identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AF5DBBB6FD2951478C1E7F8AB32201D9.text	AF5DBBB6FD2951478C1E7F8AB32201D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis albofasciata (Ter-Minasian 1953)	<div><p>Acicnemis albofasciata (Ter-Minasian, 1953)</p><p>Figs 1, 5, 6 A 1–6, 2, 32 B, D</p><p>Trachodes albofasciata Ter-Minasian, 1953: 317.</p><p>Acicnemis nigra Nakane, 1963: 37.</p><p>Type material examined</p><p>(Russia: 8; Japan: 173; Total: 181). Lectotype (designated here). Russia • Primorsky Krai, Lyanchihe (26 km from Vladivostok), 19. V. 1949, on Betula manshurica, ZIN, JHL_AREV_997 . Paralectotype. Russia • Primorsky Krai, Partizansk (formerly Suchan), VIII. 1925 (1, ZIN), JHL _AREV_996 .</p><p>Non-type material.</p><p>Russia • Primorsky Krai, Benevskoye, 43.17°N, 133.76°E, 4. VII. 2014, E. Jendek (2, CNCI), CNCCOLVG 00007390 (BOLD accession number: VVGPL 2845 -15), CNCCOLVG 00008936 (dissected male, BOLD accession number: OWEVG 891-15); • S. Primor’ye, Kedrovaya Pad Reserve, 20. V. – 2. VI. 1992, T. Nakamura (3, KUM), JHL_AREV_499 (dissected male), JHL_AREV_782, JHL_AREV_787 ; • Saghalien, Kawakami (present day Sinegorsk), 7. VIII. 1922, T. Esaki (1, KUM), JHL_AREV_899 ; Japan: Akita Prefecture: • Tamagawa, Tazawako-machi, 10. VII. 1976, S. Miyakawa (12, TUA), JHL_AREV_674 (dissected male), JHL_AREV_811 – JHL_AREV_814, JHL_AREV_935 ; • Tamagawa Onsen, 10. VII. 1976, S. Miyakawa (4, TUA), JHL_AREV_932 (dissected male), JHL_AREV_934 ; • Senboku, Nyutou-onsen, 10. VI. 2015, K. Takahashi (1, KUM), JHL_AREV_887 ; • Takada-odake, 14. VII. 1962, Y. Miyatake (1 dissected male, KUM), JHL_AREV_898 ; • Sengan-toge, 14. IX. 1974, S. Miyakawa (2, TUA), JHL_AREV_965 (dissected male) ; Aomori Prefecture: • Towada, 12. VI. 1955, K. Shimoyama (1 dissected male, ZMH), ZMH 837198 ; • Nurukawa, 12. VIII. 1950, K. Shimoyama (1, ZMH), ZMH 837199 ; • Kobokutai, 5. VIII. 1937, K. Shimoyama (1, ZMH), ZMH 837200 ; • Yunomata, Ohata-machi, Shimokita Penn., 14–30. VII. 1956, K. Morimoto (7, KUM), JHL_AREV_543 (dissected male), JHL_AREV_793 – JHL_AREV_797, JHL_AREV_846 ; Fukushima Prefecture: • Tateiwa, 22–23. VI. 1984, A. Saito (2, CMIC), CBM–ZI 68470 (dissected male), CBM–ZI 68471 ; • Minami-aizu-machi, Yaso, 24. VI. 2020, T. Saeki (5, KUM), JHL_AREV_882 – JHL_AREV_886 ; • Shindenhara, 28. V. 1979, K. Emoto (1 dissected male, KUM), JHL_AREV_850 ; Gifu Prefecture: • Gifu (1, NHMB), JHL_SYN_192 ; • Hirayu, 29. VII. – 28. VIII. 1959, M. Sato (2, KUM), JHL_AREV_102, JHL_AREV_865 ; • Yunohana, 22. VI. 1956 (1 dissected male, CMNC), JHL_AREV_112 ; Gunma Prefecture: • Sugenuma, 10–11. VII. 1975, H. Irie (1, KUM), JHL_AREV_869 ; • Nikamatazawa, Mt. Shirane, 16–18. VII. 1975, H. Irie (1 dissected male, KUM), JHL_AREV_918 ; • Fujimi-toge, 4. VIII, M. Samejima (8, TUA), JHL_AREV_961 (dissected male), JHL_AREV_962, JHL_AREV_964 ; • Hokkaido: “ Hokkaido ”, “ 6.19. 53 ”, “ Arakawa ” (1, NHMB), JHL_SYN_190 ; • Sounkyo, 11. VII 1986, S. Nomura (3, KUM), JHL_SYN_189 ; • Aizankei, 29. VII. 1964 (1, KUM), JHL_AREV_103 ; • Aizankei, Mt. Daisetsu, 27–31. VII. 1955, K. Morimoto &amp; S. Kimoto (14, KUM), JHL_AREV_501 (dissected male), JHL_AREV_798, JHL_AREV_799, JHL_AREV_801 – JHL_AREV_810, JHL_AREV_855 ; • Teshio, 4. VII. 1916, T. Isshiki (1 dissected male, HUM), JHL_AREV_101 ; • Ashoromura ( Tokachi), 29. VII. – 23. VIII. 1949, R. Matsuda (5, KUM), JHL_AREV_781, JHL_AREV_888 – JHL_AREV_891 ; • Mitsumata-rindo, Kamishihoro-cho, 25. VI. 1986, S. Miyakawa &amp; K. Morimoto (1, TUA; • 2, KUM), JHL_AREV_542, JHL_AREV_815, JHL_AREV_816 ; • Ashorobuto, Ashoro-gun, 24–31. V. 1957, M. Takahashi (3, KUM), JHL_AREV_480 (dissected male), JHL_AREV_847, JHL_AREV_848 ; • Mt. Daisetsuzan, 26. VII. 1953, Y. Kurosawa (2, NMNS), JHL_AREV_852, JHL_AREV_853 ; • Daisetsuzan, 1. VIII. 1962 (3, KUM), JHL_ACITAI_008 (dissected male), JHL_AREV_104, JHL_AREV_105 ; • Jushichinosawa, Mt. Nipesotsu, 6. VIII. 1976, H. Irie (1, KUM), JHL_AREV_856 ; • Nakayama-toge (near Sapporo), 22. VI. 1986, K. Morimoto (1, KUM), JHL_AREV_857 ; • Bibai, 12. V. 1962, K. Kamijo, collected from Populus sp. (1, KUM), JHL_AREV_858 ; • Oyukiyama, 31. VII. 1952, Yoshida (1, KUM), JHL_AREV_859 ; • Aizankei, Antaroma, 31. VII. 1952, T. Shirozu (1, KUM), JHL_AREV_861 ; • Lake Akan, Kushiro, 8. VI. 1957, M. Takahashi (1, KUM), JHL_AREV_862 ; • Wassakanai, Sohya-shichou, 26. VII. 1991, S. Ohmomo (1, KUM), JHL_AREV_863 ; • Nishiashoro ( Tokachi), 17. VIII. 1949, R. Matsuda (1, KUM), JHL_AREV_864 ; • Yukomanbetsu, Mt. Daisetsu, 25. VII. 1955, K. Morimoto (1, KUM), JHL_AREV_866 ; • Horoka, Kamishihoro-cho, 17. VII. 1976, H. Irie (1, KUM), JHL_AREV_867 ; • Kamiotoineppu ( Teshio), 24. VIII. 1922, T. Esaki (4, KUM), JHL_AREV_868, JHL_AREV_877 – JHL_AREV_879 ; • Kawakami (near Honbetsu), 17–27. VII. 1953, Y. Hirashima (1, KUM), JHL_AREV_870 ; • Aizankei, 19. VII. 1962, Y. Miyatake (1, KUM), JHL_AREV_871 ; • Aizankei ( Ishikari), 29. VII. 1952, T. Shirozu (5, KUM), JHL_AREV_872 – JHL_AREV_876 ; • Nakarikubetsu, Rikubetsu Town, Asyoro County, 24. VII. 2017, K. Narita (2, KUM), JHL_AREV_880, JHL_AREV_881 ; • Meakan-dake, Akan National Park, 5. VII. 1958, S. Miyamoto (4, KUM), JHL_AREV_892 – JHL_AREV_895 ; • Uenae, Tomakomai City, 15. VIII. 1989, H. Kojima (6, TUA), JHL_AREV_922 (dissected male), JHL_AREV_910 – JHL_AREV_914 ; • Mt. Karibayama, Shimamaki, 24–27. VII. 1989, H. Kojima (2, TUA), JHL_AREV_924, JHL_AREV_926 ; • Mt. Daisengen, Hakodate City, 23. VII. 1989, H. Kojima (1, TUA), JHL_AREV_925 ; • Poroshirisanso, Hidaka, 5. VIII. 1988, H. Kojima (2, TUA), JHL_AREV_927, JHL_AREV_928 ; • Kamishihoro-chou, Mitsumata, 17. VII. 2014, S. Yamamoto, JHL_DNA_154 (GenBank accession number: PV 255624) (1, KUM) ; Ibaraki Prefecture: • Mt. Tsukuba, 22. VIII. 1977, A. Tanaka (1, HUM), JHL_AREV_854 ; Ishikawa Prefecture: • Mt. Hakusan, 18. V. 1979, I. Togashi &amp; D. R. Smith (1 dissected male, USNM), JHL_AREV_985 ; Iwate Prefecture: • Iwate, VI. 1936, H. Yamamoto (2, NHMB), JHL_SYN_188, JHL_SYN_191 ; Miyagi Prefecture: • Nuruyu, Hanayama mura, 2–6. VIII. 1984, K. Morimoto (1 dissected male, KUM), JHL_AREV_849 ; • Mt. Funagata, Taiwa-machi, 21. VIII. 2009, J. Aoki (2, KUM), JHL_AREV_839 (dissected male) ; Nagano Prefecture: • Nakabusa, Shinano, 24. VII, T. Samejima (1 dissected male, TUA), JHL_AREV_673 ; • Bandoko, Shinano Province, 13. VII. 1935, H. Kiyosawa (4, KUM), JHL_AREV_917 (dissected male), JHL_AREV_919 – JHL_AREV_921 ; • Shirahone, 19. VII. 1956, S. Kimoto (2, KUM), JHL_AREV_593 (dissected male), JHL_AREV_916 ; • Shirahone, 12. VIII. 1960, Y. Kimura (1 dissected male, KUM), JHL_AREV_838 ; • Kamikouchi, 14. VII. 1929, M. Samejima (4, TUA), JHL_AREV_783 (dissected male), JHL_AREV_967 – JHL_AREV_969 ; Niigata Prefecture: • Sasagamine, 23. VIII. 1962, H. Koike (1 dissected male, KUM), JHL_AREV_896 ; Tochigi Prefecture: • Yumoto, Oku-nikko, 7. VII. 1950, M. Takahashi (3, KUM), JHL_AREV_902 (dissected male), JHL_AREV_906, JHL_AREV_907 ; • Yumoto, 30. VII. 1960, K. Suga (1 dissected male, KUM), JHL_AREV_903 ; • Chu-zenji, 16. VII. 1916, E. Gallois (3, HUM), JHL_AREV_851 ; • Tokura, Nikko, 16. VI, S. Miyakawa (3, TUA), JHL_AREV_960 (dissected male) ; Yamanashi Prefecture: • Daibosatsu, 22–23. VI. 1985, K. Yoshihara (6, TUA), JHL_AREV_629 (dissected male), JHL_AREV_788, JHL_AREV_789, JHL_AREV_790 – JHL_AREV_792 ; • Hikawa-rindo, Mts. Daibosatsu, 22. VII. 1983, S. Miyakawa (21, TUA), JHL_AREV_946 (dissected male), JHL_AREV_948 – JHL_AREV_958 ; • Kanayama, 1. VII. 1989, T. Tanaka (1 dissected male, TUA), JHL_AREV_923 ; • Kanayamadaira, 28–30. VI. 1991, H. Urushihara (2, TUA), JHL_AREV_930 (dissected male), JHL_AREV_933 ; • Koshu-shi, Enzankamiigiwara, 13. VIII. 2016, S. Shimamoto (2, TUA), JHL_AREV_937 (dissected male), JHL_AREV_940 ; • Koshu-shi, Enzankamiigiwara, 3. V. 2015, S. Shimamoto (1, TUA), JHL_AREV_939 ; Unknown Prefecture: • Ichi-no-hashi, 29. VII. 1975, S. Ogata (1 dissected male, KUM), JHL_AREV_502 .</p><p>Diagnosis.</p><p>Body length 5.0– 6.8 mm. Covered in black and dark gray scales, with longitudinal, black-bordered white stripes across the lateral edges of the pronotum and elytral humeri (compare with A. cryptica which has the entire lateral portion of the pronotum covered in white to yellow scales). Apical 1 / 2 of elytra with a V-shaped patch of white scales. Pronotum evenly punctured across dorsal surface, but often with shiny, impunctate longitudinal midline. Scutellum bare over anterior 1 / 2 of dorsal surface, otherwise covered in pale yellowish scales. Only odd elytral intervals with erect scales (even elytral intervals may possess erect scales at the extreme elytral apices). Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, and converging evenly at apex into a dull, rounded point. Internal sac visible in anterior 1 / 2 of aedeagus and tuberculate (roughed appearance); • internal sac also with hook-like protruding structure anteriorly (at the apex) and surrounded by tuberculate, roughened musculature (Fig. 6 A 1 – 2).</p><p>Distribution.</p><p>This species occurs in far Eastern Russia (Primorsky Krai, Saghalien) and north-central mainland Japan (from Hokkaido south to Gifu Prefecture).</p><p>Remarks.</p><p>The two A. albofasciata syntypes deposited in the ZIN Collection are apparently both females, despite the original paper describing a male (B. Korotyaev to JHL, pers. comm. 2025). Dissection of males is required to conclusively differentiate between A. albofasciata and its sister species A. cryptica . However, based on our extensive collection examinations, we found that Acicnemis albofasciata occurs in the Far East (Primorsky Krai, Russia), Hokkaido, and northern mainland Japan, while A. cryptica occurs only in mainland Japan, from Aomori Prefecture south to Kyushu (Oita Prefecture). Furthermore, examined specimens of A. albofasciata have a black-scaled, lateral stripe on the pronotum, while A. cryptica have the lateral portion of the pronotum mostly covered in yellow to white scales. We are confident that the Ter-Minasian syntypes belong to A. albofasciata (not A. cryptica) as only this species apparently occurs outside of Japan (with known records from the syntype locality (Primorsky Krai, Russia), and as the Ter-Minasian syntypes have a black-scaled lateral pronotum band. To fix the identity of A. albofasciata, we hereby designate one of the Ter-Minasian syntypes (JHL _AREV_997) as a lectotype.</p></div>	https://treatment.plazi.org/id/AF5DBBB6FD2951478C1E7F8AB32201D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
8BBE2064DEF15BE5BFE56BF7AC8123C5.text	8BBE2064DEF15BE5BFE56BF7AC8123C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis azumai Morimoto & Miyakawa 1995	<div><p>Acicnemis azumai Morimoto &amp; Miyakawa, 1995</p><p>Figs 1, 2 A, 3 A, 6 B 1–6, 2, 7</p><p>Type material examined</p><p>(Japan: 82; South Korea: 3; Total: 84). Holotype. Japan: Okinawa Prefecture: • Kudeken, Chinen-son, 22. IV. 1975, H. Irie, bears red label reading “ (HOLOTYPE) Acicnemis azumai Morimoto et. Miyakawa, 1994 ”, JHL_AREV_113, ELKU, ELKU 2962 . Paratypes. Japan: Okinawa Prefecture: • Sueyoshi, “ 1988-6 - 5 ”, T. Ueno (5, KUM), JHL_ACITAI_041 (dissected male), JHL_AREV_134 – JHL_AREV_137 ; • Yona, 9–13. VIII. 1969, H. Makihara (4, KUM), JHL_AREV_108 – JHL_AREV_111 ; • Shuri, 14. VI. 1958, O. Nakachi (1, KUM), JHL_AREV_138 ; • Shuri, 16–20. V. 1978, H. Makihara (1, KUM), JHL_AREV_117 ; • Aguni Island, 24–25. VIII. 1989, Teruhisa Ueno, JHL_ACITAI_081 (dissected male) (4, KUM), JHL_AREV_124 – JHL_AREV_126 ; • Mabuni, 11. VI. 1977, H. Irie (2, KUM) JHL_AREV_127, JHL_AREV_128 ; • Nago, 18. VI. 1984, Teiso Esaki (1, KUM), JHL_AREV_130 ; • Akajima Island, 29–30. VIII. 1980, Teruhisa Ueno (1, KUM), JHL_AREV_129 ; • Mount Banna, Ishigaki Island, 30. VII. 1970, I. Matoba (1, KUM), JHL_AREV_131 ; • Tonaki Island, 27. IX. 1989, Teruhisa Ueno (2, KUM), JHL_AREV_132, JHL_AREV_133 ; • 18. VI. 1970, H. Makihara (2, KUM), JHL_AREV_122, JHL_AREV_123; • Tamagusuku, 4. V. 1962, M. Kina (1, KUM), JHL_AREV_151 ; • Tamagusuku, 23. VIII. 1962, M. Kina (2, KUM), JHL_AREV_152, JHL_AREV_153 ; • Otomi, Iriomote Island, 25. IV. 1969, H. Makihara (1, KUM), JHL_AREV_143 ; • Shimozato, Miyako Island, 1. IX. 1958, T. Hidaka (1, KUM), JHL_AREV_144 ; • Kudeken, 31. V. 1976, T. Ogasawara (1, KUM), JHL_AREV_145 ; • Shuri, 5–9. V. 1969, H. Makihara (1, KUM), JHL_AREV_146 ; • Naha, 13. VIII. 1952, K. Hamamatsu (1, KUM), JHL_AREV_147 ; • Sueyoshi, 9. VII. 1987, Teruhisa Ueno (1, KUM), JHL_AREV_148 .</p><p>Non-type material.</p><p>Japan: Okinawa Prefecture: • Iriomote Island, Mt. Tedo, 28. III. 2022, Y. Obae (1, KUM), JHL_AREV_511 ; • Ishigaki Island, Inoda, 24. III. 2022, S. Imada, M. Maruyama, T. Nozaki (1, KUM), JHL_AREV_512 ; • Kabira Park, Ishigaki Island, 6. VI. 1968, M. Takagi (1, KUM), JHL_AREV_763 ; • Okinawa Island, Sakukawa, 8. XI. 1971, T. Terya (1, RUMC), JHL_AREV_513 ; • Okinawa Island, Nishihara, 16. VII. 1977, M. Arasaki (1, RUMC), JHL_AREV_514 ; • Okinawa Island, Yona, 24. IV. 1965, T. Kakinohana (2, RUMC), JHL_AREV_515, JHL_AREV_516 ; • Okinawa Island, Kudeken, 28. VI. 1977, K. Omato (1, RUMC), JHL_AREV_517 ; • Okinawa Island, Shuri, 7. VII. 1972, M. Kinjo (1, RUMC), JHL_AREV_518 ; • Miyako Island, 1. IX. 1958, T. Takara (3, RUMC), JHL_AREV_519 – JHL_AREV_521 ; • Okinawa Island, Shuri, 16. VI. 1961, O. Nakachi (1, RUMC), JHL_AREV_522 ; • Okinawa Island, Shuri, 24. VIII. 1962, O. Nakachi (1, RUMC), JHL_AREV_523 ; • Okinawa Island (1, OIST), JHL_AREV_984 (GenBank accession number: PV 255625) ; • Tancha, Okinawa Island, 4. III. 2025, J. H. Lewis, to residential lights (1, OIST), JHL_AREV_995 ; • Tancha, Okinawa Island, 14. III. 2025, J. H. Lewis, to residential lights (1, OIST), JHL_AREV_993 ; • Tancha, Okinawa Island, 29. VIII. 2025, J. H. Lewis, to residential lights (1, OIST), JHL_AREV_2081 ; • Sueyoshi Park, Naha, 12–26. II. 2016, (1, OIST), OKENT 0132427 ; South Korea • Oseag Ri, Ganweondo, 4–6. VII. 1984, T. Senoh (1, KUM), JHL_AREV_155 ; • Sam Jeong Li, Ma Cheong Meon, Hamyang Gun, Gyeongsamnamdo, 9–15. V. 1991, K. Morimoto (2, KUM), JHL_AREV_154, JHL_ACITAI_079 (dissected male) .</p><p>Diagnosis.</p><p>Body length 3.3–4.9 mm. Covered in pale brown scales, with alternating white and black scaled bands on odd elytral intervals. Pronotum with two exposed, bare tubercles medially at the base (one on each side) (Fig. 3 A). Scutellum covered in yellow scales. Only odd elytral intervals with erect scales. Sclerolepidia distinct and protruding. Posterior edge of prosternum lacking prominent projections contiguous with the fore-coxae (Fig. 2 A) (present in A. laeta). Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a point. Internal sac without tuberculate or roughened musculature inside the aedeagus, but with a non-hooked protruding structure (unmodified in posterior 1 / 2, with roughened in anterior 1 / 2) that is equal in length to the pedon (Fig. 6 B 1 – 2).</p><p>Distribution.</p><p>This species occurs in southern Japan (Okinawa Prefecture, Ryukyu Islands) and South Korea.</p><p>Remarks.</p><p>On Okinawa Island (Japan) this species is commonly encountered in the spring and is often attracted to residential building lights.</p></div>	https://treatment.plazi.org/id/8BBE2064DEF15BE5BFE56BF7AC8123C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
BAB38C43B84D5AD8A739F31F12F84438.text	BAB38C43B84D5AD8A739F31F12F84438.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis biarcuata Hubenthal 1919	<div><p>Acicnemis biarcuata Hubenthal, 1919</p><p>Fig. 8</p><p>Type material examined</p><p>(Taiwan: 5). Holotype. Taiwan • “ Kosempo ”, “ Formosa ”, “ IX. 09 ”, “ Sauter ”, bears red label reading “ Holotypus ”, SDEI, SDEI Coleoptera # 303414, SDEI Muncheburg COL–18569 .</p><p>Non-type material.</p><p>Taiwan • Lienhoachii, “ 1975.5. 12 ”, S. Imasaka (1, TUA), JHL_SYN_184 ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.85867&amp;materialsCitation.latitude=22.139364" title="Search Plazi for locations around (long 120.85867/lat 22.139364)">Mudan Township</a>, Pingtung County, 22°08'21.71"N, 120°51'31.23"E, 9. III. 2013, Y. Fujisawa (1, TUA), JHL_AREV_626 ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.792694&amp;materialsCitation.latitude=22.103493" title="Search Plazi for locations around (long 120.792694/lat 22.103493)">Mudan Township</a>, Pingtung County, 22°06'12.57"N, 120°47'33.71"E, 6. III. 2013, Y. Fujisawa (1, TUA), JHL_AREV_627 ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.85361&amp;materialsCitation.latitude=22.141666" title="Search Plazi for locations around (long 120.85361/lat 22.141666)">Manzhou Township</a>, Pingtung County, 22°08'30"N, 120°51'13"E, 2. V. 2019, S. Shimamoto (1, TUA), JHL_AREV_628 .</p><p>Diagnosis.</p><p>Body length 4.0– 5.2 mm. Covered in dark-gray scales, with two V-shaped, transverse black bands across middle of elytra. Pronotum with pale brown and yellow scales dorsally, but dark at base medially. Pronotum approx. sculptured and carinate. Scutellum shiny and bare in examined specimens. Only odd elytral intervals with erect scales across elytral disk (some erect scales on 2 nd interval in apical fourth of elytra). Third and fifth elytral intervals with cluster of 3–5 erect scales at elytral base. Sclerolepidia distinct and protruding. Mesosternum with distinctly raised ridge between meso-coxae, highest laterally but dipping shallowly medially (no such ridge or only weakly developed in most other species). Third tarsomeres weakly emarginate.</p><p>Distribution.</p><p>This species is currently only known from Taiwan.</p></div>	https://treatment.plazi.org/id/BAB38C43B84D5AD8A739F31F12F84438	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
A2A84396C69158FDA08B252BE9BD89DC.text	A2A84396C69158FDA08B252BE9BD89DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis bickhardti Hubenthal 1919	<div><p>Acicnemis bickhardti Hubenthal, 1919</p><p>Fig. 9</p><p>Type material examined</p><p>(Taiwan: 3). Holotype. Taiwan • Kosempo, H. Sauter, 1909, bears a red label reading “ Type ”, SNSD, JHL_ACITAI_001 .</p><p>Non-type material.</p><p>Taiwan • “ 16. VII. 20 ”, T. Kano (1, NMNS), JHL_AREV_591; • 20. VI. 1938, unknown collector (1, TUA), JHL_AREV_592 .</p><p>Diagnosis.</p><p>Body length 5.0–7.0 mm. Easily distinguished from other East Asian Acicnemis by the (1) large size (2) black to dark red cuticle largely naked, but with bright yellow scaled bands running longitudinally across the lateral edges of the pronotum and elytral humeri, as well as transversally across the apical 1 / 2 of the elytra.</p><p>Distribution.</p><p>This species is known only from Taiwan.</p><p>Remarks.</p><p>Many similarly patterned, large species that presumably form a distinct clade within Acicnemis are present throughout Southeast Asia (e. g., A. ibis Faust, 1896 and A. peduncularis Pascoe, 1872), and a number of these remain undescribed.</p></div>	https://treatment.plazi.org/id/A2A84396C69158FDA08B252BE9BD89DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
2C364F23F2EB55148197EDD143888B42.text	2C364F23F2EB55148197EDD143888B42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis cordata Hubenthal 1919	<div><p>Acicnemis cordata Hubenthal, 1919</p><p>Fig. 10</p><p>Acicnemis fukienensis Voss, 1958 .</p><p>Type material examined</p><p>(Vietnam: 1). Holotype. Vietnam • “ Mau-son, Tongking ” (modern day northern Vietnam), “ Gehr. W. Muller, Vermacht, 1909 ”, bears a red label reading “ Typus ”, SNSD, JHL_AREV_987 .</p><p>Diagnosis.</p><p>Body length 5.0 mm. Virtually indistinguishable externally from A. maculaalba (see that species). Although most A. maculaalba have the white elytral spots interrupted by brown scales along the first elytral interval, some have no interruption (as in the A. cordata holotype); therefore, this character cannot be used alone to reliably separate A. maculaalba and A. cordata .</p><p>Distribution.</p><p>This species is known from the type locality (Mau-son, Tongking, which corresponds to modern day northern Vietnam) and apparently also Kuatun, China (Voss’s specimen).</p><p>Remarks.</p><p>We examined the holotype specimen of A. cordata Hubenthal, 1919 (collected from Fujian, China; SNSD) and found it is virtually indistinguishable externally from Taiwanese and Japanese specimens of A. maculaalba . However, as the A. cordata holotype is a female, it was not possible to examine the male genitalia, which are extremely important for distinguishing members of the A. maculaalba species complex. We tentatively recognize A. cordata here, but also note that further examination of specimens from mainland Asia is necessary to work out the species in this complex.</p></div>	https://treatment.plazi.org/id/2C364F23F2EB55148197EDD143888B42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
F2EFD8DB464C5BB49665B1AE70E0D11E.text	F2EFD8DB464C5BB49665B1AE70E0D11E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis costulifera Hubenthal 1919	<div><p>Acicnemis costulifera Hubenthal, 1919</p><p>Fig. 11</p><p>Type material examined</p><p>(Unknown country: 1). Holotype. Unknown locality: “ Coll. J. Faust Ankauf 1900 ”, bears a red label reading “ Typus ”, SNSD, JHL_ACITAI_003 .</p><p>Diagnosis.</p><p>Body length 4.8 mm. This is the only species covered here with costate elytra (on the 2 nd, 4 th, and 6 th intervals). Also, is has erect, elongate scales along the odd-numbered elytral intervals (as in A. ryukyuana).</p><p>Distribution.</p><p>Acicnemis costulifera is recorded from Taiwan and Japan in Alonso-Zarazaga et al. (2023); however, Hubenthal (1919 b) states that the holotype collection locality is unknown. We examined the holotype specimen and indeed, it does lack labels with any locality data. Furthermore, no specimens of this species collected from Japan were encountered in any of the examined collections. As such, we are confident that the record of this species from Japan is erroneous, and that the record from Taiwan should be treated with suspicion until additional specimens are found.</p></div>	https://treatment.plazi.org/id/F2EFD8DB464C5BB49665B1AE70E0D11E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
183AA775DE6B51CB82458DC267CF10C2.text	183AA775DE6B51CB82458DC267CF10C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis cryptica Lewis & Kojima 2025	<div><p>Acicnemis cryptica Lewis &amp; Kojima sp. nov.</p><p>Figs 6 C 1–6, 2, 12</p><p>Type material</p><p>(Japan: 72). Holotype. Japan: Nagano Prefecture: • Otari Onsen, Otari-mura, 23. VII. 1990, M. Hasegawa, bears a red holotype label reading “ HOLOTYPE / Acicnemis cryptica / Lewis &amp; Kojima, 2025 ”, JHL_AREV_672, dissected male, deposited in KUM . Paratypes. Japan: Aomori Prefecture: • Towada, 14. VI. 1957, K. Shimoyama (1 dissected male, KUM), JHL_AREV_498 ; Fukushima Prefecture: • Yunohana, Tateiwa V., Minami-Aizu, 14. VI. 1947, “ Y. K. ” (1 dissected male, NMNS), JHL_AREV_139 ; Fukuoka Prefecture: • Hikosan (Buzen), 19. V. 1939, H. Hori (1 dissected male, KUM), JHL_AREV_981 ; Gunma Prefecture: • Mt. Buson, 13–15. VII. 1975, H. Irie (2, KUM), JHL_AREV_904 (dissected male), JHL_AREV_905 ; Kochi Prefecture: • Mt. Tebako, 7–10. VIII. 1957, K. Morimoto (13, KUM), JHL_AREV_479 (dissected male), JHL_AREV_823 – JHL_AREV_827, JHL_AREV_829 – JHL_AREV_831, JHL_AREV_833 – JHL_AREV_835, JHL_AREV_837 ; • Mt. Tebako, 18–21. VII. 1955, K. Kojima (2, KUM), JHL_AREV_828, JHL_AREV_832 ; • Monobe-mura, 26–28. VIII. 1959, K. Morimoto (2, KUM), JHL_AREV_836, JHL_AREV_845 ; Ibaraki Prefecture: • Mt. Yamizo-san, Daigo-machi, 1. XII. 1995, S. Nomura (1 dissected male, KUM), JHL_AREV_901 ; Iwate Prefecture: • Mt. Hayachine, 2–5. VIII. 1982, H. Makihara (8, KUM), JHL_AREV_594 (dissected male), JHL_AREV_817, JHL_AREV_818, JHL_AREV_822, JHL_AREV_841 – JHL_AREV_844 ; Nagano Prefecture: • Otari Onsen, Otari-mura, 23. VII. 1990, M. Hasegawa (1, TUA), JHL_AREV_819 ; Oita Prefecture: • Mt. Sobo, 29. VII. 1953, T. Matsuda (1 dissected male, KUM), JHL_AREV_544 ; Shizuoka Prefecture: • Amagi-toge, Hacchoike Izu-hanto, 14. V. 1980, J. Okuma (4, TUA), JHL_AREV_947 (dissected male), JHL_AREV_959 ; • Tokyo Metropolis: Nippara, 19. IX, 1971, H. Takizawa, JHL_AREV_118 (1, HUM) ; • Ogawadani, Okutama, “ 1966.8. 6 ”, T. Goh (1 dissected male, HUM), JHL_AREV_140 ; • Mt. Takao, 27. V. 1917, E. Gallois (3, HUM) JHL_AREV_114 (dissected male), JHL_AREV_141, JHL_AREV_142 ; • Mt. Takao, 10. V. 1914, E. Gallois (1, HUM), JHL_AREV_115 ; • Mt. Takao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.2525&amp;materialsCitation.latitude=35.62972" title="Search Plazi for locations around (long 139.2525/lat 35.62972)">Hachioji</a>, 22. VIII. 2024, 35.62972°N, 139.2525°E, R. Anderson, beating in mixed maple / beech / cypress forest, “ 2024-102 ” (2, CMNC), JHL_AREV_2035, JHL_AREV_2026 ; • Takao Musashi, V. 1912, H. Takabayashi (1, HUM), JHL_AREV_116 ; Yamanashi Prefecture: • Daibosatsu, 19. V. 1974, M. Tao (1, TUA), JHL_AREV_778 ; • Koganezawa-rindo, Ohtsuki-shi, 30–31. V. 1980, J. Okuma (5, TUA), JHL_AREV_630 (dissected male), JHL_AREV_820, JHL_AREV_821 ; • Koganezawa-rindo, 23. VI. 1974, S. Miyakawa (8, TUA), JHL_AREV_941 – JHL_AREV_945 .</p><p>Diagnosis.</p><p>Body length 4.9–7.0 mm. Covered in dark to pale gray and black scales. Pronotum covered in black scales medially, but with white to yellow scales laterally (compare with A. albofasciata which also has white lateral pronotal bands, but bordered by black scales at the lateral-most extreme of the pronotum). Apical 1 / 2 of elytra with a V-shaped patch of white scales. Pronotum evenly punctured across dorsal surface, but often with shiny, impunctate longitudinal midline. Scutellum bare over anterior 1 / 2 of dorsal surface, otherwise covered in pale yellowish scales. Only odd elytral intervals with erect scales (even elytral intervals may possess erect scales at the extreme elytral apex). Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but sinuating into an acute point (Fig. 6 C 1 – 2). Internal sac without tuberculate or roughened musculature inside the aedeagus (compare with A. albofasciata, which has a tuberculate internal sac visible inside the aedeagus), but with hook-shaped protruding structure surrounded by tuberculate, roughened musculature.</p><p>Distribution.</p><p>This species is endemic to mainland Japan (excluding Hokkaido), north from Aomori Prefecture and south to Kyushu (Oita Prefecture, Fukuoka Prefecture).</p><p>Etymology.</p><p>The specific name cryptica is a reference to the remarkable similarity of this species in external morphology to A. albofasciata and also explains why it was overlooked for decades. We also suggest the Japanese common name ハイイロカレキゾウムシ [Haiiro-kareki-zoumushi], which translates in English to “ Gray-dead-wood-weevil ”.</p><p>Remarks.</p><p>This cryptic species was previously assumed to be a pale-color morph of A. albofasciata (see Morimoto and Miyakawa 1995: 50, fig. 11), and indeed only differs externally from A. albofasciata in overall color (pale gray in A. cryptica; black to dark gray in A. albofasciata) and weakly in scale pattern (entire lateral portion of pronotum white to yellow scaled in A. cryptica; isolated white to yellow band, bordered by black lateral pronotal band in A. albofasciata). The male genitalia of A. cryptica (which were apparently not examined previously), however, are completely distinct from A. albofasciata and differ in the shape of the aedeagus as well as in internal sac morphology (Fig. 6 C 1 – 2).</p></div>	https://treatment.plazi.org/id/183AA775DE6B51CB82458DC267CF10C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
0B173776B8925A96AF787090D19F92E6.text	0B173776B8925A96AF787090D19F92E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis dividicincta subsp. dividicincta Morimoto & Miyakawa 1995	<div><p>Acicnemis dividicincta dividicincta Morimoto &amp; Miyakawa, 1995</p><p>Figs 6 D 1–6, 2, 13</p><p>Acicnemis yakushimana Morimoto &amp; Miyakawa, 1995, syn. nov.</p><p>Type material examined</p><p>(Japan: 34). Holotype. Japan: Okayama Prefecture: • Mt. Gagyuzan, Takahashi City, 19–20. V. 1975, H. Irie, bears red label reading “ (HOLOTYPE) Acicnemis dividicincta Morimoto et. Miyakawa, 1994 ”, JHL_SYN_120, ELKU, ELKU 2957 . Paratypes. Fukuoka Prefecture: • Mt. Hiko, 5. IX. 1988, K. Morimoto (1, KUM), JHL_SYN_118 ; Kochi Prefecture: • Mt. Tebako, 7–10. VIII. 1957, K. Morimoto (2, KUM), JHL_SYN_114, JHL_SYN_115 ; Nagasaki Prefecture: • Unzen, 18. VII. 1946, M. Moritsu &amp; Y. Kurosawa (3, KUM), JHL_SYN_102 (dissected male), JHL_SYN_113, JHL_SYN_119 ; Okayama Prefecture: • Mt. Gagyuzan, Takahashi City, 19–20. V. 1975, H. Irie (3, KUM), JHL_SYN_111, JHL_SYN_112, JHL_DNA_142 (dissected male) ; Wakayama Prefecture: • Mt. Ootoh, 7. VIII. 1980, I. Matoba (1, KUM), JHL_SYN_116 .</p><p>Other type material.</p><p>Kagoshima Prefecture: • Miyanoura, Yakushima Island, 26. VII. 1974, T. Mikage, bears red label reading “ (HOLOTYPE) Acicnemis yakushimana Morimoto et. Miyakawa, 1994 ”, JHL_SYN_180 (1, ELKU), A. yakushimana holotype specimen, ELKU 2959 ; • Miyanoura, Yakushima Island, 26–29. VII. 1974, T. Mikage (13, KUM), JHL _SYN_122 – JHL _SYN_124, JHL _SYN_127 – JHL _SYN_130, JHL _SYN_132, JHL _SYN_133, JHL _SYN_138, JHL _SYN_139, JHL _DNA_141 (dissected male) ; • Shiratani, Yakushima Island, 24–26. VII. 1974, T. Mikage (2, KUM), JHL _SYN_121, JHL _SYN_131 ; • Mt. Tachudake, Yakushima Island, 20. VII. 1971, H. Irie (4, KUM), JHL _SYN_125, JHL _SYN_126, JHL _SYN_134, JHL _SYN_135 ; • Ambo – Kosugidani, Yakushima Island, 29. VII. 1929, Hiroshi Hori (2, KUM), JHL _SYN_136, JHL _SYN_137 ; • Shiratani-rindo, Yakushima Island, 20. VII. 1976, M. Kaneda (1, KUM), JHL _AREV_663 .</p><p>Non-type material.</p><p>Ehime Prefecture: • Omogo, Iyu, 19. VIII. 1969, T. Nakane, (1, HUM), JHL _AREV_772 ; • Tokyo Metropolis: Hachijo-jima, Mitsune, VI. 2001, T. Kinoshita (1 dissected male, KPMNH), JHL _AREV_936 ; • Hachijo-jima, Noboryou-toge, 9. VI. 1996, I. Hirai (1, KUM), JHL _AREV_938 .</p><p>Diagnosis.</p><p>Body length 4.2–7.0 mm. Covered in black and dark gray scales, with two V-shaped, transverse white and black bands across middle of elytra. Pronotum with weakly patterned gray, black and faint yellow scales. Scutellum covered loosely in yellow scales (denuded in many specimens). Only odd elytral intervals with erect scales across elytral disk (some erect scales on 2 nd interval in apical fourth of elytra). Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a moderately acute point. Internal sac not visible inside aedeagus, but with a non-hooked protruding structure (unmodified in posterior 1 / 2, with roughened in anterior 1 / 2) that is clearly longer than pedon (Fig. 6 D 1 – 2).</p><p>Distribution.</p><p>This subspecies occurs in mainland Japan (central Japan south to Yakushima Island) and adjunctly on Hachijo Island (Tokyo Metropolis).</p><p>Remarks.</p><p>Acicnemis yakushimana was distinguished from A. dividicincta by Morimoto and Miyakawa (1995) on the basis of purported differences in the relative proportions of the funicular antennomeres and minor differences in color. Furthermore, A. yakushimana also reportedly possessed a sclerite in the internal sac at the gonopore. After dissecting several specimens of each species across their geographic range, we did not observe any differences in genital morphology between A. yakushimana and A. dividicincta . Furthermore, no observable differences in antennal funicle proportions were observed. As such, we hereby treat A. yakushimana syn. nov. as a junior subjective synonym of A. dividicincta .</p></div>	https://treatment.plazi.org/id/0B173776B8925A96AF787090D19F92E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
2923444FAAB05781AB1191A0E92145B5.text	2923444FAAB05781AB1191A0E92145B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis dividicincta subsp. okinawana Morimoto & Miyakawa 1995	<div><p>Acicnemis dividicincta okinawana Morimoto &amp; Miyakawa, 1995</p><p>Figs 1, 14</p><p>Type material examined</p><p>(Japan: 13). Holotype. Japan: Okinawa Prefecture: • Hiji, Kunigami-son, Okinawa Island, 8. VII. 1974, T. Mikage, bears red label reading “ (HOLOTYPE) Acicnemis dividicincta okinawana Morimoto et. Miyakawa, 1995 ”, JHL_SYN_110, ELKU, ELKU 2958 . Paratypes. Japan: Kagoshima Prefecture: • Torigamine, Amami Island, 30. VI. 1976, N. Morishimo (1, KUM), JHL_SYN_104 ; Okinawa Prefecture: • Hiji, Kunigami-son, Okinawa Island, 3–9. VII. 1974, T. Mikage (5, KUM), JHL_SYN_101 (dissected male), JHL_SYN_103, JHL_SYN_105 – JHL_SYN_107 .</p><p>Non-type material.</p><p>Japan: Kagoshima Prefecture: • Mt. Yuwandake, Amami-Oshima, 5–9. VII. 1968, K. Sakai (1, CMIC), CBM-ZI 68475 ; Okinawa Prefecture: • Tancha, 1–7. VI. 2022, J. H. Lewis (1, OIST), JHL_DNA_126 (GenBank accession number: PV 255626) ; • Tancha, Onna-son, Okinawa Island, 2. VI. 2022, sweeping coastal vegetation, J. H. Lewis (1, OIST), JHL_SYN_109 ; • Yona, Okinawa Island, 19. X. 1973, S. Azuma (1, RUMC), JHL_SYN_108 ; • Tancha, Onna-son, Okinawa Island, 29. VIII. 2025, J. H. Lewis, to residential lights (1, OIST), JHL_AREV_2082 ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.27191&amp;materialsCitation.latitude=26.83604" title="Search Plazi for locations around (long 128.27191/lat 26.83604)">Oku</a>, Okinawa Island, 26.83604°N, 128.27191°E, 5–19. VIII. 2016 (1, OIST), OKENT 0089525 (GenBank accession number: PV 255627) .</p><p>Diagnosis.</p><p>Body length 4.8–6.5 mm. Identical to A. dividicincta dividicincta above, differing from that species only in possessing somewhat paler gray scales.</p><p>Distribution.</p><p>This subspecies occurs on Amami-Oshima (Kagoshima Prefecture) and Okinawa Island (Okinawa Prefecture).</p><p>Remarks.</p><p>We continue to recognize this subspecies as it is significantly isolated from the mainland populations and therefore presumably does represent a distinct intraspecific lineage.</p></div>	https://treatment.plazi.org/id/2923444FAAB05781AB1191A0E92145B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
A9AD03D2B69C5CFDA239305F33D36AA1.text	A9AD03D2B69C5CFDA239305F33D36AA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis exilis Morimoto & Miyakawa 1995	<div><p>Acicnemis exilis Morimoto &amp; Miyakawa, 1995</p><p>Figs 1, 6E 1–6, 2, 15</p><p>Type material examined</p><p>(Japan: 21; Taiwan: 3; Total: 24). Holotype. Japan: Kagoshima Prefecture: • Amami-Oshima, Naze, 27. V. 1978, T. Tsutsumi, bears red label reading “ (HOLOTYPE) Acicnemis exilis Morimoto et. Miyakawa, 1994 ”, ELKU, ELKU 2963 .</p><p>Non-type material.</p><p>Japan: Kagoshima Prefecture: • Jinushi-daimyojin, Satomura, Nakano-jima Island, 7–9. VII. 2019, S. Imada (2, KUM), JHL_AREV_595, JHL_AREV_596 ; Okinawa Prefecture: • Hikawa, Yonaguni Island, 3. V. 2016, N. Tsuji (1, KUM), JHL_AREV_597 ; • Banna-koen, Ishigaki Island, 1. IX. 2015, J. Aoki (9, KUM), JHL_SYN_145 (dissected male), JHL_AREV_598, JHL_AREV_599, JHL_AREV_601 – JHL_AREV_605; • JHL_AREV_724 ; • Mt. Omoto-dake, 15–17. IV. 2014, S. Yamamoto (2, KUM), JHL_AREV_606, JHL_AREV_607 ; • Southeast Botanical Gardens, Okinawa Island, 10–24. II. 2016 (1 dissected male, OIST), OKENT 0109018 ; • Chatan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.75918&amp;materialsCitation.latitude=26.32468" title="Search Plazi for locations around (long 127.75918/lat 26.32468)">Ireibaru</a>, 26.32468°N, 127.75918°E, 11–25. XI. 2015 (1, OIST), OKENT 0105900 (GenBank accession number: PV 255629) ; • Nanjo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.75918&amp;materialsCitation.latitude=26.32468" title="Search Plazi for locations around (long 127.75918/lat 26.32468)">Tamagusuku</a>, 26.32468°N, 127.75918°E, 9–23. IX. 2015, OKENT 0056923 (GenBank accession number: PV 255628) (1, OIST) ; • Sefa-utaki, Nanjo, 15–29. VII. 2016, (1, OIST), OKENT 0065625 ; near Hiji, Okinawa Island, 16. III. 2024, J. H. Lewis (1, OIST), JHL_AREV_998 ; • Banna-dake, Ishigaki Island, 20. VII. 1973, H. Takizawa (1, HUM), JHL_ACITAI_086 ; Taiwan • Wulai (near Taipei), 27. V. 1965, K. Morimoto (1 dissected male, KUM), JHL_AREV_624 ; • Kotosho Island, 13–15. VIII. 1968, H. Makihara (2, KUM), JHL_AREV_757, JHL_AREV_758 .</p><p>Diagnosis.</p><p>Body length 2.1–3.0 mm. Covered in pale brown, white, and gray scales, with two white-scaled spots in the apical 1 / 2 of the elytra. Scutellum covered in yellow scales. Only odd elytral intervals with erect scales. Sclerolepidia small but protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a rounded point. Internal sac without tuberculate or roughened musculature inside the aedeagus, but with a non-hooked protruding structure (unmodified in posterior 2 / 3, sclerotized in anterior 1 / 3) that reaches the tip of the temo (Fig. 6 E 1 – 2).</p><p>Distribution.</p><p>This small species occurs throughout the Ryukyu Islands (southern Kagoshima Prefecture and Okinawa Prefecture, Japan) as well as Taiwan.</p><p>Remarks.</p><p>This species appears to be closely allied with the similarly minute, long hind-legged Australian species Acicnemis meriones Pascoe, 1872, A. arachnopus Hubenthal, 1919, and the Sumatran species A. ambigua Hubenthal, 1919; however, phylogenetic analysis is required to confirm this. Preliminary observations suggest that some of the names in this group are synonyms.</p></div>	https://treatment.plazi.org/id/A9AD03D2B69C5CFDA239305F33D36AA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
D31A94491053581F892CFEBD2F63C355.text	D31A94491053581F892CFEBD2F63C355.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis Fairmaire 1849	<div><p>Acicnemis Fairmaire, 1849</p><p>Oplocnemus Dejean, 1835 (Type species: Oplocnemus mucronatus Buquet nomen nudum).</p><p>Hoplocnemus Agassiz, 1846 (unjustified emendation of Oplocnemus).</p><p>Acienemis Motschulsky, 1863 (incorrect subsequent spelling).</p><p>Berethia Pascoe, 1872 (Type species: Berethia medinotata Pascoe, 1872).</p><p>Acicnemus Haly, 1890 (incorrect subsequent spelling).</p><p>Type species.</p><p>Acicnemis variegata Fairmaire, 1849 (by subsequent designation).</p><p>Gender.</p><p>Feminine.</p><p>Description.</p><p>Body length 2.0–12.0 mm. Scale pattern: Usually cryptically patterned with contrasting brown, black, gray, and dull yellow scales. Pronotum and elytra often with erect scales. Head: Rostrum evenly curved in most species, and variably longer in females than in males; eyes large, ovate, and non-contiguous; antennae with seven-articled funicle. Prothorax: Densely punctate; pronotum often with tufts of erect scales; prosternum lacking well defined prosternal canal. Elytra: Scutellum varied (large, plate-like, naked scutellum in some species, or small and covered in scales in others); erect scales often only on odd-numbered elytral intervals; elytra unmodified in most species, costate in a few; elytra with apical projections in some species ( A. laticollis Pascoe, 1885); low tubercles along elytral intervals in some species, absent in most; volant, hind wings functional. Thorax: Sclerolepidia varied: prominent and protruding in some species, forming low, indistinct tubercles in others. Abdomen: First and second sternites variably concave medially in males (convex or flat in females). Legs: Femora all bearing a large ventral tooth (simple in most species, serrated in a few); forelegs elongate in some species (e. g., A. longimana Hubenthal, 1922); third tarsomeres emarginate (bilobed) in most species, truncate in a few; tarsal claws simple.</p><p>Distribution.</p><p>Acicnemis species are broadly distributed across the Palearctic Region (Azerbaijan, China, Georgia, Japan, Korean archipelago, Russia, Taiwan), the Oriental Region (Cambodia, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Thailand, Vietnam), the Australasian Region (Australia, Fiji, New Guinea, Samoa, Tahiti, Tonga), and apparently also the Afrotropical Region (Nigeria) (Hubenthal 1919 a, b; Zimmerman 1967; Alonso-Zarazaga and Lyal 1999; Kojima and Morimoto 2004; Oziegbe and Faluyi 2010; Pullen et al. 2014; Alonso-Zarazaga et al. 2023).</p><p>Ecology and natural history.</p><p>The larvae of Trachodini mine through dead wood (Lyal 2014). Host plant associations are not well known in Acicnemis; however, a few records exist ( Acicnemis palliata Pascoe, 1872 with Wisteria floribunda (Willd.) DC. and W. sinensis (Sims) DC. ( Fabaceae) (Morimoto and Miyakawa 1995; Lee and Morimoto 1996; Zhang 2021); Acicnemis suturalis Roelofs, 1875 with Wisteria brachybotrys Siebold &amp; Zucc. (Morimoto and Miyakawa 1995; Lee and Morimoto 1996); Acicnemis crassiusculus Fairmaire, 1878 with Hibiscus tiliaceus L. (1753) ( Malvaceae) and Ludwigia spp. ( Onagraceae) (Zimmerman 1943; Oziegbe and Faluyi 2010); the larvae of one unidentified Acicnemis species were taken from a dead stem of Castanopsis sieboldii (Makino) Hatus ( Fagaceae) (Lee and Morimoto 1996). Some species are attracted to lights (e. g., A. azumai) (JHL pers. obs.).</p></div>	https://treatment.plazi.org/id/D31A94491053581F892CFEBD2F63C355	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
CE8417E50CC25A60A94704F37FED721C.text	CE8417E50CC25A60A94704F37FED721C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis kiotoensis Nakane 1963	<div><p>Acicnemis kiotoensis Nakane, 1963</p><p>Figs 2 C, 3 B, 4 A, 6 F 1–6, 2, 16</p><p>Type material examined</p><p>(Japan: 73; Taiwan: 4; Total: 77). Holotype. Japan: Kyoto Prefecture: • Kurama, 17. VI. 1954, T. Nakane, bears red label reading “ HOLOTYPE ”, bears pink label reading “ 185-5 ”, Syst. Ent. Hokkaido Univ Japan SEHU 0000005125, HUM . Paratype. Japan: Kagoshima Prefecture: • Yakushima Island, Miyanoura, 28. IV. 1954, Y. Kurosawa, bears red label reading “ PARATYPE ” (1, HUM) .</p><p>Non-type material.</p><p>Japan: Kagoshima Prefecture: • Kirishima, 18. IX. 2007, J. Aoki (2, KUM), JHL_AREV_161, JHL_AREV_162 ; • Mt. Kirishima, 22. VII. 1956, J. Nagao, (1, KUM), JHL_AREV_775 ; • Amami Oshima, Hatsuno, 25. V. 1960, T. Shibata (1, KUM), JHL_AREV_177 ; • Chuorindo, Amami Oshima, 27. VI. 1980, A. Izumi (1, LBM), LBM 5117 ; • Chuorindo, Amami Oshima, 30. VI. 1980, Yoshikawa (1, LBM), LBM 5119 ; • Hatsuno, Amami Oshima, 1–8. VII. 1973, M. Ito (1, LBM), LBM 5130 ; • Yuwan, Amami Oshima, 7. VIII. 1964, M. Nagai (1, HUM), JHL_AREV_645 ; • Amami Oshima, 21. VII. 1955, T. Shirozu (1, KUM), JHL_AREV_660 ; • Ishihara-rindo, Amami Oshima, 9–11. VI. 1978, R. Noda (1, KUM), JHL_AREV_661 ; • Nonigawa, Amami Oshima, 17. VI. 1963, J. Nagao (1, KUM), JHL_AREV_662 ; • Hatsuno, Amami Oshima, 27. V. 1960, T. Shibata (1, ZMH), ZMH 837205 ; • Yudomari-rindo, Yakushima Island, 26. VII. 1982, A. Seki (2, LBM), LBM 5120, 5121 ; • Miyanoura, Yakushima Island, 17. VII. 1968, H. Nomura (1, LBM), LBM 5124 ; • Miyanoura, Yakushima Island, 21–22. VII. 1974, K. Emoto (2, LBM), LBM 5126, 5127 ; • Miyanoura, Yakushima Island, 15–26. VII. 1972, H. Huzita (2, LBM), LBM 5129, 5131 ; • Miyanoura, Yakushima Island, 26–29. VII. 1974, T. Mikage (5, KUM), JHL_AREV_648 – JHL_AREV_650 ; • Miyanoura, Yakushima Island, 29. VIII. 1952, Takeya &amp; Hirashima (1, KUM), JHL_AREV_653 ; • Miyanoura, Yakushima Island, 18. VI. 1975, Y. Kurosawa (1, NMNS), JHL_AREV_679 ; • Okawa-rindo, Yakushima Island, 1–2. VIII. 1974, T. Mikage (2, KUM), JHL_AREV_646, JHL_AREV_647 ; • Okawa, Yakushima Island, 22. VII. 1975, H. Akiyama (1 dissected male, NMNS), JHL_AREV_157 ; • Shiratani, Yakushima Island, 26–30. VII. 1979, M. Miura (8, HUM), JHL_AREV_634 – JHL_AREV_641 ; • Shiratani, Yakushima Island, 24–25. VII. 1974, T. Mikage (5, KUM), JHL_AREV_657 – JHL_AREV_659 ; • Kosugidani, Yakushima Island, 19. VII. 1974, T. Mikage (1, KUM), JHL_AREV_656 ; • Kosugidani, Yakushima Island, 23. VII. 1950, T. Shirozu (1, KUM), JHL_AREV_655 ; • Kosugidani, Yakushima Island, 9. – 10. VII. 1952, Y. Kurosawa (4, NMNS), NSMT-I-C 10267 –10269, NSMT-I-C 10301 ; • Hanayama Natural Forest, Yakushima Island, 21. VIII. 1983, Sk. Yamane (3, HUM), JHL_AREV_642 – JHL_AREV_644 ; • Mt. Hoyoshi, Minamiosumi-cho, 24–25. III. 2018, R. Ito (1, KUM), JHL_AREV_840 ; • Mt. Yuwan, Amami-Oshima, 30. VII. 1963, L. Gressitt (1 dissected male, KUM), JHL_SYN_152 ; Kyoto Prefecture: • Mt. Kurama, 30. V. 1982, K. Masaki (1, LBM), LBM 5118 ; • same locality, 8. X. 1978, K. Masaki (1, LBM), LBM 5122; Miyazaki Prefecture: • Miyakonojo-shi, Natsu-cho, Miike, 22. VI. 2019, R. Ito (1, KUM), JHL_AREV_175 ; • Mt. Aoidake, 7. VIII. 1974, K. Morimoto (1 dissected male, KUM), JHL_DNA_147 ; Nagasaki Prefecture: • Mt. Taterasan, Izuhara, Tsushima Island, 24–27. VII. 1985, A. Saito (15, CMIC), CBM–ZI 68476 –68490 ; • Mt. Tatera, Tsushima Island, 18–20. V. 1961, H. Kamiya (2, KUM), JHL_AREV_651, JHL_AREV_652 ; Nara Prefecture: • Kasugayama, 25. VII. 1981, K. Masaki (1, LBM), LBM 5116 ; • Kasugayama, 10. V. 1964, K. Kinugasa (1, LBM), LBM 5133 ; • Kasugayama, 20. V. 1951, K. Sawada (1, KUM), JHL_AREV_654 ; • Mt. Kasuga, 9. V. 1952, K. Sawada (1 dissected male, KUM), JHL_SYN_153 ; Okinawa Prefecture: • Okinawa Island, Yonahadake, 8–29. VI. 1977, H. Irie (2, KUM), JHL_AREV_172, JHL_AREV_173 ; • Iriomote Island, Shirahama, 23–24. VI. 1970, H. Makihara (2, KUM), JHL_AREV_158, JHL_AREV_159 ; • same locality, 31. VIII – 5. IX. 1969, H. Makihara (6, KUM), JHL_AREV_163 – JHL_AREV_168; • same locality, 26. VII. 1963, Y. Miyatake (1, KUM), JHL_AREV_169; • Okinawa Island, Yona, 9–13. VIII. 1969, H. Makihara (1, KUM), JHL_AREV_176 ; • Iriomote Island, Nakara River, 25–28. VI. 1970, H. Makihara (2, KUM), JHL_AREV_170, JHL_AREV_171 ; • Iriomote Island, Taketomi-cho, Mt. Tedo, 2. X. 2017, K. Narita (1, KUM), JHL_AREV_174 ; • Iriomote Island, 28. VI. 1978, M. Kinjo (1, RUMC), JHL_AREV_524 ; • Iriomote Island, Funaura, 8. X. 1977, M. Arasaki (1, RUMC), JHL_AREV_525 . Taiwan • Mt. Hsin Kao, 1. VII. 1961, S. Ueno, JHL_AREV_160 (1, KUM) ; • Meifeng, 15. V. 1983, H. Townes (1, CMNC), JHL_AREV_667 ; • Anmashan, Taichung Hsien, 11–15. V. 1992, A. Smetana (2, CMNC), JHL_AREV_707, JHL_AREV_708 .</p><p>Diagnosis.</p><p>Body length 4.5–7.0 mm. Covered in pale and dark brown scales, with two obliquely shaped pale brownish, black-bordered spots across middle of elytra. Second funicular antennomere longer than funicular antennomeres 3 + 4 (Fig. 2 C) (compare with A. koguma, which has the second funicular antennomere shorter than funicular antennomeres 3 + 4). Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum diagnostic: large, shiny, and lacking scales (Fig. 3 B). Odd and 2 nd elytral intervals with erect scales (scales beginning at middle of elytra on 2 nd interval; from elytral base on odd intervals). Sclerolepidia indistinct, forming low, rounded, non-protruding tubercles (Fig. 4 A). Third tarsomeres emarginate (not truncate as in A. koguma). Pedon with lateral edges approx. parallel in dorsal view but converging evenly into a wide, blunt point. Internal sac not visible inside the aedeagus, but with a non-hooked protruding structure (unmodified in posterior 1 / 2, with roughened in anterior 1 / 2) that is clearly longer than the pedon (Fig. 6 F 1 – 2).</p><p>Distribution.</p><p>This distinctive species occurs in Japan (mainland, throughout the Ryukyu Islands) and Taiwan.</p></div>	https://treatment.plazi.org/id/CE8417E50CC25A60A94704F37FED721C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
6FA0DC04D9645807AD2E8B6E193B52DE.text	6FA0DC04D9645807AD2E8B6E193B52DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis koguma Lewis & Kojima 2025	<div><p>Acicnemis koguma Lewis &amp; Kojima sp. nov.</p><p>Figs 2 D, 3 C, 17</p><p>Type material examined</p><p>(Japan: 3). Holotype. Japan: Wakayama Prefecture: • Mt. Otoh, 20. VII. 1980, I. Matoba, bears a red holotype label reading “ HOLOTYPE / Acicnemis koguma / Lewis &amp; Kojima, 2025 ”, JHL_DNA_143, female, deposited in KUM . Paratypes. Japan • Tokyo Metropolis: Mt. Takao, 12. VIII. 1969, H. Fujita, LBM accession # 5128 (1, LBM), JHL_SYN_141 ; • Mt. Takao, 3–4. VII. 1965, K. Sakai (1, CMIC), CBM-ZI 68474 .</p><p>Diagnosis.</p><p>Body length 4.5–7.0 mm. Covered in pale and dark brown scales, with two obliquely shaped pale brownish, black-bordered spots across middle of elytra. Second funicular antennomere shorter than funicular antennomeres 3 + 4 (Fig. 2 D) (longer in A. kiotoensis). Pronotum with impunctate longitudinal midline (absent in A. kiotoensis). Scutellum large (somewhat narrower than A. kiotoensis) and bare dorsally but bordered laterally by pale yellow scales (Fig. 3 C). Only odd elytral intervals with erect scales (some erect scales on 2 nd interval in apical fifth of elytra) ( A. kiotoensis with erect scales on the 2 nd elytral interval). Sclerolepidia indistinct, forming low, rounded, non-protruding tubercles. Third tarsomeres truncate (emarginate in A. kiotoensis).</p><p>Distribution.</p><p>This rare species is only known from two localities in mainland Japan, Mt. Takao (Tokyo Metropolis) and Mt. Otoh (Wakayama Prefecture).</p><p>Etymology.</p><p>The specific name koguma means little bear in Japanese, and is a reference to the apparent mountain-loving preference and small, brown nature of this rarely observed weevil species. We also suggest the Japanese common name コグマカレキゾウムシ [Koguma-kareki-zoumushi], which in English means “ Little-bear-dead-tree-weevil ”.</p><p>Remarks.</p><p>This cryptic species was clearly overlooked due to its remarkable similarity with A. kiotoensis in overall color and appearance, but differs consistently in the five characters listed in the diagnosis. The male remains unknown.</p></div>	https://treatment.plazi.org/id/6FA0DC04D9645807AD2E8B6E193B52DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
BBBD59AA596D5FC7A6ECA6D76421B4F8.text	BBBD59AA596D5FC7A6ECA6D76421B4F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis laeta Hubenthal 1919	<div><p>Acicnemis laeta Hubenthal, 1919</p><p>Figs 1, 2 B, 6 G 1–6, 2, 18</p><p>Type material examined</p><p>(Taiwan: 71). Lectotype (designated here). Taiwan • Fuhosho, “ IX. 07–09 ”, Sauter, SDEI, SDEI Muncheberg Col 20204, bears the original red label reading “ Syntypus ” as well as a new, red lectotype label reading “ LECTOTYPE / Acicnemis laeta / Hubenthal, 1919 / Det. Lewis &amp; Kojima, 2025 ” . Paralectotypes. Taiwan • Fuhosho, “ IX. 07–09 ”, Sauter (22, SDEI), SDEI Muncheberg Col 18617–18623 &amp; 20199–20203 &amp; 20205–20214, all bear a red label reading “ Syntypus ” and new, yellow paralectotype labels ; • Hoozan, “ 08–10 ”, Sauter (18, SDEI), SDEI Muncheberg Col 18624–18632 &amp; 20215–20223, all bear a red label reading “ Syntypus ” and new, yellow paralectotype labels ; • Kosempo, Sauter (2, SDEI), SDEI Muncheberg Col 18633–18634, bear a red label reading “ Syntypus ” and new, yellow paralectotype labels .</p><p>Non-type material.</p><p>Taiwan • Lingui, 18. VI. 1979, S. Nakamura, W. Chen (1, KUM), JHL_AREV_526 ; • Pi Shan Spa, Tai Tung Hsein, 2. VI. 1987, K. Baba (2, KUM), JHL_AREV_527, JHL_AREV_528 ; • Guandaoxi, I. VI. 1973, K. Kojima (2, KUM), JHL_AREV_529, JHL_AREV_530 ; • Jihyueh-tan, Nantou Hsein, 12. V. 1977, H. Fujita (1, KUM), JHL_AREV_531 ; • Kenting Park, Pingtung Hsein, 11. V. 1970, M. Takagi (1, KUM), JHL_AREV_532 ; • Nanshanchi, Nantou Hsein, 7. V. 1977, T. Niisato (1, KUM), JHL_AREV_533 ; • Liu Kui, 28. III. – 10. IV. 1986, K. Baba (2, KUM), JHL_AREV_534, JHL_AREV_535 ; • Kenting, Pingtung Hsein, 2. IV. 1965, S. Ueno (2, KUM), JHL_AREV_536, JHL_AREV_537 ; • Nanshanchi, 30. IV. 1965, T. Nakane, T. Shirozu (2, KUM), JHL_AREV_538, JHL_AREV_539 ; • Nanshanchi, Nantou Hsein, 12. VII. 1966 (1, KUM), JHL_AREV_540 ; • Shyk Shan, near Liu Kui, 25. IV. 1986, K. Baba (1 dissected male, KUM), JHL_AREV_541 ; • Nanshanchi, 3. V. 1979, K. Emoto (1, KUM), JHL_AREV_664 ; • Lienhoachi, 12. V. 1975, S. Imasaka (1, TUA), JHL_AREV_665 ; • “ community park ”, 28. X. 1976 (1, TUA), JHL_AREV_666 ; • Nantou, 24. V. 1983, K. Ra, (1, HUM), JHL_AREV_773; • Wushe, 29. IV. – 3. V. 1983, H. Townes (3, CMNC), JHL_AREV_669 – JHL_AREV_671 ; • Liukei, Kaohshiiung, 29. IV. – 8. V. 1982, H. Takizawa (3, HUM), JHL_AREV_675 – JHL_AREV_677 ; • near Liukuei, Kaohsiung Hsien, 5–9. IV. 1995, H. Kojima (1, TUA), JHL_AREV_690 ; • “ Formosa ”, T. Kano (1, NMNS), JHL_AREV_680 ; • Kao; • Nanshan Riv. Trail, Ren’ai Township, Nantou County, 18–19. X. 2017, K. Narita (2, KUM), JHL_DNA_124 (GenBank accession number: PV 255630), JHL_AREV_125 (GenBank accession number: PV 255631) .</p><p>Diagnosis.</p><p>Body length 4.1–5.8 mm. Covered in pale brown scales, with alternating white and black scaled bands on odd elytral intervals. Scutellum covered in yellow to brown scales. Only odd elytral intervals with erect scales. Sclerolepidia distinct and protruding. Posterior edge of prosternum with prominent projections contiguous with the fore-coxae (Fig. 2 B). Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a moderately acute point in apical 1 / 2. Internal sac protruding and with two bead-shaped sclerotized structures in basal 1 / 2, and a vase-shaped structure in apical 1 / 2 with roughened musculature (Fig. 6 G 1 – 2).</p><p>Distribution.</p><p>We only encountered this species from Taiwan. The records from Japan and the Kuril Islands (Russia / Japan) noted in Alonso-Zarazaga et al. (2023) are almost certainly erroneous and this species may have been confused with A. azumai . This species may plausibly occur in mainland Asia (China) (listed as occurring there by Alonso-Zarazaga et al. (2023)), but we have yet to examine any specimens of A. laeta from this region.</p><p>Remarks.</p><p>This species is superficially quite similar to A. azumai, but differs in pronotal, prosternal, and genital morphology.</p></div>	https://treatment.plazi.org/id/BBBD59AA596D5FC7A6ECA6D76421B4F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
DDDC73717A1C5A5B942A07135233633F.text	DDDC73717A1C5A5B942A07135233633F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis luteomaculata Morimoto & Miyakawa 1995	<div><p>Acicnemis luteomaculata Morimoto &amp; Miyakawa, 1995</p><p>Figs 1, 2E, 6 H 1–6, 2, 19</p><p>Type material examined</p><p>(Japan: 45). Holotype. Japan: Fukuoka Prefecture: • Hiko-san, 13–14. VI. 1957, K. Morimoto, bears red label reading “ (HOLOTYPE) Acicnemis luteomaculata Morimoto et. Miyakawa, 1994 ”, ELKU, ELKU 2961 . Paratypes. Japan: Ehime Prefecture: • Hojo-shi, Mt. Takanawa, 18. V. 2003, T. Kurihara (1, KUM), JHL_AREV_420 ; Fukuoka Prefecture: • Hiko-san, 8. VI. – 13. VII. 1957, K. Morimoto (11, KUM), JHL_AREV_181 – JHL_AREV_190, JHL_ACITAI_090 (dissected male) ; • same locality, 4. VI. 1958, K. Morimoto (2, KUM), JHL_AREV_192, JHL_AREV_193; • same locality, 10. VII. 1938, Hori, Kawahara &amp; Yasumatsu (1, KUM), JHL_AREV_194; • Mt. Wakasugi, 18. XII. 1955, K. Oshima (1, KUM), JHL_AREV_120 ; • Wakasugi, 28. VII. 1940, T. Shirozu (2, KUM), JHL_AREV_405, JHL_AREV_406 ; Fukushima Prefecture: • Mt. Shizukura-yama, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.6266&amp;materialsCitation.latitude=37.3756" title="Search Plazi for locations around (long 139.6266/lat 37.3756)">Mishima Town</a>, 37.3756°N, 139.6266°E, 11–20. VII. 2018, K. Narita (1, KUM), JHL_AREV_412 ; Kochi Prefecture: • Mt. Tebako, 7–10. VIII. 1957, K. Morimoto (2, KUM), JHL_AREV_196, JHL_AREV_197 ; Nara Prefecture: • Mt. Odaigahara, 2. VII. 1978, H. Hiramatsu (1, KUM), JHL_AREV_198 ; • Kasugayama, 25. VII. 1959, T. Hozumi (1, KUM), JHL_AREV_121 ; • Kasuga, 5–31. V. 1959, K. Ueda (3, KUM), JHL_AREV_401 – JHL_AREV_403 ; • Kasuga, 30. IV. 1961, K. Ueda (1, KUM), JHL_AREV_404 ; Okayama Prefecture: • Takahashi City, Mt. Gagyuzan, 19–20. V. 1975, H. Irie (3, KUM), JHL_AREV_178 – JHL_AREV_180 ; Wakayama Prefecture: • Mt. Gomadan, 28. VII. 1979, I. Matoba (1, KUM), JHL_AREV_191 .</p><p>Non-type material.</p><p>Japan: Ehime Prefecture: • Hojo-shi, Mt. Takanawa, 18. V. 2003, T. Kurihara (1, KUM), JHL_AREV_420 ; • Ishizuchi Mt., Omogo Valley, 18–25. VIII. 1980, S. &amp; J. Peck (1, CMNC), JHL_AREV_668 ; Fukuoka Prefecture: • Hiko-san, 23–26. VII. 1968, M. Chujo (1, KUM), JHL_AREV_195 ; Hiroshima Prefecture: • Sandankyo, 12–13. V. 1969, M. Chujo (1, KUM), JHL_AREV_418 ; Miyazaki Prefecture: • Miyakonojo-shi, Natsuo-cho, Miike, 22. VI. 2019, R. Ito (6, KUM), JHL_AREV_407 – JHL_AREV_411, JHL_DNA_117 (GenBank accession number: PV 255632) ; Nara Prefecture: • Kasuga, 10. V. 1959, T. Shibata (1, KUM), JHL_ACITAI_091 ; Oita Prefecture: • Mt. Kuro-dake, 28. V. 1990, S. Ogata (4, KUM), JHL_AREV_413 – JHL_AREV_416 ; • Tsukahara, Yufuin-cho, Yufu-shi, 21. XI. 2016, R. Ito (1, KUM), JHL_AREV_721 ; • Tsukahara, Yufuin-cho, Yufu-shi, 27. V. 2018, R. Ito (1, KUM), JHL_DNA_123 (GenBank accession number: PV 255633) ; Wakayama Prefecture: • Komoridani, 21. VIII. 1941, K. Sakaguchi (1, HUM), JHL_AREV_678 ; Yamagata Prefecture: • Tozawa-mura, Mt. Haguro, 23. VI. 2003, J. Aoki (1, KUM), JHL_AREV_417 .</p><p>Diagnosis.</p><p>Body length 4.0– 5.4 mm. Covered in pale brown scales, with two obliquely shaped tan-colored, black-bordered spots across middle of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum sparsely covered in pale yellow scales. All elytral intervals with erect scales. Sclerolepidia indistinct, forming low, rounded, non-protruding tubercles. Third tarsomeres distinctly truncate (Fig. 2 E) (compare with the superficially similar A. kiotoensis, A. maculaalba, and A. sauteri, which all have emarginate third tarsomeres). Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a blunt, rounded point. Internal sac not visible inside the aedeagus, but with a basal projection that bears a distinctive rectangular sclerite anteriorly at its apex (Fig. 6 H 1 – 2).</p><p>Distribution.</p><p>This species is found in mainland Japan (except Hokkaido) and has also been reported from South Korea (Park et al. 2009).</p></div>	https://treatment.plazi.org/id/DDDC73717A1C5A5B942A07135233633F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
874918711E505A9BAD08F51AD234E18C.text	874918711E505A9BAD08F51AD234E18C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis maculaalba Roelofs 1875	<div><p>Acicnemis maculaalba Roelofs, 1875</p><p>Figs 1, 2 G, 3 D, 6 I 1–6, 2, 20</p><p>Type material examined</p><p>(Japan: 50). Holotype. Japan • unknown locality, “ 1910–320 ”, G. Lewis, bears a circular white, red-bordered label reading “ Type H. T. ”, NHMUK 015024192 .</p><p>Non-type material.</p><p>Japan: Fukuoka Prefecture: • Mt. Homan, 15. V. 1975, H. Irie (1, KUM), JHL_AREV_583, misidentified A. luteomaculata paratype ; Kagoshima Prefecture: • Sata (Osumi), Magome-Odomari, 28. V. 1952, Esaki &amp; Hirashima (1, KUM), JHL_AREV_556 ; • Kuchinoshima, 26. VI. – 3. VII. 1969, H. Makihara (3, KUM), JHL_AREV_557 – JHL_AREV_559 ; • Akuseki Island, 30. VII. 1969, M. Sakai (4, KUM), JHL_AREV_560 – JHL_AREV_563 ; • Nakanoshima, 25. IV. – 2. V. 1975, H. Irie (4, KUM), JHL_AREV_564 – JHL_AREV_567 ; • Amami-Oshima, Hatsuno, 12. VI. 1963, J. Nagao (1, KUM), JHL_AREV_570 ; • Yakushima Island, Miyanoura, 26. VII. 1974, T. Mikage (1, KUM), JHL_AREV_571 ; • Funayuki, Yakushima Island, 26. VII. 1989, M. Satoshi (1, CMIC), CBM ZI 159703 ; Kochi Prefecture: • Okinoshima ( Tosa Province), 28. VI. – 2. VIII. 1953, K. Morimoto &amp; K. Sugimoto (5, KUM), JHL_AREV_551 – JHL_AREV_555 ; Mie Prefecture: • Mie University Forest, Ichishi-gun, 11. VI. 1956, M. Ichihashi (2, KUM), JHL_AREV_546, JHL_AREV_547 ; Miyazaki Prefecture: • Mt. Aoidake, 6. V. 1976, K. Morimoto, Z. Kuranaga, A. Iwasaki, N. Yoshida (1, KUM), JHL_AREV_590 ; Okinawa Prefecture: • Komi, Iriomote Island, 14. III. 1997, M. Chujo (1, KUM), JHL_SYN_143 ; • Iriomote Island, 28. V. 1962, K. Kojima (1, KUM), JHL_AREV_545 ; • Yonahadake, 8–29. VI. 1977, H. Irie (2, KUM), JHL_AREV_568, JHL_AREV_569 ; • Hiji, Okinawa Island, 3. VII. 1974, T. Mikage (2, KUM), JHL_AREV_572, JHL_AREV_573 ; • Yona, Okinawa Island, 15. VI. 1970, H. Makihara (1, KUM), JHL_SYN_142 ; • Yona, Okinawa Island, 29. VI. 1976, H. Makihara (1, KUM), JHL_AREV_574 ; • Mt. ufudaki, Kohama Is., 29. V. 2022, S. Imada, (1, KUM), JHL_AREV_777 ; • Mantabaru, Yonaguni Island, 25–27. V. 2022, S. Imada (3, KUM), JHL_AREV_575 – JHL_AREV_577 ; • Omoto-dake, Ishigaki Island, 30. VI. 1974, T. Mikage (1, KUM), JHL_AREV_578 ; • Mt. Banna, Ishigaki Island, 23–26. V. 1990, K. Morimoto (1, KUM), JHL_AREV_579 ; • Shirahama, Iriomote Island, 31. VIII. – 5. IX. 1969, H. Makihara (3, KUM), JHL_AREV_580 – JHL_AREV_582 ; • Funaura, Iriomote Island, 1. X. 1978, S. Azuma (1, RUMC), JHL_AREV_713 ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.79491&amp;materialsCitation.latitude=26.28557" title="Search Plazi for locations around (long 127.79491/lat 26.28557)">Nakagusuku Park</a>, Okinawa Island, 26.28557°N, 127.79491°E, 3–17. II. 2016, OKENT 0068361 (GenBank accession number: PV 255635) (1, OIST) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.23598&amp;materialsCitation.latitude=26.73974" title="Search Plazi for locations around (long 128.23598/lat 26.73974)">Yona Field</a>, Okinawa Island, 26.73974°N, 128.23598°E, 8–22. VII. 2016, OKENT 0050489 (GenBank accession number: PV 255634) (1, OIST) ; • Toyko District: Oji-ike, Miyake Island, 5–7. IV. 2010, J. Aoki (6, KUM), JHL_AREV_584 – JHL_AREV_589 ; • Sato, Mt. Kurosa – Kitakao-san, Mikura-jima Island, 2. VI. 1979, J. Okuma (1 dissected male, KUM), JHL_AREV_426 ; Wakayama Prefecture: • near Mt. Ohtoh, 13. VI. – 7. VIII. 1980, I. Matoba (2, KUM), JHL_AREV_548, JHL_AREV_549 ; • Mt. Ohtoh, 16–19. VII. 1982, H. Makihara (1, KUM), JHL_AREV_500 .</p><p>Diagnosis.</p><p>Body length 2.6–5.0 mm. Covered in pale and dark brown scales, with two obliquely shaped pale brownish, black-bordered spots across middle of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum small, bordered by pale, yellow scales, and weakly bare dorsally or covered in scales (Fig. 3 D) (compare with A. kiotoensis which possesses a large, broad, bare scutellum). Odd, 2 nd, and 4 th elytral intervals with erect scales. Sclerolepidia indistinct, forming low, rounded, non-protruding tubercles. Third tarsomeres emarginate (Fig. 2 G) (as opposed to truncate, as in A. koguma). Pedon with lateral edges approx. parallel in dorsal view but converging in distal 1 / 2 evenly into a blunt point. Pedon stout (pedon length / total aedeagus length including temo = 0.2). Internal sac protruding and with roughened musculature across length; with two bent, tube-like sclerotized structures anteriorly at its apex (Fig. 6 I 1 – 2).</p><p>Distribution.</p><p>This species occurs broadly throughout Japan (mainland Japan and Ryukyu Islands) and Taiwan and is also recorded from China by Alonso-Zarazaga et al. (2023).</p><p>Remarks.</p><p>Acicnemis maculaalba forms a species group with A. sauteri . The association of A. maculaalba and A. sauteri is supported by morphology and weakly (BS: 38) by our CO 1 gene tree. See also A. cordata Remarks section.</p></div>	https://treatment.plazi.org/id/874918711E505A9BAD08F51AD234E18C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
3A98A0486003545BA3018AA6A9CF6822.text	3A98A0486003545BA3018AA6A9CF6822.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis nobilis Hubenthal 1919	<div><p>Acicnemis nobilis Hubenthal, 1919</p><p>Figs 6 J 1–6, 2, 21</p><p>Type material examined</p><p>(Taiwan: 10). Holotype. Taiwan • Hoozan, “ H. Sauter 1910 ”, bears small yellow label reading “ 1911 14 ”, bears red label reading “ Typus ”, SNSD .</p><p>Non-type material.</p><p>Taiwan • Penpuchi, Nantou Hs., 13. VII. 1966, H. Sasaji (2, KUM), JHL_SYN_151 (dissected male), JHL_AREV_492 ; • Sangkang, Nantou Hs., 14. VII. 1966, H. Sasaji (3, KUM), JHL_SYN_150 (dissected male), JHL_AREV_491, JHL_AREV_493 ; • Lushan, Nantou Hsein, 8. VI. 1976, H. Makihara (1, KUM), JHL_AREV_494 ; • Nanshanchi, Wenchuan Hs., 27. IV. – 4. V. 1977, T. Mikage (1, KUM), JHL_AREV_495 ; • Hsi-Leng, 3. V. 1981, S. Tsuyuki (1, TUA), JHL_AREV_496 ; • Kaohsiung, 5. VIII. 1978, T. Niisato (1, TUA), JHL_AREV_497 .</p><p>Diagnosis.</p><p>Body length 4.1–5.2 mm. Covered in dark gray and black scales, with a pale-yellow scale, V-shaped, transverse band across middle of elytra. Basal patch of black scales surrounding scutellum approx. V-shaped and extending prominently onto the second elytral intervals (as opposed to the similar A. nohirai which has an elongate I-shaped patch of black scales on the first elytral interval that barely extends on the second elytral interval basally). Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum covered in pale scales. Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but sinuating into a blunt, rounded point. Internal sac protruding from aedeagus, with roughened musculature posteriorly and a club-shaped sclerotized structure anteriorly at its apex (Fig. 6 J 1 – 2).</p><p>Distribution.</p><p>This species is currently only known from Taiwan.</p></div>	https://treatment.plazi.org/id/3A98A0486003545BA3018AA6A9CF6822	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
99B40C3B45FA56D283181925B0964925.text	99B40C3B45FA56D283181925B0964925.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis nohirai Morimoto & Miyakawa 1995	<div><p>Acicnemis nohirai Morimoto &amp; Miyakawa, 1995</p><p>Figs 1, 6 K 1–6, 2, 22</p><p>Type material examined</p><p>(Japan: 23). Holotype. Japan: Gifu Prefecture: • Hirayu, 19. VI. 1963, T. Nohirai, bears a red label reading “ (Holotype) Acicnemis nohirai Morimoto et Miyakawa, 1994 ”, ELKU, ELKU 2956 . Paratypes. Japan: Aomori Prefecture: • Kawauchi, Shimokita-gun, 25. VII. 1984, S. Yamauchi (1, KUM), JHL_AREV_435 ; Ehime Prefecture: • Omogokei, 15. VI. 1981, S. Naomi (1, KUM), JHL_AREV_437 ; Gifu Prefecture: • Hirayu, 19. VI. 1963, T. Nohirai (5, KUM), JHL_AREV_429 – JHL_AREV_433 ; • Hokkaido: Horoka, Kamishihoro-cho, 17. VII. 1976, H. Irie (1, KUM), JHL_AREV_434 ; • Uchida, 18. V. 1930 (1, HUM), JHL_AREV_454 ; • Sapporo, “ 3. VIII. 80 ” (1, KUM), JHL_AREV_460 ; Nagano Prefecture: • Tobira Spa, Matsumoto, 7. VIII. 1973, T. Mikage (1 dissected male, KUM), JHL_SYN_147 ; Niigata Prefecture: • Kurokawa, 30. VII. 1962, K. Baba (1 dissected male, KUM), JHL_SYN_146 ; Oita Prefecture: • Kurasame, Kuju, 15. VI. 1975, H. Irie (2, KUM), JHL_AREV_438, JHL_AREV_439 ; Tochigi Prefecture: • Chuzenji-Yumoto, Shimotsuke, 22. VII. 1923, T. Esaki (1, KUM), JHL_AREV_436 ; Yamanashi Prefecture: • Fuji-rindo, Mt. Fuji, 10–12. VIII. 1976, H. Irie (1, KUM), JHL_AREV_440 ; • Mt. Kaikoma, 13. VIII. 1962, K. Oshima (1, KUM), JHL_AREV_441 ; • Masutomi, Nirasaki, 16. VII. 2006, H. Takizawa (1, HUM), JHL_AREV_461 .</p><p>Non-type material.</p><p>Japan: Gifu Prefecture: • Hirayu, 19. VI. 1963, T. Nohirai (1, KUM), JHL_AREV_442 ; Nagano Prefecture: • Otaki-mura, 31. V. 2010, K. Takahashi (1, KUM), JHL_DNA_119 (PV 255636) ; Oita Prefecture: • Yuhu-shi, Yuhuin-chou, Tsukahara, 18. IX. 2016, R. Ito (1, KUM), JHL_DNA_122 (PV 255637) ; Yamanashi Prefecture: • Fuji-rindo, Mt. Fuji, 3. X. 1992, T. Mikage (1, KUM), JHL_AREV_443 ; • Mt. Daibosatsu, Hagiwara, 12. VIII. 2005, T. Kurihara (1, KUM), JHL_AREV_720 ; • Masutomi, 24–25. V. 1968, H. Takizawa, (1, HUM), JHL_AREV_774 .</p><p>Diagnosis.</p><p>Body length 4.9–6.0 mm. Covered in dark gray and black scales, with a white-scaled, V-shaped, transverse band across middle of elytra. Elytra with elongate I-shaped patch of black scales on the first elytral interval (surrounding the scutellum) that barely extends onto the second elytral interval. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum covered in dark brown scales. Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a blunt, rounded point. Internal sac not visible inside aedeagus, but with protruding basal structure that bears a prong-shaped sclerite anteriorly at its apex (Fig. 6 K 1 – 2).</p><p>Distribution.</p><p>This species is apparently endemic to mainland Japan and occurs from Hokkaido south to Kyushu (Oita Prefecture).</p></div>	https://treatment.plazi.org/id/99B40C3B45FA56D283181925B0964925	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
2D859A7357DE559B94B24FCD616645CF.text	2D859A7357DE559B94B24FCD616645CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis palliata Pascoe 1872	<div><p>Acicnemis palliata Pascoe, 1872</p><p>Figs 3 E, 4 B, 6 L 1–6, 2, 23</p><p>Acicnemis dorsonigrita Voss, 1941, syn. nov.</p><p>Type material examined</p><p>(China: 2; Japan: 20: Total: 22). Holotype. Japan: Nagasaki Prefecture: • Nagasaki, Pascoe Coll. 93–60, bears a circular white, red-bordered label reading “ Type H. T. ”, NHMUK 015026632 . Paratypes. China • Tienmuschan, bears a red label reading “ Type ” (1 dissected male, NHMB), Acicnemis dorsonigrita type specimen, JHL_ACITAI_010 ; • Tienmuschan, bears a red label reading “ Cotypus Acicnemis dorsonigrita ” (1, ZMH), ZMH 841827 .</p><p>Non-type material.</p><p>Japan: Chiba Prefecture: • Ichihara-shi, Kongouuchi, 28. V. 1989, M. Satoshi (2, CMIC), CBM-ZI 159709, CBM-ZI 159710 ; Ehime Prefecture: • Higashino, Matsuyama-shi, 1–22. VII. 2016, Y. Hisasue (2, KUM), JHL_AREV_484, JHL_AREV_485 ; Fukuoka Prefecture: • Chikuzen, Kashii, Kusaba Kosen, 5. VI. 1935, Esaki, Hori, Ho, Fujino, Yasumatsu, Nomura &amp; Kawahara (1, KUM), JHL_ACITAI_046 ; • Mt. Homan, 2. VII. 1978, R. Noda (1 dissected male, KUM), JHL_ACITAI_007 ; • Nokonosima, 8–29. V. 2005, H. Hirano (2, KUM), JHL_AREV_486, JHL_AREV_487 ; Fukushima Prefecture: • 1. VII. 2000, K. Takahashi (1, KUM), JHL_AREV_483; • Kaneyama-machi, Okuriyama, 2. VI. 2012, J. Aoki (3, KUM), JHL_AREV_488 ; • Aizu, Kaneyama, 20–21. VII. 2012, J. Aoki (4, KUM), JHL_AREV_489 ; Kagawa Prefecture: • Takamatsu-shi, 15. V. 1961, M. Chujo (1, KUM), JHL_ACITAI_051 ; • Miki-cho, 16. VI. 1961, M. Chujo (1, KUM), JHL_ACITAI_050 ; • Kinuki-shi, 25. V. 2015, J. Aoki (1, KUM), JHL_AREV_490 ; • Kita-gun, Hirai-cho, 13. V. 1949, K. Iwata (1, ZMH), ZMH 837201 ; Nagasaki Prefecture: • Tsushima, Izuhara, 15. V. 1971, H. Irie (1, KUM), JHL_ACITAI_006 ; Nara Prefecture: • Kasuga, 4. VI. 1961, K. Ueda (1, KUM), JHL_ACITAI_049 ; Oita Prefecture: • Tsukahara, Yufuin-machi, Yufu-shi, 18. VI. 2016, R. Ito, (1, KUM), JHL_AREV_779 ; Osaka Prefecture: • Mt. Iwawaki, 18. VI. 1966, H. Kimura (1, KUM), JHL_ACITAI_048 .</p><p>Diagnosis.</p><p>Body length 6.1–7.0 mm. Covered in pale brown, dark gray, and black scales, with a tan-colored, V-shaped, transverse band across middle of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum bordered by pale brown scales, but conspicuously bare and shiny dorsally (Fig. 3 E). Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding (Fig. 4 B). Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a blunt, rounded point. Internal sac not visible inside aedeagus, but with a protruding basal structure that bears two hook-like structures anteriorly at the apex that are curved 90 degrees inward (Fig. 6 L 1 – 2).</p><p>Distribution.</p><p>This species occurs in Japan (mainland) and China, and has also been reported from South Korea (Park et al. 2009) and the Kuril Islands (Alonso-Zarazaga et al. 2023).</p><p>Remarks.</p><p>An examination of the holotype of A. palliata (NHMUK), the holotype of A. dorsonigrita (NHMB), a paratype of A. dorsonigrita (ZMH), as well as comparisons of the original descriptions of these two species revealed that A. dorsonigrita syn. nov. is a junior synonym of A. palliata . In particular, the holotypes of A. palliata and A. dorsonigrita (1) both have scutella which are bare dorsally, bordered by scales around the perimeter (2) are more than 5 mm in length (measured from the apex of the elytra to the base of the rostrum), and (3) bear the same characteristic pattern of scales dorsally. Furthermore, the original descriptions of these species certainly appear to refer to the same species, and notably the body length measurements given for A. palliata (“ 3 lin. ” = 6.25 mm) and A. dorsonigrita (5–5.8 mm) are approximately equal. The much smaller species (at most 4.4 mm) previously referred to as A. dorsonigrita by Morimoto and Miyakawa (1995) is new and is described below as A. squamata .</p></div>	https://treatment.plazi.org/id/2D859A7357DE559B94B24FCD616645CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
C54830051F165B59A4CFC62B2EA1A456.text	C54830051F165B59A4CFC62B2EA1A456.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis postica Hubenthal 1919	<div><p>Acicnemis postica Hubenthal, 1919</p><p>Figs 1, 6 M 1–6, 2, 24</p><p>Acicnemis cruciata Kleine, 1924, syn. nov.</p><p>Type material examined</p><p>(Japan: 31; Taiwan: 5; Total: 36). Lectotype (designated here): Taiwan • “ Formosa Is. ”, “ Coll. Pape ”, SDEI Müncheberg Col – 18570 (1, SDEI), bears red label reading “ Syntypus ” and a new, red lectotype label reading “ LECTOTYPE / Acicnemis postica / Hubenthal, 1919 / Det. Lewis &amp; Kojima, 2025 ” . Paralectotypes. Taiwan • Fuhosho, “ IX- 7–09 ”, H. Sauter, SDEI Müncheberg Col – 18571 (1, SDEI), bears red label reading “ Syntypus ” and a new, yellow paralectotype label ; • Kankau ( Koshun), VI. 1912, H. Sauter, SDEI Müncheberg Col – 18572 (1, SDEI), bears red label reading “ Syntypus ” and a new, yellow paralectotype label .</p><p>Other type material.</p><p>• Kosempo, “ VII – 19 ”, H. Sauter, Acicnemis cruciata holotype (1, SDEI), SDEI Coleoptera # 303422, SDEI Müncheberg Col – 18568.</p><p>Non-type material.</p><p>Taiwan • Taoyuan Hsein, Paling, 24. V. 1977, H. Fujita (1, KUM), JHL_ACITAI_057 ; • Wenchuan Hsein, Lushan, 4. VII. 1978, T. Mikage (1, KUM), JHL_ACITAI_058 ; • Ping Tung Hsein, Kenting, 2. IV. 1965, S. Ueno (1, KUM), JHL_ACITAI_060 ; • Kenting, 31. III. 1970, Y. Miyake (2 dissected males, KUM), JHL_ACITAI_082, JHL_SYN_179 ; • Chihpen Wenchuan, Taitung Hsien, 7. VI. 1980, H. Makihara (1, KUM), JHL_AREV_735 ; • Jiuyuehtan, 31. V. 1978, K. Kusama (1, KUM), JHL_AREV_736 ; • Taidong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.031&amp;materialsCitation.latitude=22.8062" title="Search Plazi for locations around (long 121.031/lat 22.8062)">Lijia road</a>, 22.8062°N, 121.0310°E, 19. VIII. 2013, V. Grebennikov (1, CNCI), CNCCOLVG 00006901 (BOLD accession number: MEBLA 191-14) ; Japan: Kagoshima Prefecture: • Shinokawa, Setouchi-cho, Amami Oshima, 10. VII. 2008, K. Ozaki (1, CMIC), CBM ZI 136615 ; • Nakanoshima Island, 1–2. V. 1975, H. Irie (1, KUM), JHL_AREV_750 ; • Nagata, Yakushima Island, 17. V. 1960, J. Nagao (1, KUM), JHL_AREV_754 ; • Mt. Kindake, Nakano Is., 7–9. VI. 2019, S. Imada, (1, KUM), JHL_AREV_780 ; Okinawa Prefecture: • Ishigaki Island, Mt. Omoto, 18–22. VI. 1991, K. Morimoto (1, KUM), JHL_ACITAI_055 ; • Mt. Omoto, 21. V. 1977, H. Irie (1, KUM), JHL_AREV_755 ; • Mt. Omoto, 18–21. IV. 1975, H. Irie (1 dissected male, KUM), JHL_ACITAI_062 ; • Ushikumori, Iriomote Island, 7. X. 1963, K. Morimoto (1, KUM), JHL_AREV_756 ; • Yonaguni Island, Urabu-dake, 25–26. VI. 1974, T. Mikage (6, KUM), JHL_ACITAI_056, JHL_AREV_751 – JHL_AREV_753 ; • Okinawa Island, Hiji, 9. VII. 1974, T. Mikage (2, KUM), JHL_ACITAI_059 ; • Fusato, Okinawa Island, 5. VI. 1970 (1, KUM), JHL_AREV_762 ; • Omotodake, Ishigaki Island, 8. VII. 1968, M. Takagi (1, KUM), JHL_AREV_761 ; • Funaura, Iriomote Island, 2. X. 1978 (2, RUMC), JHL_AREV_709, JHL_AREV_710 ; • Banna, Ishigaki Island, 14. III. 1964, S. Azuma (1, RUMC), JHL_AREV_712 ; • Haneji, Okinawa Island, 10. VIII. 1979, T. Kohama (1, RUMC), JHL_AREV_711 ; • Mt. Inbi-take, Yonaguni Island, 2. VI. 2021, S. Imada (1, KUM), JHL_AREV_723 ; • Kuburadake, Yonaguni Island, 15. VI. 2001, L. Hirano (1, KUM), JHL_AREV_725 ; • Shirahama, Iriomote Island, 31. VIII – 5. IX. 1969, H. Makihara (9, KUM), JHL_AREV_726 – JHL_AREV_734 ; • Shirahama, Iriomote Island, 4. X. 1963, S. Ueno (1, KUM), JHL_AREV_737 ; • upper Nakara, Iriomote Island, 12. III. 1964, Y. Miyatake (1, KUM), JHL_AREV_738 ; • Komi, Iriomote Island, 12. VII. 1963, Y. Miyatake (1, KUM), JHL_AREV_739 ; • Sonai, Yonaguni Island, 25–29. VIII. 1969, H. Makihara (10, KUM), JHL_AREV_740 – JHL_AREV_749 ; • Okinawa Island (1, OIST), JHL_AREV_983 (GenBank accession number: PV 255638); • Tancha, Okinawa Island, 18. V. 2024, J. H. Lewis, sweeping vegetation (1, OIST), JHL_AREV_994 .</p><p>Diagnosis.</p><p>Body length 3.2–5.2 mm. Covered in orange, white, dark and pale gray scales. Elytra with inner five or six intervals black and gray; outer intervals with pale brown to orange scales; with V-shaped band of white scales along middle three or four intervals in center of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum small and covered in erect, hair-like yellow scales. Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres emarginate. Metatibia evenly curved, not sinuate as in most other species. Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a blunt, rounded point. Internal sac not visible inside aedeagus, but protruding and with two sclerotized cylinder-shaped structures posteriorly near the base (Fig. 6 M 1 – 2).</p><p>Distribution.</p><p>This species occurs commonly throughout southern Japan (Ryukyu Islands) and Taiwan.</p><p>Remarks.</p><p>Examination of the A. cruciata Kleine, 1924 syn. nov. holotype (SDEI Coleoptera # 303422) revealed that this is a junior synonym of A. postica as it possesses all of the key characteristics that define this distinctive species (same scale pattern, evenly curved metatibiae, small, scaled scutellum). We designate one of the Hubenthal A. postica syntypes (SDEI Müncheberg Col – 18570) as a lectotype to fix the identity of this species.</p></div>	https://treatment.plazi.org/id/C54830051F165B59A4CFC62B2EA1A456	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
9B3885269B1E5202BB6E58F07231B225.text	9B3885269B1E5202BB6E58F07231B225.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis ryukyuana Lewis 2023	<div><p>Acicnemis ryukyuana Lewis, 2023</p><p>Figs 1, 2 F, 6 N 1–6, 2, 25, 32 A, C</p><p>Type material examined</p><p>(Japan: 28). Holotype. Japan: Okinawa Prefecture: • Ishigaki, Omoto-dake, 2. VII. 1991, K. Hidetada, bears black-bordered red label reading “ HOLOTYPE / Acicnemis ryukyuana / Lewis, 2023 ”, JHL_ACI_004, dissected male, deposited in KUM . Paratypes. Japan: Okinawa Prefecture: • Ishigaki Island, Yarabu-dake, 28. V. 1998, K. Takahashi (1, KUM), JHL_ACI_001 ; • Ishigaki Island, Omoto-dake, 2. VII. 1991, K. Hidetada (9, KUM), JHL_ACI_002, JHL_ACI_003, JHL_ACI_005 – JHL_ACI_011 ; • Ishigaki Island, Mt. Yarabudake, 17–20. IV. 1998, S. Ohmomo (1, KUM), JHL_ACI_012 ; • Ishigaki Island, Omotodake, 30. VI. 1974 (1, KUM), JHL_ACI_013 ; • Okinawa Island, Kunigami, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.2363&amp;materialsCitation.latitude=26.73972" title="Search Plazi for locations around (long 128.2363/lat 26.73972)">Yambaru National Park, Yona Field</a> (26.73972°N, 128.23630°E), 4–18. IX. 2015, A. Miyagi, Y. Tamaki, I. Maehira, L. Iha, S. Iriyama, T. Yoshida (2, OIST), OKENT 0063788, OKENT 0063789 (dissected male) ; • Okinawa Island, Kunigami, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.2363&amp;materialsCitation.latitude=26.73972" title="Search Plazi for locations around (long 128.2363/lat 26.73972)">Yambaru National Park, Yona Field</a> (26.73972°N, 128.23630°E), 7–21. VIII. 2015, M. Yoshimura, M. Ogasawara (1, OIST), OKENT 0053190 (GenBank accession number: PV 255639) ; • Okinawa Island, Kunigami, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.2372&amp;materialsCitation.latitude=26.73894" title="Search Plazi for locations around (long 128.2372/lat 26.73894)">Yambaru National Park, Yona Field</a> (26.73894°N, 128.23720°E), 21. VIII – 4. IX. 2015, Y. Tamaki, T. Yoshida (2, OIST), OKENT 0055037, OKENT 0053264 ; • Okinawa Island, Kunigami, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.2363&amp;materialsCitation.latitude=26.73972" title="Search Plazi for locations around (long 128.2363/lat 26.73972)">Yambaru National Park, Yona Field</a> (26.73972°N, 128.23630°E), 18. IX – 16. X. 2015, A. Miyagi, I. Maehira, T. Yoshida (1, OIST), OKENT 0053498 (GenBank accession number: PV 255640) ; • Okinawa Island, Hiji, 8. VII. 1974, T. Mikage (1, KUM), JHL_ACI_014 .</p><p>Non-type material.</p><p>Japan: Okinawa Prefecture: • Takeda-rindo, Ishigaki Island, 19–22. II. 2007, H. Makihara &amp; E. Kagaya (1, KUM), JHL _AREV_625 .</p><p>Diagnosis.</p><p>Body length 3.2–5.0 mm. Covered in dark gray, black, and bright yellow scales, with a black-bordered, yellow spot at the center of each elytron. Apex of elytra covered in yellow scales, and contrasting with overall dark dorsal surface of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma), but with an M-shaped band of yellow scales. Scutellum small, red, shiny, and largely bare over dorsal surface. Elytra with long, erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres weakly emarginate to truncate (Fig. 2 F). Pedon with lateral edges approx. parallel in dorsal view, but converging evenly into a blunt, rounded point. Internal sac not visible inside aedeagus, but with protruding basal structure that bears a distinctive U-shaped formation anteriorly at its apex (Fig. 6 N 1 – 2).</p><p>Distribution.</p><p>This species is known only from Okinawa Prefecture, Japan (Yambaru National Park (Okinawa Island) and forested regions of Ishigaki Island.</p><p>Remarks.</p><p>This species is very distinct from other Japanese Acicnemis and its range-restricted, endemic presence in pristine, subtropical rain forest dominated parts of Okinawa Prefecture was hitherto a mystery. However, preliminary morphological and molecular evidence suggest a sister relationship between A. ryukyuana and an undescribed Southeast Asian (Thai) species that possesses emarginate third tarsomeres. Acicnemis ryukyuana likely represents a lineage that was isolated from the undescribed Thai species lineage when the Ryukyu Islands separated from mainland Asia (see Osozawa et al. 2011) and evolved truncate third tarsomeres as derived character.</p></div>	https://treatment.plazi.org/id/9B3885269B1E5202BB6E58F07231B225	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
5D5F21B1AD665E6F806788A6CA815813.text	5D5F21B1AD665E6F806788A6CA815813.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis sauteri Hubenthal 1919	<div><p>Acicnemis sauteri Hubenthal, 1919</p><p>Figs 1, 6O 1–6, 2, 26</p><p>Type material examined</p><p>(Japan: 21; Taiwan: 18; Total: 39). Lectotype (designated here). Taiwan • Hoozan, 1910, H. Sauter, bears a red label reading “ Typus ” and a new, red lectotype label reading “ LECTOTYPE / Acicnemis sauteri / Hubenthal, 1919 / Det. Lewis &amp; Kojima, 2025 ”, SNSD, JHL_ACITAI_004 . Paralectotypes. Taiwan • Hoozan, “ 08–10 ”, Sauter (3, SDEI), bear a red label reading “ Syntypus ”, SDEI Muncheberg Col 18573, 18574, 18583 ; • Kosempo, Sauter (4, SDEI), bear a red label reading “ Syntypus ”, SDEI Muncheberg Col 18575, 18576, 18577, 18578 ; • Fuhosho, “ VIII- 09 ”, Sauter (3, SDEI), bear a red label reading “ Syntypus ”, SDEI Muncheberg Col 18580, 18581, 18582 ; • Taihorinsho, “ X- 09 ”, Sauter (1, SDEI), bears a red label reading “ Syntypus ”, SDEI Muncheberg Col 18579 .</p><p>Non-type material.</p><p>Taiwan • Lushan Wenchuan, Nantou Hsien, 6–7. VI. 1976, H. Makihara (2, KUM), JHL_AREV_609, JHL_AREV_611 ; • Sun Moon Lake, Nantou Hsien, 6. XII. 2013, J. Aoki (10, KUM), JHL_SYN_148 (dissected male), JHL_SYN_149 (dissected male), JHL_AREV_613 – JHL_AREV_620 ; • Wukongshan, Liu-kuei, Kaohsiung Hsein, 2. V. 1996, H. Takizawa (1, HUM), JHL_ACITAI_085 ; Japan: Kagoshima Prefecture: • Nakanoshima Island, 25–30. IV. 1975, H. Irie (1, KUM), JHL_AREV_608 ; • Jinushi-daimyojin, Satomura, Nakano-jima Island, 7–9. VII. 2019, S. Imada (1, KUM), JHL_AREV_610 ; • Sato, Nakano-jima Island, 7–9. VII. 2019, N. Tsuji (3, KUM), JHL_AREV_691 – JHL_AREV_693 ; • Aden, Kikai-jima Island, 6. V. 2022, R. Ito (3, KUM), JHL_AREV_623, JHL_AREV_908, JHL_AREV_909 ; • Kikai-jima Island, 4. III. 2012, J. Aoki (1, KUM), JHL_AREV_695 ; • Torigamine, Amami-Oshima, 3. VIII. 1976, N. Morishima (1, LBM), LBM 5123 ; • Hatsuno, Amami-Oshima, 24–30. VI. 1973, M. Ito (1 dissected male, LBM), LBM 5132 ; • Sumiyo, Amami Oshima, 27. V. 2004, K. Takahashi (1, KUM), JHL_AREV_694 ; • Akuseki-jima Island, Ue-shuraku, Sakamori-jinja, 2–6. VII. 2019, S. Imada (1, KUM), JHL_DNA_121 (GenBank accession number: PV 255642) ; • Nakano-jima Island, Satomura, Jinushi-daimyojin, 7–9. VII. 2019, S. Imada (1, KUM), JHL_DNA_120 (GenBank accession number: PV 255641) .</p><p>Diagnosis.</p><p>Body length 4.9–6.2 mm. Covered in pale and dark brown scales, with two obliquely shaped white, black-bordered spots across middle of elytra. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum small, bordered by pale yellow scales, and weakly bare antero-dorsally (compare with A. kiotoensis which possesses a large, broad, bare scutellum). Odd and 2 nd elytral intervals with erect scales (4 th elytral interval also occasionally with 1–2 scales near white spot). Sclerolepidia indistinct, forming low, rounded, non-protruding tubercles. Third tarsomeres emarginate (as opposed to truncate, as in A. koguma). Pedon with lateral edges approx. parallel in dorsal view, but converging in distal 1 / 2 evenly into a blunt point. Pedon elongate (pedon length / total aedeagus length including temo = 0.26). Internal sac protruding and unmodified in the posterior 1 / 2, but with roughened musculature in anterior 1 / 2 (Fig. 6 O 1 – 2).</p><p>Distribution.</p><p>This species occurs in southern Japan (Kagoshima Prefecture) and Taiwan, and has also been recorded from South Korea (Alonso-Zarazaga et al. 2023). The apparent lack of this species in the southern Ryukyu Islands (Okinawa Prefecture) is odd; however, similar patterns are known in other weevils (e. g., Aphanerostethus magnus Lewis &amp; Kojima, 2024)</p><p>Remarks.</p><p>This species is closely allied with A. maculaalba (see Remarks for that species). We designate one of Hubenthal’s syntypes (SNSD, JHL _ACITAI_004) as a lectotype to fix the identity of this species.</p></div>	https://treatment.plazi.org/id/5D5F21B1AD665E6F806788A6CA815813	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
F6287459457D5125A8A12D3A16A2CFE4.text	F6287459457D5125A8A12D3A16A2CFE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis shibatai Voss 1971	<div><p>Acicnemis shibatai Voss, 1971</p><p>Figs 6 P 1–6, 2, 27</p><p>Type material examined</p><p>(Japan: 17; South Korea: 1; Taiwan: 2; Total: 20). Holotype. Japan: Kagoshima Prefecture: • Ikari, Amami-Oshima, 28. V. 1960, T. Shibata, KUM, bears red label that reads “ Holotype ” . Paratypes. Japan: Kagoshima Prefecture: • Ikari, Amami-Oshima, 6. V. – 4. VI. 1960, T. Shibata, JHL_AREV_444 – JHL_AREV_451, JHL_ACITAI_064 (9, KUM), bear red label that reads “ Paratype ” .</p><p>Non-type material.</p><p>Japan: Chiba Prefecture: • Mt. Kiyosumi, 9. VI. 1963, K. Morimoto (1, KUM), JHL_AREV_459 ; • Mt. Kiyosumi, 14. VI. 1992, T. Ito (1, CMIC), CBM ZI 63020 ; Kagoshima Prefecture: • Yuwan, Amami-Oshima, 31. VII. 1963, Y. Hirashima (1, KUM), JHL_AREV_457 ; • Mt. Yuwan, Amami-Oshima, 30. VII. 1963, L. Gressitt (1 dissected male, KUM), JHL_ACITAI_070 ; • Ishihara-rindo, 9–11. VI. 1978, R. Noda (1, KUM), JHL_AREV_458 ; • Mikyo, Tokunoshima, 27. VII. 1963, Y. Hirashima (1, KUM), JHL_AREV_453 ; • Nishinakama, Amami-Oshima, 2. VI. 1970, H. Makihara, (1, KUM), JHL_AREV_776 ; Okinawa Prefecture: • Shirahama, Iriomote Island, 31. VIII – 5. IX. 1969, H. Makihara (1, KUM), JHL_AREV_452 ; South Korea • Kwangnung, Pochon-gun, Kyonggi-do, 16–19. VII. 1992, K. Morimoto (1, KUM), JHL_AREV_419 ; Taiwan • Wulai, 27. V. 1965, K. Morimoto (2, KUM), JHL_AREV_455, JHL_AREV_456 .</p><p>Diagnosis.</p><p>Body length 3.0– 3.8 mm. Covered in dark and pale brown, white, and black scales, with a white-scaled, V-shaped band in the center of the elytron. Pronotum lacking distinct longitudinal, impunctate midline (present in A. koguma). Scutellum covered in pale scales. Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres emarginate. Pedon tube-shaped with lateral edges approx. parallel in dorsal view, but sinuating evenly into a blunt, rounded point. Internal sac with a small sclerotized structure posteriorly, and two prong-like structures anteriorly at its apex (Fig. 6 P 1 – 2).</p><p>Distribution.</p><p>This species occurs in Japan (mainland, Ryukyu Islands south to Amami-Oshima), South Korea, and Taiwan.</p></div>	https://treatment.plazi.org/id/F6287459457D5125A8A12D3A16A2CFE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
B87BD5220EA3561BA8C43A55BE18EDB3.text	B87BD5220EA3561BA8C43A55BE18EDB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis shigematsui Morimoto & Miyakawa 1995	<div><p>Acicnemis shigematsui Morimoto &amp; Miyakawa, 1995</p><p>Figs 3 F, 6 Q 1–6, 2, 28</p><p>Type material examined</p><p>(Japan: 9). Holotype. Japan: Okinawa Prefecture: • Ishigaki Island, Omoto-dake, 1. VIII. 1989, K. Shigematsu, bears red label reading “ (HOLOTYPE) Acicnemis shigematsui Morimoto et. Miyakawa, 1994 ”, JHL_ACITAI_052, ELKU, ELKU 2960 . Paratypes. Okinawa Prefecture: • Iriomote Island, Shirahama, 31. VIII – 5. IX. 1969, H. Makihara (2, KUM), JHL_ACITAI_005 (dissected male), JHL_ACITAI_047 ; • Iriomote Island, Mt. Ushiku, 4. XI. 1963, G. A. Samuelson (1, KUM), JHL_ACITAI_053 ; • Iriomote Island, Upper Nakara River, 12. III. 1964, Y. Miyatake (1, KUM), JHL_ACITAI_045 .</p><p>Non-type material.</p><p>Japan: Okinawa Prefecture: • Ishigaki Island, Yarabu-dake, 3. V. 1998, K. Takahashi (1, KUM), JHL_ACITAI_054 ; • Omoto-dake, 1–4. VII. 2009, R. Ito (1, KUM), JHL_AREV_481 ; • Omoto-dake, 29. III. 2022, T. Nozaki (1, KUM), JHL_AREV_482 ; • Okinawa Island, Naha, Shuri, “ Sumiyoshi Park ” (probably referring to Sueyoshi Park), 23. XI. 1990, K. Morimoto (1, KUM), JHL_ACITAI_044 .</p><p>Diagnosis.</p><p>Body length 5.1–7.0 mm. Covered in pale brown, dark gray, and black scales, with a tan-colored, V-shaped, transverse band across middle of elytra. Pronotum lacking distinct longitudinal, impunctate midline (present in A. koguma). Scutellum covered in pale scales (Fig. 3 F). Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges approx. parallel in dorsal view, but weakly sinuating into a blunt, subtriangular point. Internal sac not visible inside aedeagus, but with a protruding basal structure extending to the end of the temo and tuberculate (roughened) musculature anteriorly at the apex (Fig. 6 Q 1 – 2).</p><p>Distribution.</p><p>This species is endemic to Okinawa Prefecture, Japan, and is currently only known from Ishigaki, Iriomote, and apparently also Okinawa Island.</p><p>Remarks.</p><p>This species is quite similar in size and color to A. palliata; however, it was differentiated by Morimoto and Miyakawa (1995) on the basis of elytral color differences alone. To verify the validity of this species the type series of A. shigematsui and A. palliata were compared, and several significant differences in external and genital morphology were discovered which are all newly reported in the dichotomous key below. Furthermore, although A. shigematsui was previously only known from Ishigaki and Iriomote Islands, a specimen deposited in the KUM was collected on Okinawa Island (400 km away from the type localities). No specimens of A. shigematsui have since been collected on the island. Further field studies are required to ascertain whether this rarely collected species is truly established on the island.</p></div>	https://treatment.plazi.org/id/B87BD5220EA3561BA8C43A55BE18EDB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
DE46B977D1385A19A1AF1AE7EC20886C.text	DE46B977D1385A19A1AF1AE7EC20886C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis squamata Lewis & Kojima 2025	<div><p>Acicnemis squamata Lewis &amp; Kojima sp. nov.</p><p>Figs 1, 6S 1–6, 2, 30</p><p>Type material</p><p>(Japan: 46). Holotype. Japan: Tokushima Prefecture: • Mt. Tsurugi, Awa, 2. VI. 1957, S. Ueno, JHL_ACITAI_068, dissected male, deposited in KUM . Paratypes. Japan: Aomori Prefecture: • Oyama-nagane, Nakadomari-machi, Kitatsugaru-gun, 7. VI. 2010, K. Morimoto (1, KUM), JHL_AREV_702 ; Fukui Prefecture: • Kinome-toge, 25. VI. 1982, K. Morimoto (1, KUM), JHL_ACITAI_065 ; Fukushima Prefecture: • Yunohana, south Aizu, 8. VIII. 1949, K. Nagayama (1, HUM), JHL_AREV_474 ; • Yunohana, south Aizu, 7. VI. 1981, Y. Kurosawa (1, NMNS), JHL_AREV_682 ; • Yunohana, south Aizu, 20. VII. 1974, Y. Kurosawa (2, NMNS), JHL_AREV_683 ; • Masuzawa Valley, Tateiwa-mura, 6. VI. 2005, K. Morimoto (1, KUM), JHL_AREV_698 ; • Odayama Mt., Monden V., Kita-Aizu, 22. VI. 1948, Y. Kurosawa (1 dissected male, NMNS), JHL_AREV_107 ; Gifu Prefecture: • unknown locality, 24. V. 1949 (1, NHMB), JHL_ACITAI_077; • Hinaga, 22. VII. 1946 (1, NHMB), JHL_ACITAI_078 ; • Hida, 1. VI. 1957, T. Nakane (1, HUM), JHL_AREV_472 ; • Kawai-Idani, Hida, 1–2. VI. 1957, T. Nakane (3, HUM), JHL_AREV_476 – JHL_AREV_478 ; Gunma Prefecture: • Osawa, Katashina, 5. VI. 1988, T. Shinbori (1, KPMNH), JHL_AREV_684 ; • Numata, 30. V. 1953, T. Takei (1, ZMH), ZMH 837203 ; • Hokkaido: Sapporo, “ Takano ” &amp; H. Kono (2, HUM), JHL_AREV_462, JHL_AREV_465 ; • Rausu Shiretoko, 8. VII. 1958, T. Nakane (2, HUM), JHL_AREV_475 ; • Chitose, 1. VIII. 2011, J. Aoki (1, KUM), JHL_AREV_699 ; Ibaraki Prefecture: • Mt. Tsukuba, 11. VIII. 1989, M. J. Sharkey (2, CMNC), JHL_AREV_3000, JHL_AREV_3001 ; Ishikawa Prefecture: • Yamanaka, Kaga, 20. VII. 1910, T. Otsuka (1, HUM), JHL_AREV_463 ; Iwate Prefecture: • Shizukuishi, 7. VI. 2010, Aoki Junichi (1, KUM), JHL_ACITAI_072 ; Kanagawa Prefecture: • Kuno, Odawara, 8. VI. 2014, K. Watanabe (3, OIST), JHL_AREV_687 – JHL_AREV_689 ; • Mt. Ono, Yamakita, 10. V. 1988, N. Niwa, (1, KUM), JHL_AREV_771 ; • Kochi Prefecture: • Engyoji, 22. VII. 1952, Y. Wada (1, KUM), JHL_ACITAI_066 ; • Kumamoto Prefecture: • Mt. Hakucho, 14. V. 1983, M. Chujo (1, KUM), JHL_ACITAI_063 ; • Kyoto Prefecture: • Miyama, Kyoto University Research Forest, 31. V. 2010, Junichi Aoki (1, KUM), JHL_ACITAI_069 (GenBank accession number: PV 255643) ; Nagano Prefecture: • Kaida Highlands, Kiso, 5. IX. 2009, J. Aoki (1, KUM), JHL_AREV_696 ; • “ Noziri Hotel ”, 29. V. 1941, T. Nakane (1, HUM), JHL_AREV_471 ; Nara Prefecture: • Mt. Omine, 23. V. 1974, T. Hattori (1, KUM), JHL_AREV_467 ; Niigata Prefecture: • Mt. Yakemine, 3. VII. 1949, H. Koike (1, KUM), JHL_ACITAI_061 ; • Kurokawa, 28. V. 1955, K. Baba (1, ZMH), ZMH 837202 ; Okayama Prefecture: • Tsuki, Soja, 18. IV. 1976, T. Aono (1, HUM), JHL_AREV_469 ; Osaka Prefecture: • Minoh, 30. V. 1940, K. Taniguchi (1, HUM), JHL_AREV_470 ; Shizuoka Prefecture: • Kakegawa, Ijiri, 3. V. 2010, J. Aoki (1, KUM), JHL_ACITAI_073 ; • Kawanehon-machi, 28. VIII. 2009, J. Aoki (1, KUM), JHL_AREV_697 ; Tokushima Prefecture: • Mt. Tsurugi, 27. VII. 1961, M. T. Chujo (1, KUM), JHL_ACITAI_067 ; Tochigi Prefecture: • Nikko-shi, Chugushi, Ryuzuno-taki Fall, 19. VI. 2014, S. Yamamoto (1, KUM), JHL_ACITAI_076 (GenBank accession number: PV 255644) ; • Ashio, Funeishi-toge, 2. VII. 2006, K. Takahashi (1, KUM), JHL_AREV_701 ; • Tokyo Metropolis: Niijima Island, Mt. Mukoyama, 15. V. 1979, H. Fujita (1, KUM), JHL_ACITAI_071 ; • Kogezawa, “ 1954.5. 10 ” (1 dissected male, KUM), JHL_ACITAI_009 ; • Tokura, 6. VIII. 1950, S. Ueno (2, HUM), JHL_AREV_468 ; • Mt. Takao, 14. IV. 1985, A. Tanaka (1 dissected male, HUM), JHL_AREV_106 ; Wakayama Prefecture: • Mt. Kohya, VII. 1949, M. Hayashi (1, HUM), JHL_AREV_473 ; Yamagata Prefecture: • Yonezawa, 22. VI. 1948, Y. Kurosawa (1, HUM), JHL_AREV_464 ; • Naderayama, 23. V. 1943, Y. Kurosawa (1, NMNS), NSMT-I-C 07303 ; • Seki, near Yonezawa, 2. VII. 1944, Y. Kurosawa (1, NMNS), NSMT-I-C 07304 ; Yamanashi Prefecture: • Masutomi, 13–14. VI. 1970, H. Takizawa (1, HUM), JHL_AREV_466 ; • Masutomi, “ 10. VI. 2603 ” (1, NMNS), NSMT-I-C 07305 ; • Akeno, 6. VI. 2010, T. Shinbori (2, KPMNH), JHL_AREV_685, JHL_AREV_686 ; • Kami-yoshida, Fuji-yoshida city, 15. VII. 2024, J. H. Lewis (1, OIST), JHL_AREV_986 .</p><p>Diagnosis.</p><p>Body length 3.0– 4.4 mm. Covered in pale yellow, white, gray, and black scales. Pronotum with central longitudinal band of dark gray to black scales, bordered laterally by pale brown scales. Elytra with first intervals alternating black (at base of scutellum) and yellow; second to sixth intervals gray in posterior 1 / 2 of elytra, humeri with yellow to pale scales; apical 1 / 2 of elytra with transverse white V-shaped band extending from first to fifth intervals; apex of elytra largely with pale to yellow scales. Pronotum lacking impunctate longitudinal midline (present in A. koguma). Scutellum covered in bright yellow to dull gray scales. Only odd elytral intervals with erect scales. Sclerolepidia numerous, dense, and nub-like, protruding laterally outward. Third tarsomeres emarginate. Apex of pedon with distinct sinuation and pointed (Fig. 6 S 1 – 2). Internal sac of aedeagus distinctly scaly in appearance, protruding, and not extending to midpoint of temo; anterior apex of internal sac ending in two scaly feather-like tips (Fig. 6 S 1 – 2).</p><p>Distribution.</p><p>This species is only known from mainland Japan, from Hokkaido south to Kyushu (Kumamoto Prefecture).</p><p>Etymology.</p><p>The specific name squamata is a reference to the diagnostic internal sac of the aedeagus which bears scaly patterning similar to snakeskin.</p><p>Remarks.</p><p>See A. palliata Remarks section for taxonomic history. Acicnemis squamata differs from A. palliata (= A. dorsonigrita) in size ( A. squamata is 3.0– 4.4 mm in length whereas A. palliata is 6.1–7.0 mm in size), genitalia ( A. squamata with internal sac of aedeagus scaly in appearance, protruding, and ending in two scaly featherlike tips; A. palliata without scaly internal sac, and with two protruding hook-like structures curved inwards at 90 °) and scutellum scaling ( A. squamata with scutellum covered in scales; A. palliata with scutellum prominently bare along the dorsal surface). See also the dichotomous key below.</p></div>	https://treatment.plazi.org/id/DE46B977D1385A19A1AF1AE7EC20886C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
CDF7E3820987565CA9374527DCFB4285.text	CDF7E3820987565CA9374527DCFB4285.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis suturalis Roelofs 1875	<div><p>Acicnemis suturalis Roelofs, 1875</p><p>Figs 6 T 1–6, 2, 31</p><p>Type material examined</p><p>(Japan: 24). Holotype. Japan: Hyogo Prefecture: • Kobe, “ 1910–320 ”, G. Lewis, NHMUK 015026634, bears a circular, white and red label reading “ Type H. T. ” .</p><p>Non-type material.</p><p>Japan: Aomori Prefecture: • Nei, Misawa-shi, 8. VI. 2010, K. Morimoto (1, KUM), JHL_AREV_507 ; Chiba Prefecture: • Nagara, Chosei-gun, 12. VI. 1988, M. Satoshi (2, CMIC), CBM ZI 159815, 159816 ; Gunma Prefecture: • Minakami, 6–7. VI. 2008, K. Morimoto (6, KUM), JHL_AREV_504, JHL_AREV_505 ; Ehime Prefecture: • Mt. Takanawa, Matsuyama-shi, 11–12. VIII. 2015, K. Kuroda (1, KUM), JHL_AREV_506 ; Fukuoka Prefecture: • Mt. Wakasugi ( Chikuzen), 23. V. 1954, K. Morimoto, JHL_AREV_422, JHL_AREV_423, JHL_ACITAI_092 (dissected male) (3, KUM) ; • Mt. Hiko, 15. VII. 1955, H. Kamiya, JHL_AREV_424 ; Hyogo Prefecture: • Kobe, Mt. Maya, 3. IX. 1963, K. Morimoto, JHL_AREV_421 (1, KUM) ; Kanagawa Prefecture: • Mimasutoge, 6. V. 1990, H. Takizawa (1, HUM), JHL_AREV_632 ; • Hatano, 7. VI. 1983, H. Takizawa (1, HUM), JHL_AREV_633 ; Kyoto Prefecture: • Katsura-gawa, 12. VI. 1918, M. Suzuki, JHL_AREV_425 (1, KUM) ; • Kyoto University Ashu Forest, Miyama-cho, Tanann City, 23. V. 2009, K. Morimoto (1, KUM), JHL_AREV_503 ; Tochigi Prefecture: • Nanma, Kanuma, 4. V. 1993, H. Takizawa (1, HUM), JHL_AREV_631 ; Niigata Prefecture: • Kamikawa-cho, 3. VI. 2000, K. Takahashi (2, KUM), JHL_AREV_508 .</p><p>Diagnosis.</p><p>Body length 4.0– 4.7 mm. Covered in pale brown, dark and pale gray, pale yellow, and black scales, with two small white spots on the elytra (one on each side) in the apical 1 / 2 of the elytra. Pronotum lacking impunctate longitudinal carina (present in A. koguma). Scutellum covered in pale scales. Elytra with erect scales only on odd intervals. Sclerolepidia distinct and protruding. Third tarsomeres distinctly emarginate. Pedon with lateral edges converging in dorsal view, sinuating into an elongate, tapered tip. Internal sac protruding from aedeagus, and with two cylinder-shaped sclerotized structures anteriorly at the apex (Fig. 6 T 1 – 2).</p><p>Distribution.</p><p>This distinctive species occurs in Japan (mainland), Taiwan, and has also been reported from South Korea and the Kuril Islands (Russia) (Park et al. 2009; Alonso-Zarazaga et al. 2023).</p></div>	https://treatment.plazi.org/id/CDF7E3820987565CA9374527DCFB4285	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
F14219CE481D553D900D14BA7FAE6024.text	F14219CE481D553D900D14BA7FAE6024.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acicnemis undefined-1	<div><p>Acicnemis sp. 1</p><p>Figs 6 R 1–6, 2, 29</p><p>Material examined</p><p>(Myanmar: 2; Taiwan: 7; Thailand: 3; Vietnam: 2; Total: 14). Myanmar • Tamu Village, Kachin State, 7. VII. 2003, A. Abe (2, KUM), JHL_SYN_169 (dissected male), JHL_AREV_717 ; Taiwan • “ Formosa ”, T. Kano (1, NMNS), JHL_AREV_681 ; • His-Leng, 5. V. 1981, S. Tsuyuki (5, TUA), JHL_SYN_175 (dissected male), JHL_SYN_176 (dissected male), JHL_AREV_714 – JHL_AREV_716 ; • Lushan Wenchuan, Nantou Hsien, 6. VI. 1976, H. Makihara (1, KUM), JHL_AREV_719 ; Thailand • Pupin-Meo, 11. X. 1993, T. Senoh (1 dissected male, KUM), JHL_SYN_170 ; • Chiang Rai, Wiang Pa Pao, 12. XI. 2012, K. Takahashi (2, KUM), JHL_AREV_718 (EGP 0160 - H 9), JHL_AREV_722 ; • Doi Suthep, 23. VI. 1993, T. Senoh, (1, KUM), JHL_AREV_766 ; Vietnam • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.86583&amp;materialsCitation.latitude=22.538889" title="Search Plazi for locations around (long 105.86583/lat 22.538889)">Phia Oac National Park</a> near Phia Den, Cao Bang Province, 22°32'20"N, 105°51'57"E, 19–22. III. 2012, S. D. Gaimari, M. Hauser, H. T. Pham (2, CMNC), JHL_AREV_703, JHL_AREV_704 ; • “ IIea ” Binh, Tonkin, bears red label reading “ simpliciata Typus H. ”, bears brown label reading “ A. simpliciata m ♂ Heller det. 1930 ” (1, SNSD), JHL_ACITAI_093 .</p><p>Diagnosis.</p><p>Body length 4.1–5.0 mm. Covered in pale and dark brown scales, with bright X-shaped band of white scales across apical 1 / 2 of elytra, which extends up to the elytra humeri. Distinctive bright white scale bands also present on hind femora. Pronotum lacking distinct longitudinal, impunctate midline (present in A. koguma). Scutellum small, bordered by pale, yellow scales, and weakly bare dorsally (or covered in scales); compare with A. kiotoensis which possesses a large, broad, bare scutellum. Only odd elytral intervals with broad, erect scales. Sclerolepidia numerous, dense, and nub-like, protruding laterally outward (unlike A. maculaalba, A. kiotoensis, and A. koguma which have indistinct, low, rounded sclerolepidia). Third tarsomeres emarginate (as opposed to truncate, as in A. koguma). Pedon with lateral edges approx. parallel in dorsal view, but sinuating in distal 1 / 2 into a distinct point. Internal sac protruding and clear across posterior 2 / 3; with two bean-shaped and U-shaped sclerotized structure anteriorly at apex (Fig. 6 R 1 – 2).</p><p>Distribution.</p><p>This widely distributed species occurs in Vietnam, Thailand, Myanmar, and Taiwan.</p><p>Remarks.</p><p>One of K. M. Heller’s specimen of this species in the SNSD collection has a holotype label with the name “ A. simpliciata ” written (JHL _ACITAI_093); however, we were unable to find a formal description of this species in any of Heller’s works. Heller often attached type labels to specimens without formally publishing the descriptions (Olaf Jäger – SNSD Collection Manager to JHL, pers. comm. 2025; Katsura Morimoto – KUM Curator to HK pers. comm., 2025). Thus, this species is likely undescribed; however, there are a number of other similar Southeast Asian species for which type material should be compared with Acicnemis sp. 1 before formal description. This falls outside of the scope of the current work (study area); however, we still acknowledge the presence of this species in Taiwan and treat it here as an unnamed species.</p></div>	https://treatment.plazi.org/id/F14219CE481D553D900D14BA7FAE6024	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lewis, Jake H.;Kojima, Hiroaki;Campbell, Erin O.	Lewis, Jake H., Kojima, Hiroaki, Campbell, Erin O. (2025): A Japan-focused review of East Asian Acicnemis Fairmaire, 1849 (Coleoptera, Curculionidae) reveals cryptic diversity in a well-studied region. ZooKeys 1259: 103-167, DOI: 10.3897/zookeys.1259.164450
