taxonID	type	description	language	source
962B8793FF89FF82968585DB2EE754C7.taxon	description	Figs. 1 – 10	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF89FF82968585DB2EE754C7.taxon	materials_examined	Type species, by original designation: Lycocerus particularis Pic, 1923 (Fig. 2 A).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF89FF82968585DB2EE754C7.taxon	diagnosis	Diagnosis (after Kasantsev 1999). Walteriella can be distinguished from other cantharine genera by the combination of following characters: usually reddish to yellowish-brown elytra (Figs. 1, 2 A, D – E, 3), densely covered with pubescence; serrate (Figs. 1 A – B, 3 A, D – E) or filiform antennae, with antennomeres simple or slightly or strongly flattened (Figs. 1 C – F, 2 A, 3 B – C, F – M); subquadrate (Fig. 2 B) to trapezoidal (Fig. 2 C) pronotum, with anterior angles and margin rounded and somewhat expanded, lateral margins absent on posterior half in some species and disc distinctly convex; broadly expanded elytra (Figs. 1 A – B, 2 A, 3 A, D – E), with lateral sides rounded or slender, parallel-sided (Figs. 1 C – F, 3 B – C, F – M); strongly (Figs. 1 A – B, 2 A, 3 A, D – E) or weakly (Figs. 1 C – F, 3 B – C, F – M) developed elytral costae; legs with long bristling setae; simple claws or anterior claw of each leg with a small basal tooth; posterior margin of abdominal ventrite VII of female (Fig. 7) subrounded or subtruncate laterally, with a weakly sclerotised membrane behind the middle emargination; oval aedeagus, with the dorsal plate of each paramere distinctly curved inwards, forming a broad middle emargination (Figs. 5 C, G, K, 6 C) and an apical lock (Figs. 5 D, H, L, 6 D).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF88FF86968587B72F8D52BF.taxon	description	Figs. 1 A – B, 2 B, D, F, 3 A, D – E, 5 A – D, 7 A, 8 A, 9 A, 10	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF88FF86968587B72F8D52BF.taxon	materials_examined	Type material examined. Holotype (Fig. 3 A): ♂: “ (Near Chihtuan) / Taoyuan Hsien, / alt. 1200 m TAIWAN / 31 st March, 1980 / Coll. T. Shimomura // HOLOTYPUS / des. S. Kasantsev // Walteria / taiwana n. sp. / S. Kasantsev det. 1997 // Walteria / taiwana n. sp. / S. Kasantsev det. 1997 // CANTHARIDAE / CANTH 00002833 ” (NHMB). Additional material examined. CHIAYI COUNTY: 1 ♂, 18 Provincial Rd., 67.5 km, 26. IV. 2004, By Sweeping, J. - H. Chen & C. - Y. Huang leg. (NMNS: NMNS ENT 4809 - 40). HSINCHU COUNTY: 1 ♀, Mamei, Jianshih, 19. IV. 2017, S. - H. Chang leg. (YHC). KAOHSIUNG CITY: 1 ♀, Tengjhih, Tauyuan, 28. III. 2015, W. - C. Liao leg. (YHC). NANTOU COUNTY: 1 ♂, 1 ♀, Sungkang, 17. IV. 1995, C. Lou leg. (KURA); 1 ♂, Guandaoshan, 9. IV. 1996, R. Matsumoto leg. (KURA); 2 ♂, Fenghuangshan, 3. III. 2012, Y. - T. Wang leg. (YHC); 1 ♂, 23.5147279, 120.9095722, Shalixian Forest Road, Xinyi, 17. IV. 2025, By Sweeping, M. Geiser leg. (YHC).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF88FF86968587B72F8D52BF.taxon	diagnosis	Diagnosis. Adults of this species can be readily distinguished from its Taiwanese congeners by the following characters (states of W. brunnea, W. sanguinea brevemarginata and W. kasugana in parentheses): red pronotum, covered with reddish pubescence (Figs. 1 A – B, 3 A, D – E) (black to reddish brown, covered with pale pubescence; Figs. 1 C – F, 3 B – C, F – M); red elytra (Figs. 1 A – B, 3 A, D – E) (yellowish brown in W. kasugana and some specimens of W. brunnea; Figs. 1 F, 3 B, G, L – M); antennae serrate (Figs. 1 A – B, 3 A, D – E) (filiform; Figs. 1 C – F, 3 B – C, F – M); subquadrate pronotum, with lateral margins complete (Fig. 2 B) (usually trapezoidal, with anterior angles and margin rounded and somewhat expanded, lateral margins absent in posterior half; Fig. 2 C); strongly expanded elytra, with four distinctly developed costae (Figs. 1 A – B, 2 D, 3 A, D – E) (parallel-sided or only weakly widened apically, with two weakly developed costae; Figs. 1 C – F, 2 E, 3 B – C, F – M); anterior claw of each leg with a small basal tooth (Fig. 2 F) (simple; Fig. 3 G); aedeagus: dorsal plate of each paramere with two apical teeth from caudal view (Fig. 5 D) and one basal tooth on the inner surface (Fig. 5 C), laterophyses distinctly separated from each other (Fig. 5 C), with apex acute, not exceeding the middle emargination of dorsal plate (dorsal plate of each paramere with only one apical tooth from caudal view (Figs. 5 H, L, 6 D), without basal tooth on the inner surface, laterophyses closely next to each other (Fig. 5 G, K, 6 C), with apex rounded, largely exceeding the middle emargination of dorsal plate); female abdominal ventrite VII: terminal margin subrounded laterally, with a well-developed and distinctly sclerotised membrane behind the middle emargination (Fig. 7 A) (truncate or slightly sinuate laterally, with a small and weakly sclerotised membrane behind the middle emargination; Fig. 7 B – D); female genitalia: pyriform vagina, with surface smooth, and extremely short spermathecal duct (Fig. 8 A) (stout and rounded vagina, with surface wrinkled, and longer and well-developed spermathecal duct; Fig. 8 B – D). Supplementary description	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF88FF86968587B72F8D52BF.taxon	description	Male. Aedeagus (Fig. 5 A – D). Oval; globular and swollen dorsally at base. Ventral process of each paramere subcylindrical, moderately narrowed apically, bent inwards; dorsal plate of each paramere with two well-developed teeth from caudal view and a large basal tooth on inner surface, nearly as long as ventral process, distinctly curved inwards, rounded apically, forming a broad, oval middle emargination; laterophyses distinctly separated from each other, bent dorsad, apically acute, not exceeding the middle emargination of dorsal plate. Female. Abdominal ventrite VII (Fig. 7 A) with lateral margins weakly arcuate, terminal margin subrounded laterally and concave medially, with a rounded and distinctly sclerotised membrane behind the middle emargination. Female genitalia (Fig. 8 A) with vagina pyriform; diverticulum moderately spiralling, much longer than spermathecal duct; spermatheca with a moderately spiral and very long tube, slightly longer than diverticulum; accessory gland very long, about 4.0 times as long as diverticulum.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF88FF86968587B72F8D52BF.taxon	distribution	Distribution and phenology. This species is widely distributed in the mountainous regions of Taiwan’s main island (Fig. 9 A), with occurrence records ranging from early March to late May, peaking in middle April (Fig. 10). Based on the available data, this species is sympatric with three other Walteriella species in Taiwan.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF8DFF88968586DE2E2F54C0.taxon	description	Figs. 1 C – D, 3 B, F – H, 4 A – B, E, 5 E – H, 6 E, 7 B, 8 B, 9 B, 10	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF8DFF88968586DE2E2F54C0.taxon	materials_examined	Type material examined. Holotype (Fig. 3 B): ♂: “ Taiheizan / Formosa. / V – 6 – 32 // L. Gressitt / Collection // Holotypus // Podabrinus / brunneus / Wittm. / det. W. Wittmer // California Academy / of Sciences / Type No. 7784 ” (CAS). Additional material examined. KAOHSIUNG CITY: 1 ♂, 1 ♀, Daguihu, 11. V. 2023, C. - H. Ma & I. - L. Lee leg. (YHC). NEW TAIPEI CITY: 1 ♀, 25.020332, 121.940902, Wazhi, Gongliao, 2. III. 2019, HID Light Trap, B. - H. Ho & J. - J. Cherng leg. (YHC). NANTOU COUNTY: 1 ♀, Sungkang, 28. IV. 1996, C. Lou leg. (KURA); 1 ♂, Piluchi [= Pilu Creek, Renai], 7 – 15. V. 1997, Malaise Trap, M. - M. Yang leg. (NMNS: NMNS ENT 2708 - 305); 1 ♂, Meifeng, 7. V. – 11. VI. 2002, Malaise Trap (KCN), C. - S. Lin & W. - T. Yang leg. (NMNS: NMNS ENT 4912 - 828); 1 ♀, 23.6593442, 120.8053385, Fenghuang Forest Road, Lugu, 17. IV. 2025, By Hand, Y. - T. Chung leg. (YHC). TAICHUNG CITY: 1 ♀, Dasyueshan National Forest Recreation Area, 24 ° 15 ′ N, 121 ° 00 ′ E, 2100 – 2300 m, Heping, 4. VI. 2010, T. Tsuru leg. (KURA). TAOYUAN CITY: 1 ♂, Lalashan, 2. V. 1981, S. Tsuyuki leg. (KURA); 2 ♂, Lalashan, 14. V. 2009, C. - F. Lee leg. (TARI). PINGTUNG COUNTY: 1 ♀, Dahenshan [= Tahanshan Forest Road], 6. IV. 2013, W. - R. Liang leg. (NHMUK); 1 ♂, Mt. Dahanshan [= Tahanshan], ca. 1300 m, Chunri, 26. III. 2018, T. Niisato leg. (KURA); 1 ♂, same locality, 27. III. 2018, T. Niisato leg. (KURA); 1 ♀, 22.4071 N 120.7575 E, Jinshuiying Historic Trail, 0 – 2 km, alt. 1470 m, Chunri, 11. IV. 2025, By Sweeping, B. - H. Ho & M. Geiser leg. (BHHC). YILAN COUNTY: 1 ♂, 1 ♀, Dabaishan [= Tabaishan], 14. IV. 2015, L. Huang leg. (YHC).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF8DFF88968586DE2E2F54C0.taxon	diagnosis	Diagnosis. Adults of this species are externally extremely similar to W. sanguinea brevemarginata but distinguishable from it by the following characters (states of W. sanguinea brevemarginata in parentheses): semilustrous head and pronotum (Fig. 4 A – B) (head and pronotum shiny; Fig. 4 C – D); head with more elongate vertex and surface densely punctate and pubescent (Fig. 4 A – B) (shorter vertex and surface relatively sparsely punctate and pubescent; Fig. 4 C – D); males with relatively smaller and less protruding eyes, with ratio of eye diameter to interocular space 1.0: 2.8 – 3.7 (Fig. 4 A – B) (eyes large and strongly protruding, with ratio of eye diameter to interocular space 1.0: 2.2 – 2.5; Fig. 4 C – D); antennae usually elongate and slender (Fig. 4 E) (usually slightly flattened; Fig. 4 F); elytra subparallel-sided (Fig. 3 F – H) (usually weakly to moderately widened apically; Fig. 3 I – K); aedeagus: oval, dorsal plate of each paramere with rounded or obtuse apex, middle emargination oval (Fig. 5 E – H) (trapezoidal, dorsal plate of each paramere with acute apex, middle emargination with upper part distinctly widened; Fig. 5 I – L); female genitalia: stout vagina, with accessory gland about 2.0 times as long as diverticulum (Fig. 8 B) (rounded vagina, accessory gland about 3.0 times as long as diverticulum; Fig. 8 C). Supplementary description	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF8DFF88968586DE2E2F54C0.taxon	description	Male. Aedeagus (Fig. 5 E – H). Oval; globular and swollen dorsally at base. Ventral process of each paramere subcylindrical, slightly narrowed apically, slightly bent inwards; dorsal plate of each paramere with one well-developed apical tooth from caudal view, slightly longer than ventral process, distinctly curved inwards, rounded apically, forming a broad, oval middle emargination; laterophyses closely next to each other, distinctly bent dorsad, apically rounded, distinctly exceeding the middle emargination of dorsal plate. Female. Abdominal ventrite VII (Fig. 7 B) with lateral margins weakly arcuate, terminal margin slightly sinuate laterally, with submedian rounded lobe on each side, angularly emarginate medially, with an angular, apically rounded and weakly sclerotised membrane behind the middle emargination. Female genitalia (Fig. 8 B) with vagina elongate and stout apically; diverticulum weakly spiralling, much longer than spermathecal duct; spermatheca with a weakly spiral and long tube, longer than diverticulum; accessory gland long, about 2.0 times as long as diverticulum.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF8DFF88968586DE2E2F54C0.taxon	distribution	Distribution and natural history. This species occurs extensively in the mountainous areas of Taiwan’s main island (Fig. 9 B), with records spanning from late February to middle June, and most observations falling between middle April and early May (Fig. 10). It is found in sympatry with three other Walteriella species in Taiwan.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	description	Figs. 1 E, 2 C, E, G, 3 C, I – K, 4 C – D, F, 5 I – N, 7 C, 8 C, 9 C, 10	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	materials_examined	Type material examined. Holotype (Fig. 3 C): ♀: “ Taiwan 1977. 23. 4 / J. & S. Klapperich // Fenchihu / 1400 m // 144 // HOLOTYPUS // Athemellus / sanguineus / brevemarginatus Wittm. / det. W. Wittmer // Naturhistorisches / Museum Basel / Coll. W. Wittmer // CANTHARIDAE / CANTH 00002910 ” (NHMB). Additional material examined. CHIAYI COUNTY: 1 ♂, Arishan [= Alishan], 29. IV. 1917, T. Shiraki leg. (TARI). HUALIEN COUNTY: 1 ♀, Pilu, 25. V. 1999, J. Lai leg. (KURA); 1 ♀, Tsukeshan [= Chike Mountain], 9. III. 2016, Y. - M. Hsu leg. (YHC). NANTOU COUNTY: 1 ♂, Nanshanchi [= Nanshan Creek, Renai], 31. III. 1978, S. - C. Hung leg. (NTUE); 1 ♂, same locality, 19. III. 1983, Y. Kusakabe leg. (KURA); 1 ♀, same locality, 30. IV. 1983, M. Hasegawa leg. (KURA); 1 ♀, Sungkang, 24. V. 1988, C. - C. Lo leg. (NMNS: NMNS ENT 4463 - 1037); 1 ♀, Nanshanchi [= Nanshan Creek, Renai], 8. IV. 1991, Y. Okushima leg. (KURA); 1 ♀, Hohuanchi [= Hehuan Creek, Renai], 30. IV. 1994, T. Kishimoto & T. Shimizu leg. (KURA); 1 ♂, Sungkang, 28. IV. 1996, C. Lou leg. (KURA); 1 ♀, Lushan, 2. V. 1996, C. Lou leg. (KURA); 1 ♂, Piluchi [= Pilu Creek, Renai], 7 – 15. V. 1997, Malaise Trap, M. - M. Yang leg. (NMNS: NMNS ENT 2708 - 128); 1 ♂, Chunyang, 7. III. – 13. IV. 2006, Malaise Trap (KCN), C. - S. Lin & W. - T. Yang leg. (NMNS: NMNS ENT 7299 - 88); 1 ♀, Sungkang, 21. IV. 2014, P. - H. Huang (YHC). NEW TAIPEI CITY: 1 ♀, Wulai, 5. IV. 1981, C. - S. Chen leg. (NTUE); 1 ♂, 1 ♀, Fushan, 26. II. 1999, M. - S. Lin leg. (NTUE); 1 ♀, Wulai, 18. V. 1999, K. - L. Hsu leg. (NTUE); 2 ♀, Dadaoshan Forest Road, Wulai, 7. IV. 2025, M. Geiser & Y. Hsiao leg. (YHC). TAICHUNG CITY: 1 ♀, Anmashan Cottage, 8. VI. 2005, By Hand, Y. - L. Chen leg. (NMNS: NMNS ENT 5017 - 93). TAITUNG COUNTY: 2 ♀, Lichia-Lintao [= Lichia Forest Road], 26. III. 2016, C. Hsu, C. - F. Hsu & P. - C. Chou leg. (1 in NHMUK, 1 in YHC); 1 ♂, 1 ♀, Lijia-Rindao [= Lichia Forest Road], ca. 600 m, Beinan, 29. III. 2018, T. Niisato leg. (KURA). TAOYUAN CITY: 1 ♀, Piyaway, 7. IV. 1972, B. - S. Chang leg. (NMNS: 1282 - 26518); 1 ♀, Lalashan, 2. V. 1981, S. Tsuyuki leg. (KURA); 1 ♀, nr. Hsileng [= Sileng], 14. V. 1983, M. Hasegawa leg. (KURA); 1 ♀, Suling [= Sileng] — Chihtuan, 14. IV. 1984, Y. Kusakabe leg. (KURA); 1 ♂, Lalashan, 13 – 16. IV. 1984, Y. Kusakabe leg. (KURA); 1 ♂, Baling, 5. IV. 1991, Y. Okushima leg. (KURA); 1 ♂, Lalashan, 14. V. 2009, C. - F. Lee leg. (TARI); 1 ♂, Baehen [= North-Cross Island Highway], 9. IV. 2014, L. Huang leg. (YHC); 1 ♀, Shuewunao, Fushing, 26. IV. 2015, L. Huang leg. (YHC). YILAN COUNTY: 1 ♀, Chiduan [= Chihtuan], 12. IV. 1981, B. - S. Chang leg. (NMNS: 1282 - 26218); 1 ♂, Fushan, 30. III. 1995, Light Trap, Site: D, 2200, A. Warneke leg. (TFRI: 00084382); 1 ♂, 1 ♀, Fushan, 30. III. 1995, Light Trap, Site: A, 2200, J. - J. Hsiao leg. (TFRI: 00048588, 00084381); 1 ♀, Fushan, 30. III. 1995, Light Trap, Site: C, 2200, W. - T. Jou leg. (TFRI: 00084379); 1 ♂, Fushan, 31. III. 1995, Light Trap, Site: C, 0000, W. - T. Jou leg. (TFRI: 00084380); 1 ♂, Fushan, 21. III. 1996, Light Trap, Site: A, 0200, A. Warneke leg. (TFRI: 00114675); 1 ♂, 1 ♀, Fushan, 3. IV. 2003, Y. - B. Fan leg. (TFRI: 00084379); 1 ♂, 2 ♀, Shiaojiaoxi, Jiaoxi, 4. III. 2017, F. - S. Hu leg. (1 in NHMUK, 2 in YHC); 1 ♀, Dajiaoxi Plantation, Jiaoxi, 12. III. 2018, Flight Interception Trap S. - P. Lin leg. (YHC); 1 ♂, Yulan Industrial Road, 17. III. 2018, S. - P. Lin leg. (YHC).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	diagnosis	Diagnosis. This species is externally very similar to W. brunnea but differs from it as detailed in the diagnosis of W. brunnea above. Supplementary description	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	description	Male. Aedeagus (Fig. 5 I – L). Oval; globular and swollen dorsally at base. Ventral process of each paramere clavate, slightly expanded apically, slightly bent inwards; dorsal plate of each paramere with one well-developed apical tooth from caudal view, slightly longer than ventral process, distinctly curved inwards, acute apically, forming a broad, oval middle emargination; laterophyses closely next to each other, distinctly bent dorsad, apically rounded, distinctly exceeding the middle emargination of dorsal plate. Inner sac (Fig. 5 M – N) weakly swollen apically, slightly shorter than tegmen, surface with a rounded dorsal process near the apex of dorsal plate and a tooth-like process near the apex of ventral process. Female. Abdominal ventrite VII (Fig. 7 C) with lateral margins weakly arcuate, terminal margin obliquely truncate laterally, with submedian rounded lobe on each side, angularly emarginate medially, with an angular, apically rounded and weakly sclerotised membrane behind the middle emargination. Female genitalia (Fig. 8 C) with vagina elongate and rounded apically; diverticulum weakly spiralling, much longer than spermathecal duct; spermatheca with a weakly spiral and long tube, shorter than diverticulum; accessory gland very long, about 3.3 times as long as diverticulum.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	distribution	Distribution and natural history. This species is widely recorded from the mountainous areas of Taiwan’s main island (Fig. 9 C), with occurrences ranging from late February to early June and mostly concentrated in April to early May (Fig. 10). Despite its broad distribution, most records are from central to northern Taiwan. It is sympatric with W. brunnea and W. taiwana.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF82FF8C968582622EFB51DF.taxon	discussion	Remarks. Wittmer (1983) noted the similarity between this taxon and Walteriella sanguinea sanguinea (Wittmer, 1951) from Fukien, China, and treated it as a subspecies of W. sanguinea sanguinea. The only diagnostic difference proposed by Wittmer (1983) was the less protuberant submedian rounded lobes on the terminal margin of the female abdominal ventrite VII (Fig. 3 C). However, our morphological study reveals variation in this character, with some specimens exhibiting distinctly protuberant submedian rounded lobes (Fig. 7 C). Further comparative analyses of the aedeagus and female genitalia are urgently needed to clarify the taxonomic status of this species.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF87FF8F9685843E2B8052C3.taxon	description	Figs. 1 F, 3 L – M, 6 A – D, F, 7 D, 8 D, 9 D, 10	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF87FF8F9685843E2B8052C3.taxon	materials_examined	Type material. Holotype: ♂: “ 11. IV. 2025 / TAIWAN: Jinshuiying, 1555 m / 22 ° 24 ’ 17 ” N, 120 ° 45 ’ 31 ” E / Tahanshan, Chunri, Pingtung / M. Geiser, M. Fikáček / B. - H. Ho & G. - J. Phang leg. // HOLOTYPE / Walteriella kasugana / Hsiao & Okushima, 2025 ” (NMNS). Paratypes: 2 ♂: “ 27. IV. 2014 / Jinshueiying, Pingtung / Taiwan / W. - C. Liao leg. // PARATYPE / Walteriella kasugana / Hsiao & Okushima, 2025 ” (1 in NHMUK, 1 in YHC); 2 ♂: “ Taiwan: Pingtung / Tahanshan / 30. IV. 2014 leg. Y. - T. Chung // PARATYPE / Walteriella kasugana / Hsiao & Okushima, 2025 ” (TARI); 1 ♀: “ 2. V. 2020 / Tahanshan, Chunri, / Pingtung, Taiwan / F. - S. Hu & I. - L. Lee leg. ” (NMNS).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF87FF8F9685843E2B8052C3.taxon	diagnosis	Diagnosis. Adults of this species can be readily distinguished from other Taiwanese Walteriella species by their unique yellowish-brown elytra with black head, thorax and legs (Figs. 1 F, 3 L – M). This species is similar to W. brunnea in general shape of the aedeagus and female genitalia and some specimens of W. brunnea exhibit more or less yellowish coloration (Fig. 3 B, G). However, in addition to its characteristic elytral coloration, W. kasugana can be further distinguished from W. brunnea by the following characters (states of W. brunnea in parentheses): shorter elytra in males, about 3.4 – 3.6 times as long as wide (more elongate, about 3.8 – 4.0 times as long as wide); aedeagus: the base of median lobe smaller in lateral view (Fig. 6 B, F) (larger and more globular and swollen; Figs. 5 F, 6 E); female genitalia: accessory gland about 1.3 times as long as diverticulum (Fig. 8 D) (about 2.0 times as long as diverticulum; Fig. 8 B).	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF87FF8F9685843E2B8052C3.taxon	description	Description Male (Fig. 3 L). Length: 7.7 – 8.2 mm (holotype: 8.2); width: 1.5 – 1.7 mm (holotype: 1.7). Eyes black. Head black, lateral sides of clypeus pale. Antennae dark brown to black, with antennomere I brown basally. Mandibles brown, with apices darker; maxillary and labial palpomeres black, with apices of terminal maxillary palpomeres brown. Pronotum black, with posterior margin yellowish brown. Scutellum black. Elytra yellowish brown; mesoand metaventrites black; abdomen dark brown to black. Legs black, with coxae pale distally. Body densely covered with fine yellowish pubescence; anterior margin of clypeus fringed with yellowish bristles; antennae, elytra and legs with some yellowish bristles intermingled with primary pubescence, but sparsely at humeri. Head as long as wide, eyes moderately protruding, approximate ratio of eye diameter to interocular space 1.0: 2.7 – 4.3; dorsum slightly swollen posteriorly, depressed along anterior margin of clypeus and laterally anterior to eyes; anterior margin of clypeus arcuate, slightly emarginate medially; surface densely, finely punctate, each side with rectangular impression posterior to antennal socket; antennae filiform and long, extending to apical fourth of elytra, antennomere I clavate, II short and expanded apically, III – XI subcylindrical, with a small groove on each segment, approximate ratio of antennomere lengths as follows: 20.8: 10.0: 20.0: 27.5: 26.3: 27.5: 25.6: 25.6: 22.5: 21.3: 20.6. Pronotum trapezoidal, about 0.71 times as wide as head, about 1.18 times as long as wide; anterior and posterior margins arcuate; lateral sides widening anteriad, distinctly constricted in anterior third, marginated in anterior half; anterior angles somewhat expanded and rounded; posterior angles slightly protruding and obtuse; disc swollen, with pair of rounded elevations posterolaterally; antero-lateral areas hollowed; medio-longitudinal furrow distinct in centre of disc; surface semilustrous, finely and sparsely punctate. Scutellum triangular with obtuse apex. Elytra collectively about 1.5 times as wide as pronotum, about 3.4 – 3.6 times as long as wide, lateral margins subparallel, moderately narrowed apically; surface densely and coarsely punctate. Prosternal process concave apically. Mesoventrite distinctly convex medially. Legs very slender; femora nearly straight; tibiae nearly straight but somewhat bent; tarsi with tarsomere I – III emarginate and IV bilobed; claws simple. Aedeagus (Figs. 6 A – D, F). Oval; globular and swollen dorsally at base. Ventral process of each paramere subcylindrical, slightly narrowed apically, slightly bent inwards; dorsal plate of each paramere with one well-developed apical tooth from caudal view, slightly longer than ventral process, distinctly curved inwards, rounded apically, forming a broad, oval middle emargination; laterophyses closely next to each other, distinctly bent dorsad, apically rounded, distinctly exceeding the middle emargination of dorsal plate. Female (Fig. 3 M). Length: 9.7 mm; width: 1.9 mm. Similar to male. Body and pronotum wider than male; head as long as wide, eyes less protruding than in male, approximate ratio of eye diameter to interocular space 1.0: 4.1; antennae shorter than male, extending to elytral midlength, approximate ratio of antennomere lengths as follows: 15.0: 10.0: 17.0: 16.0: 15.0: 15.0: 16.0: 15.0: 14.0: 12.0: 12.0. Pronotum about 0.90 times as wide as head, about 1.05 times as long as wide. Elytra collectively about 1.4 times as wide as pronotum, about 3.7 times as long as wide. Abdominal ventrite VII (Fig. 7 D) with lateral margins weakly arcuate, terminal margin slightly sinuate laterally, with submedian rounded lobe on each side, angularly emarginate medially, with an oval, apically rounded and weakly sclerotised membrane behind the middle emargination. Female genitalia (Fig. 8 D) with vagina elongate and stout apically; diverticulum weakly spiralling, much longer than spermathecal duct; spermatheca with a weakly spiral and long tube, longer than diverticulum; accessory gland long, about 1.3 times as long as diverticulum. Derivation of name. The specific epithet is a Latinised adjective derived from ‘ Kasuga’, the Paiwan name for Chunri Township, referring to the only currently known distribution of the species. Chunri is an indigenous mountain township in Pingtung County, Taiwan, primarily inhabited by the Paiwan people.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF87FF8F9685843E2B8052C3.taxon	distribution	Distribution and natural history. This species is known only from Mt. Tahan, Chunri Township, Pingtung County, Taiwan (Fig. 9 D), with occurrence records spanning from middle April to early May and primarily concentrated in late April (Fig. 10). Although it appears to be sympatric with its closely related W. brunnea at Mt. Tahan, population of W. brunnea at Mt. Tahan is primarily recorded from late March to middle April, whereas the majority of specimens of W. kasugana were collected in late April to early May, with only a single record dated April 11.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F968585022A6A508A.taxon	materials_examined	Type species, by monotypy: Leiothorax kashmirensis Wittmer, 1978. (Fig. 11)	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F968585022A6A508A.taxon	discussion	Remarks. The generic name, Leiothorax Wittmer, 1978, is preoccupied by the Late Triassic true shrimp genus Leiothorax Pinna, 1974 (Malacostraca: Decapoda: Caridea). In order to correct the homonymy, Neoleiothorax nom. nov. (gender masculine) is proposed to replace Leiothorax Wittmer, 1978. The new generic name is derived from the Latin prefix Neo-, meaning “ new, ” combined with the root - leiothorax, retained from the original spelling, referring to the replacement of Leiothorax. The four species previously placed in Leiothorax are hereby transferred as follows.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F968583132AE557CB.taxon	distribution	Distribution. Kashmir.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F9685820A2AF956DC.taxon	distribution	Distribution. Nepal.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F968581002AF955F7.taxon	distribution	Distribution. Nepal.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
962B8793FF84FF8F968580162AF954C8.taxon	distribution	Distribution. Nepal.	en	Hsiao, Yun, Okushima, Yûichi (2025): A taxonomic review of Walteriella Kazantsev, 2001 (Coleoptera: Cantharidae) from Taiwan, with description of a new species and a nomenclatural act on Leiothorax Wittmer, 1978. Zootaxa 5692 (1): 57-76, DOI: 10.11646/zootaxa.5692.1.3, URL: https://doi.org/10.11646/zootaxa.5692.1.3
