taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461829	https://doi.org/10.3897/zookeys.1259.164450.figure5	Figure 5. Acicnemis albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 781). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 5. Acicnemis albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 781). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461857	https://doi.org/10.3897/zookeys.1259.164450.figure32	Figure 32. X-ray μCT generated 3 D models of two Acicnemis species. The colored parts in the segmented models below represent hindwing (in blue), metanotum + dorso-longitudinal flight muscles (in green), and dorso-ventral flight muscles (in orange). A. A. ryukyuana Lewis, 2023 (OKENT 0053498) dorsal view with scales removed, showing a relatively smooth pronotum and distantly spaced punctures; B. A. albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 782) dorsal view with scales removed, showing a moderately rugose to roughened pronotum with interspaces between punctures much smaller; C. A. ryukyuana segmented model with the right elytron removed, showing a comparively larger metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 30 %); D. A. albofasciata showing a comparatively smaller metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 23 %).	Figure 32. X-ray μCT generated 3 D models of two Acicnemis species. The colored parts in the segmented models below represent hindwing (in blue), metanotum + dorso-longitudinal flight muscles (in green), and dorso-ventral flight muscles (in orange). A. A. ryukyuana Lewis, 2023 (OKENT 0053498) dorsal view with scales removed, showing a relatively smooth pronotum and distantly spaced punctures; B. A. albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 782) dorsal view with scales removed, showing a moderately rugose to roughened pronotum with interspaces between punctures much smaller; C. A. ryukyuana segmented model with the right elytron removed, showing a comparively larger metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 30 %); D. A. albofasciata showing a comparatively smaller metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 23 %).	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461831	https://doi.org/10.3897/zookeys.1259.164450.figure7	Figure 7. Acicnemis azumai Morimoto & Miyakawa, 1995 (OKENT 0132427). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 7. Acicnemis azumai Morimoto & Miyakawa, 1995 (OKENT 0132427). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
BAB38C43B84D5AD8A739F31F12F84438.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461832	https://doi.org/10.3897/zookeys.1259.164450.figure8	Figure 8. Acicnemis biarcuata Hubenthal, 1919 (JHL _ AREV _ 628). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 8. Acicnemis biarcuata Hubenthal, 1919 (JHL _ AREV _ 628). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
A2A84396C69158FDA08B252BE9BD89DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461833	https://doi.org/10.3897/zookeys.1259.164450.figure9	Figure 9. Acicnemis bickhardti Hubenthal, 1919 (JHL _ AREV _ 591). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 9. Acicnemis bickhardti Hubenthal, 1919 (JHL _ AREV _ 591). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2C364F23F2EB55148197EDD143888B42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461835	https://doi.org/10.3897/zookeys.1259.164450.figure10	Figure 10. Acicnemis cordata Hubenthal, 1919 holotype (JHL _ AREV _ 987). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 10. Acicnemis cordata Hubenthal, 1919 holotype (JHL _ AREV _ 987). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F2EFD8DB464C5BB49665B1AE70E0D11E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461836	https://doi.org/10.3897/zookeys.1259.164450.figure11	Figure 11. Acicnemis costulifera Hubenthal, 1919 (JHL _ ACITAI _ 003). A. Lateral; B. Oblique. Scale bar: 1 mm (applies only to A). Note that the silver region is an entomological pin.	Figure 11. Acicnemis costulifera Hubenthal, 1919 (JHL _ ACITAI _ 003). A. Lateral; B. Oblique. Scale bar: 1 mm (applies only to A). Note that the silver region is an entomological pin.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
183AA775DE6B51CB82458DC267CF10C2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
183AA775DE6B51CB82458DC267CF10C2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
183AA775DE6B51CB82458DC267CF10C2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461837	https://doi.org/10.3897/zookeys.1259.164450.figure12	Figure 12. Acicnemis cryptica Lewis & Kojima, sp. nov. (JHL _ AREV _ 778). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 12. Acicnemis cryptica Lewis & Kojima, sp. nov. (JHL _ AREV _ 778). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
0B173776B8925A96AF787090D19F92E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
0B173776B8925A96AF787090D19F92E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
0B173776B8925A96AF787090D19F92E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461838	https://doi.org/10.3897/zookeys.1259.164450.figure13	Figure 13. Acicnemis dividicincta dividicincta Morimoto & Miyakawa, 1995 (JHL _ AREV _ 772). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 13. Acicnemis dividicincta dividicincta Morimoto & Miyakawa, 1995 (JHL _ AREV _ 772). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2923444FAAB05781AB1191A0E92145B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2923444FAAB05781AB1191A0E92145B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461839	https://doi.org/10.3897/zookeys.1259.164450.figure14	Figure 14. Acicnemis dividicincta okinawana Morimoto & Miyakawa, 1995 (JHL _ SYN _ 106). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 14. Acicnemis dividicincta okinawana Morimoto & Miyakawa, 1995 (JHL _ SYN _ 106). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
A9AD03D2B69C5CFDA239305F33D36AA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
A9AD03D2B69C5CFDA239305F33D36AA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
A9AD03D2B69C5CFDA239305F33D36AA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
A9AD03D2B69C5CFDA239305F33D36AA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461840	https://doi.org/10.3897/zookeys.1259.164450.figure15	Figure 15. Acicnemis exilis Morimoto & Miyakawa, 1995 (OKENT 0065625). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 15. Acicnemis exilis Morimoto & Miyakawa, 1995 (OKENT 0065625). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461828	https://doi.org/10.3897/zookeys.1259.164450.figure4	Figure 4. Sclerolepidia in Acicnemis. A. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing low, rounded sclerolepidia; B. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing protruding, erect sclerolepidia.	Figure 4. Sclerolepidia in Acicnemis. A. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing low, rounded sclerolepidia; B. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing protruding, erect sclerolepidia.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461841	https://doi.org/10.3897/zookeys.1259.164450.figure16	Figure 16. Acicnemis kiotoensis Nakane, 1963 (JHL _ AREV _ 775). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 16. Acicnemis kiotoensis Nakane, 1963 (JHL _ AREV _ 775). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
6FA0DC04D9645807AD2E8B6E193B52DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
6FA0DC04D9645807AD2E8B6E193B52DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
6FA0DC04D9645807AD2E8B6E193B52DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461842	https://doi.org/10.3897/zookeys.1259.164450.figure17	Figure 17. Acicnemis koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 17. Acicnemis koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
BBBD59AA596D5FC7A6ECA6D76421B4F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
BBBD59AA596D5FC7A6ECA6D76421B4F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
BBBD59AA596D5FC7A6ECA6D76421B4F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
BBBD59AA596D5FC7A6ECA6D76421B4F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461843	https://doi.org/10.3897/zookeys.1259.164450.figure18	Figure 18. Acicnemis laeta Hubenthal, 1919 (JHL _ AREV _ 773). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 18. Acicnemis laeta Hubenthal, 1919 (JHL _ AREV _ 773). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DDDC73717A1C5A5B942A07135233633F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DDDC73717A1C5A5B942A07135233633F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DDDC73717A1C5A5B942A07135233633F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DDDC73717A1C5A5B942A07135233633F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461844	https://doi.org/10.3897/zookeys.1259.164450.figure19	Figure 19. Acicnemis luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 409). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 19. Acicnemis luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 409). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461845	https://doi.org/10.3897/zookeys.1259.164450.figure20	Figure 20. Acicnemis maculaalba Roelofs, 1875 (JHL _ AREV _ 777). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 20. Acicnemis maculaalba Roelofs, 1875 (JHL _ AREV _ 777). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
3A98A0486003545BA3018AA6A9CF6822.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
3A98A0486003545BA3018AA6A9CF6822.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
3A98A0486003545BA3018AA6A9CF6822.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461846	https://doi.org/10.3897/zookeys.1259.164450.figure21	Figure 21. Acicnemis nobilis Hubenthal, 1919 (JHL _ AREV _ 496). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 21. Acicnemis nobilis Hubenthal, 1919 (JHL _ AREV _ 496). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
99B40C3B45FA56D283181925B0964925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
99B40C3B45FA56D283181925B0964925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
99B40C3B45FA56D283181925B0964925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
99B40C3B45FA56D283181925B0964925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461847	https://doi.org/10.3897/zookeys.1259.164450.figure22	Figure 22. Acicnemis nohirai Morimoto & Miyakawa, 1995 (JHL _ AREV _ 774). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 22. Acicnemis nohirai Morimoto & Miyakawa, 1995 (JHL _ AREV _ 774). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461828	https://doi.org/10.3897/zookeys.1259.164450.figure4	Figure 4. Sclerolepidia in Acicnemis. A. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing low, rounded sclerolepidia; B. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing protruding, erect sclerolepidia.	Figure 4. Sclerolepidia in Acicnemis. A. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing low, rounded sclerolepidia; B. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing protruding, erect sclerolepidia.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461848	https://doi.org/10.3897/zookeys.1259.164450.figure23	Figure 23. Acicnemis palliata Pascoe, 1872 (JHL _ AREV _ 779). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 23. Acicnemis palliata Pascoe, 1872 (JHL _ AREV _ 779). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
C54830051F165B59A4CFC62B2EA1A456.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
C54830051F165B59A4CFC62B2EA1A456.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
C54830051F165B59A4CFC62B2EA1A456.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
C54830051F165B59A4CFC62B2EA1A456.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461849	https://doi.org/10.3897/zookeys.1259.164450.figure24	Figure 24. Acicnemis postica Hubenthal, 1919 (JHL _ AREV _ 780). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 24. Acicnemis postica Hubenthal, 1919 (JHL _ AREV _ 780). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461850	https://doi.org/10.3897/zookeys.1259.164450.figure25	Figure 25. Acicnemis ryukyuana Lewis, 2023 (OKENT 0063788). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 25. Acicnemis ryukyuana Lewis, 2023 (OKENT 0063788). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461857	https://doi.org/10.3897/zookeys.1259.164450.figure32	Figure 32. X-ray μCT generated 3 D models of two Acicnemis species. The colored parts in the segmented models below represent hindwing (in blue), metanotum + dorso-longitudinal flight muscles (in green), and dorso-ventral flight muscles (in orange). A. A. ryukyuana Lewis, 2023 (OKENT 0053498) dorsal view with scales removed, showing a relatively smooth pronotum and distantly spaced punctures; B. A. albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 782) dorsal view with scales removed, showing a moderately rugose to roughened pronotum with interspaces between punctures much smaller; C. A. ryukyuana segmented model with the right elytron removed, showing a comparively larger metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 30 %); D. A. albofasciata showing a comparatively smaller metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 23 %).	Figure 32. X-ray μCT generated 3 D models of two Acicnemis species. The colored parts in the segmented models below represent hindwing (in blue), metanotum + dorso-longitudinal flight muscles (in green), and dorso-ventral flight muscles (in orange). A. A. ryukyuana Lewis, 2023 (OKENT 0053498) dorsal view with scales removed, showing a relatively smooth pronotum and distantly spaced punctures; B. A. albofasciata (Ter-Minasian, 1953) (JHL _ AREV _ 782) dorsal view with scales removed, showing a moderately rugose to roughened pronotum with interspaces between punctures much smaller; C. A. ryukyuana segmented model with the right elytron removed, showing a comparively larger metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 30 %); D. A. albofasciata showing a comparatively smaller metanotum and dorso-longitudinal flight muscles relative to the elytra (~ 23 %).	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
5D5F21B1AD665E6F806788A6CA815813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
5D5F21B1AD665E6F806788A6CA815813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
5D5F21B1AD665E6F806788A6CA815813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
5D5F21B1AD665E6F806788A6CA815813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461851	https://doi.org/10.3897/zookeys.1259.164450.figure26	Figure 26. Acicnemis sauteri Hubenthal, 1919 (JHL _ AREV _ 623). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 26. Acicnemis sauteri Hubenthal, 1919 (JHL _ AREV _ 623). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F6287459457D5125A8A12D3A16A2CFE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F6287459457D5125A8A12D3A16A2CFE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F6287459457D5125A8A12D3A16A2CFE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461852	https://doi.org/10.3897/zookeys.1259.164450.figure27	Figure 27. Acicnemis shibatai Voss, 1971 (JHL _ AREV _ 776). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 27. Acicnemis shibatai Voss, 1971 (JHL _ AREV _ 776). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
B87BD5220EA3561BA8C43A55BE18EDB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461827	https://doi.org/10.3897/zookeys.1259.164450.figure3	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	Figure 3. Scutella and pronota of various Acicnemis species. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing two bare (unscaled) tubercles baso-medially on the pronotum; B. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing a broad, unscaled scutellum; C. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing a bare, but more slender scutellum than A. kiotoensis; D. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing a small scutellum covered in scales; E. A. palliata Pascoe, 1872 (JHL _ AREV _ 779) showing a larger scutellum that is shiny and bare dorsally, and surrounded along the perimeter by yellow scales; F. A. shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052) showing a smaller scutellum covered in scales.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
B87BD5220EA3561BA8C43A55BE18EDB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
B87BD5220EA3561BA8C43A55BE18EDB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
B87BD5220EA3561BA8C43A55BE18EDB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461853	https://doi.org/10.3897/zookeys.1259.164450.figure28	Figure 28. Acicnemis shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 28. Acicnemis shigematsui Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 052). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DE46B977D1385A19A1AF1AE7EC20886C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461825	https://doi.org/10.3897/zookeys.1259.164450.figure1	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	Figure 1. Maximum likelihood tree of twelve Acicnemis species based on a 300 base-pair long CO 1 gene fragment constructed in IQ-TREE v. 1.6. 12 using a GTR + F + I + G 4 model. Branch support values represent standard nonparametric bootstraps (1000 replicates). Bootstrap values less than 75 are not displayed on the tree. The codes MEBLA 191-14, VVGPL 2845-15, and OWEVG 891-15 are BOLD Systems accession numbers, whereas all other codes are GenBank accession numbers. Associated Acicnemis weevil figures on the right of the tree are approximately to scale.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DE46B977D1385A19A1AF1AE7EC20886C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DE46B977D1385A19A1AF1AE7EC20886C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
DE46B977D1385A19A1AF1AE7EC20886C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461855	https://doi.org/10.3897/zookeys.1259.164450.figure30	Figure 30. Acicnemis squamata Lewis & Kojima, sp. nov. (JHL _ AREV _ 771). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 30. Acicnemis squamata Lewis & Kojima, sp. nov. (JHL _ AREV _ 771). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CDF7E3820987565CA9374527DCFB4285.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CDF7E3820987565CA9374527DCFB4285.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
CDF7E3820987565CA9374527DCFB4285.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461856	https://doi.org/10.3897/zookeys.1259.164450.figure31	Figure 31. Acicnemis suturalis Roelofs, 1875 (JHL _ AREV _ 421). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 31. Acicnemis suturalis Roelofs, 1875 (JHL _ AREV _ 421). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F14219CE481D553D900D14BA7FAE6024.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461830	https://doi.org/10.3897/zookeys.1259.164450.figure6	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	Figure 6. Aedeagi of Acicnemis. A 1–2. A. albofasciata (Ter-Minasian 1953); B 1–2. A. azumai Morimoto & Miyakawa, 1995; C 1–2), A. cryptica Lewis & Kojima, sp. nov.; D 1–2. A. dividicincta Morimoto & Miyakawa, 1995; E 1–2. A. exilis Morimoto & Miyakawa, 1995; F 1–2. A. kiotoensis Nakane, 1963; G 1–2. A. laeta Hubenthal, 1919; H 1–2. A. luteomaculata Morimoto & Miyakawa, 1995; I 1–2. A. maculaalba Roelofs, 1875; J 1–2. A. nobilis Hubenthal, 1919; K 1–2. A. nohirai Morimoto & Miyakawa, 1995; L 1–2. A. palliata Pascoe, 1872; M 1–2. A. postica Hubenthal, 1919; N 1–2. A. ryukyuana Lewis, 2023; O 1–2. A. sauteri Hubenthal, 1919; P 1–2. A. shibatai Voss, 1971; Q 1–2. A. shigematsui Morimoto & Miyakawa, 1995; R 1–2. Acicnemis sp. 1; S 1–2. A. squamata Lewis & Kojima, sp. nov.; T 1–2. A. suturalis Roelofs, 1875.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F14219CE481D553D900D14BA7FAE6024.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461826	https://doi.org/10.3897/zookeys.1259.164450.figure2	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	Figure 2. Diagnostic morphological characters in Acicnemis. A. A. azumai Morimoto & Miyakawa, 1995 (JHL _ ACITAI _ 081) showing lack of prominent posterior prosternal projections; B. A. laeta Hubenthal, 1919 (JHL _ AREV _ 533) showing prominent prosternal projections; C. A. kiotoensis Nakane, 1963 (JHL _ AREV _ 982) showing antenna with the second funicular antennomere longer than antennomeres 3 + 4; D. A. koguma Lewis & Kojima, sp. nov. (JHL _ SYN _ 141) showing antenna with the second funicular antennomere shorter than antennomeres 3 + 4; E. A. luteomaculata Morimoto & Miyakawa, 1995 (JHL _ AREV _ 189) showing truncate third tarsomere; F. A. ryukyuana Lewis, 2023 (JHL _ ACI _ 011) showing weakly emarginate third tarsomere; G. A. maculaalba Roelofs, 1875 (JHL _ AREV _ 577) showing strongly emarginate third tarsomere.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
F14219CE481D553D900D14BA7FAE6024.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461854	https://doi.org/10.3897/zookeys.1259.164450.figure29	Figure 29. Acicnemis sp. 1 (JHL _ AREV _ 766). A. Lateral; B. Dorsal. Scale bar: 1 mm.	Figure 29. Acicnemis sp. 1 (JHL _ AREV _ 766). A. Lateral; B. Dorsal. Scale bar: 1 mm.	2025-11-10	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.		Zenodo	biologists	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.			
