGURE 1. The diversity of Walteriella cantharid beetles in Taiwan. A–B. W. taiwana (Kasantsev, 1999), A: male, B: female; C–D. W. brunnea (Wittmer, 1955), C: male, D: female; E. W. sanguinea brevemarginata (Wittmer, 1983), male; F. W. kasugana sp. nov., male.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322118/files/figure.png	https://doi.org/10.5281/zenodo.17322118	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	FIGURE 3. Habitus, dorsal view. A, D–E. W. taiwana (Kasantsev, 1999), A: holotype male, D: male, E; female; B, F–H. W. brunnea (Wittmer, 1955), B: holotype male, F–G: colour variations of males, H; female; C, I–K. W. sanguinea brevemarginata (Wittmer, 1983), C: holotype female, I: male, J–K: body size variations of females. L–M. W. kasugana sp. nov., L: holotype male, M: paratype female. Scale bars = 5.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322128/files/figure.png	https://doi.org/10.5281/zenodo.17322128	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	FIGURE 6. Male genitalia of W. kasugana sp. nov. A–D. Aedeagus, A: ventral view, B: lateral view, C: dorsal view, D: caudal view; E–F. comparison of laterophyses and basal part of median lobe between W. brunnea (Wittmer, 1955) and W. kasugana, E: W. brunnea, F: W. kasugana. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322130/files/figure.png	https://doi.org/10.5281/zenodo.17322130	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	FIGURE 7. Female abdominal ventrite VII. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322138/files/figure.png	https://doi.org/10.5281/zenodo.17322138	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	FIGURE 8. Female genitalia.A. W.taiwana (Kasantsev, 1999); B. W.brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. Scale bars: 1.0 mm. Abbreviations: ag—accessory gland; di—diverticulum; sd— spermathecal duct; sp—spermatheca; va—vagina.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322140/files/figure.png	https://doi.org/10.5281/zenodo.17322140	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 9. Distribution maps of four Walteriella species from Taiwan. A. W. taiwana (Kasantsev, 1999); B. W. brunnea (Wittmer, 1955); C. W. sanguinea brevemarginata (Wittmer, 1983); D. W. kasugana sp. nov. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322147/files/figure.png	https://doi.org/10.5281/zenodo.17322147	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	FIGURE 10. Seasonal activity of four Walteriella species from Taiwan. The number of observations is based on examined specimens, literature records and citizen science data.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF87FF8F9685843E2B8052C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322126/files/figure.png	https://doi.org/10.5281/zenodo.17322126	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	FIGURE 5. Male genitalia. A–D. W. taiwana (Kasantsev, 1999); E–H. W. brunnea (Wittmer, 1955); I–L. W. sanguinea brevemarginata (Wittmer, 1983). Aedeagus: A–K; inner sac: M–N; ventral view: A, E, I, M; lateral view: B, F, J, N; dorsal view: C, G, K; caudal view: D, H, L. Scale bars: 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
962B8793FF84FF8F968585022A6A508A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17322151/files/figure.png	https://doi.org/10.5281/zenodo.17322151	FIGURE. 11. Holotype of Leiothorax kashmirensis Wittmer, 1978 (housed in NHMB). A. Habitus, dorsal view; B–C. aedeagus, A: ventral view, B: lateral view; D. label data. Scale bars: (A) 5.0 mm; (B–C) 0.5 mm.	FIGURE. 11. Holotype of Leiothorax kashmirensis Wittmer, 1978 (housed in NHMB). A. Habitus, dorsal view; B–C. aedeagus, A: ventral view, B: lateral view; D. label data. Scale bars: (A) 5.0 mm; (B–C) 0.5 mm.	2025-09-11	Hsiao, Yun;Okushima, Yûichi		Zenodo	biologists	Hsiao, Yun;Okushima, Yûichi			
