taxonID	type	description	language	source
8834D75AAF7252FCB4F9ECA135D771BC.taxon	description	Figs 60 C, 62, 63	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8834D75AAF7252FCB4F9ECA135D771BC.taxon	diagnosis	Diagnosis. Labrum trapezoid; dorsal surface of labrum with distal row multiple; tarsal claw hooked with seven teeth; abdominal tergites predominantly medium to dark brown; central area of upper surface of femora with numerous short, stout, and blunt setae, shorter on hind femur; posterior margin of abdominal segment VII without denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8834D75AAF7252FCB4F9ECA135D771BC.taxon	description	Description. Head. Labrum trapezoid (Fig. 62 A). Ratio labrum width / clypeus width 1.25. Ratio labrum width / insertion width 1.64. Medial emargination shallow with flat denticles. Proximal row of setae present, shifted to distal margin. Number of setae on proximal row ~ 115. Distal row of setae multiple. Median part of labrum in ventral view with scattered thin setae (Fig. 62 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with 11 setae below the mola (Fig. 62 C). Maxillae. Posterior margin of cardo with few stout and medium size setae together with some hair-like setae and two or three stout and long setae in submarginal position. Proximo-lateral margin of stipes with a bunch of ~ 15 medium size and stout setae. Apical-ventral row with 36 pectinate setae. Maxillary palp segment II 0.75 as long as segment I. Maxillary palp segment III 0.63 as long as segment I (Fig. 62 D). Stout setae on outer margin of segment I absent. Segment II inner margin with four stout setae, outer margin without stout setae. Segment III inner margin with four stout setae, dorsal surface with few setae at apex. Segment III 1.73 × as long as base width. Labium. Labial palp segment II 0.92 as long as segment I. Labial palp segment III 0.65 as long as segment I (Fig. 62 E). Segment I inner margin with 11 stout setae, outer margin with 18 stout setae. Segment II outer margin with ten stout setae. Segment III dorsal face with five stout setae. Segment III 2.77 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 62 F). Thorax. Dorsal margin of femur with a row of very long, stout, pointed setae (Fig. 63 A); an irregular submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin, more regular on hind femur; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae, except on hind leg where the submarginal row is regular (Fig. 63 B); central area of upper surface with numerous short, stout, blunt setae. Fore tibia with two or three rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, outer margin with numerous hair-like setae; hind tibia with two or three rows of simple long, stout setae on ventral margin, lower surface with few simple, thin setae, outer margin with short and long pointed stout setae and hair-like setae. Tarsal claw hooked with seven denticles progressively larger apically, except distal much smaller (Fig. 63 C). Abdomen. Gill I with both lamellae. Gills II to VII with dorsal and ventral lamellae of the same size. Gill I with ~ 12 filaments on each lamella, gills II – VI with ~ 20 filaments on each lamella (Fig. 63 D), gill VII with ~ ten filaments on each lamella. Filamentous projections on apex. Posterior margin of terga IX with denticles. Posterior margin of terga VIII with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8834D75AAF7252FCB4F9ECA135D771BC.taxon	etymology	Derivatio nominis. The specific name is to acknowledge the Asaro tribe or “ Mudmen ” and their living traditions, who inhabit close to where the specimens were collected.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2AB4154D4F4354D9B6B4D837701847FB.taxon	description	Figs 34 K, 60 A, 65, 66	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2AB4154D4F4354D9B6B4D837701847FB.taxon	diagnosis	Diagnosis. Labrum cordiform; median part of labrum in ventral view with a row of stout and short setae; abdominal tergites predominantly orange with dark lateral marks; gills II – VI with ~ 15 filaments on each lamella; posterior margin of abdominal segment VII with minute denticles; posterolateral expansions present on abdominal segments V – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2AB4154D4F4354D9B6B4D837701847FB.taxon	description	Description. Head. Labrum cordiform (Fig. 65 A). Ratio labrum width / clypeus width 0.96. Ratio labrum width / insertion width 1.24 – 1.30. Medial emargination narrow with pointed denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 82 – 93. Distal row of setae multiple. Median part of labrum in ventral view with stout setae in row (Fig. 65 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 15 – 22 setae below the mola (Fig. 65 C). Maxillae. Posterior margin of cardo with many hair-like and long setae, together with two or three long and stout setae in submarginal position. Proximo-lateral margin of stipes with a bunch of ten long hair-like setae (Fig. 34 K). Apical-ventral row with 14 – 15 pectinate setae. Maxillary palp segment II 0.80 – 0.81 as long as segment I. Maxillary palp segment III 0.57 – 0.65 as long as segment I (Fig. 65 D). Stout setae on outer margin of segment I absent. Segment II inner margin with 1 – 3 stout setae, outer margin without stout setae. Segment III inner margin with four stout setae, dorsal surface half covered with setae. Segment III 1.53 – 1.56 × as long as base width. Labium. Labial palp segment II 0.88 as long as segment I. Labial palp segment III 0.76 – 0.80 as long as segment I (Fig. 65 E). Segment I inner margin with 25 – 35 stout setae, outer margin with 28 – 30 stout setae. Segment II outer margin with 12 stout setae. Segment III dorsal face with six or seven stout setae. Segment III 2.53 – 2.71 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 65 F). Thorax. Dorsal margin of femur with a row of very long, thin, pointed setae; irregular submarginal rows of pointed setae as long as or a little bit shorter than those on the dorsal margin, these setae decrease in size from the fore- to the hind femur (Fig. 66 A – B); ventral margin with short, stout setae and irregular submarginal rows of long, pointed stout setae, except on hind leg where those on ventral margin are shorter; central area of upper surface with numerous long, stout, pointed setae, shorter on hind femur. Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with two rows of long, simple, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with several rows of simple and feathered long, stout setae on ventral margin, lower surface covered with simple and feathered setae, outer margin with short and long pointed stout setae and hair-like setae. Tarsal claw hooked with eight or nine denticles progressively larger apically (Fig. 66 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size. Gill I with ~ 10 filaments on each lamella, gills II – VI with ~ 15 filaments on each lamella, gill VII minute with 5 – 7 filaments on each lamella. Filamentous projections on apex. Posterior margin of terga IX with denticles. Posterior margin of terga VII – VIII with minute denticles. Posterior margin of terga V – VI without denticles. Posterolateral projections present on segments V – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2AB4154D4F4354D9B6B4D837701847FB.taxon	etymology	Derivatio nominis. This species is dedicated to Dr Michael Balke (ZSM, Germany), collector of most of the species treated here.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
555CC751CA2D52E9B7464EF76066940C.taxon	description	Figs 34 L, 61 A, 67, 68	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
555CC751CA2D52E9B7464EF76066940C.taxon	diagnosis	Diagnosis. Labrum cordiform; emargination narrow with pointed denticles; tarsal claw straight with nine or ten teeth; abdominal tergites predominantly medium to dark brown; posterolateral expansions present on abdominal segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
555CC751CA2D52E9B7464EF76066940C.taxon	description	Description. Head. Labrum cordiform (Fig. 67 A). Ratio labrum width / clypeus width 1.06 – 1.08. Ratio labrum width / insertion width 1.28. Medial emargination narrow with pointed denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 120 – 140. Distal row of setae multiple. Median part of labrum in ventral view with scattered thin setae (Fig. 67 B). Mandibles. Outer margin with row of setae on entire margin. Ending of the row near incisor. Right mandible with 14 – 16 setae below the mola (Fig. 67 C). Maxillae. Posterior margin of cardo with many hair-like setae. Proximo-lateral margin of stipes with a bunch of ~ 15 stout, long, pointed setae (Fig. 34 L). Apical-ventral row with 22 – 24 pectinate setae. Maxillary palp segment II 0.82 – 0.88 as long as segment I. Maxillary palp segment III 0.75 – 0.85 as long as segment I (Fig. 67 D). Stout setae on outer margin of segment I present. Segment II inner margin with 1 – 5 stout setae, outer margin with 2 – 7 stout setae. Segment III inner margin with four or five stout setae, dorsal surface fully covered with setae. Segment III 1.67 – 1.77 × as long as base width. Labium. Labial palp segment II 0.77 – 0.83 as long as segment I. Labial palp segment III 0.60 – 0.62 as long as segment I (Fig. 67 E). Segment I inner margin with 23 – 25 stout setae, outer margin with 33 – 38 stout setae. Segment II outer margin with 21 – 22 stout setae. Segment III dorsal face with three or four stout setae. Segment III 2.56 – 2.67 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 67 F). Thorax. Dorsal margin of femur with a row of very long, thin, pointed setae; irregular submarginal rows of pointed setae as long as or a little bit shorter than those on the dorsal margin, ventral margin with short, stout setae and submarginal rows of long, pointed stout setae, except on hind leg where those on ventral margin are shorter; central area of upper surface with numerous long, stout, pointed setae, shorter on hind femur (Fig. 68 A – C). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with several rows of long, simple, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with several rows of simple and feathered long, stout setae on ventral margin, lower surface covered with simple and feathered setae, outer margin with short and long pointed stout setae and hair-like setae. Tarsal claw hooked with nine or ten denticles progressively larger, except distal much smaller (Fig. 68 D). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size. Gill I with nine or ten filaments on each lamella (Fig. 68 J), gills II – VI with ~ 15 filaments on each lamella (Fig. 68 L), gill VII with three or four filaments on each lamella (Fig. 68 K). Filamentous projections on apex. Posterior margin of terga VII – IX with denticles (Fig. 68 G – I). Posterior margin of terga V – VI with minute denticles (Fig. 68 E – F). Posterolateral projections present on segment VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
555CC751CA2D52E9B7464EF76066940C.taxon	etymology	Derivatio nominis. The specific name is to recognise the tribe Baruya living in the valleys of Wonemara and Marawaka, where some of the specimens come from, and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9C67D26A68C35566A388366F02EE5DE4.taxon	description	Figs 34 D, 60 B, 69, 70	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9C67D26A68C35566A388366F02EE5DE4.taxon	diagnosis	Diagnosis. Labrum trapezoid; median part of labrum in ventral view with scattered thin and long setae; abdominal tergites predominantly orange with dark lateral marks; posterior margin of abdominal segment VII without denticles; posterolateral expansions present on abdominal segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9C67D26A68C35566A388366F02EE5DE4.taxon	description	Description. Head. Labrum trapezoid (Fig. 69 A). Ratio labrum width / clypeus width 1.26 – 1.27. Ratio labrum width / insertion width 1.69 – 1.71. Medial emargination shallow with flat denticles. Proximal row of setae simple, shifted to distal margin. Number of setae on proximal row ~ 123 – 130. Distal row of setae multiple. Median part of labrum in ventral view with scattered thin setae (Fig. 69 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with 8 – 13 setae below the mola (Fig. 69 C). Maxillae. Posterior margin of cardo with few stout and medium size setae (Fig. 34 D). Proximo-lateral margin of stipes with a bunch of eight long and stout setae. Apical-ventral row with 34 – 44 pectinate setae. Maxillary palp segment II 0.77 – 0.86 as long as segment I. Maxillary palp segment III 0.77 – 0.86 as long as segment I (Fig. 69 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two stout setae, outer margin without or with up to two stout setae. Segment III inner margin with six stout setae, dorsal surface with few setae at apex. Segment III 1.67 – 2.00 × as long as base width. Labium. Labial palp segment II 0.84 – 0.87 as long as segment I. Labial palp segment III 0.60 – 0.64 as long as segment I (Fig. 69 E). Segment I inner margin with 8 – 10 stout setae. Segment I outer margin with 12 – 16 stout setae. Segment II outer margin with 6 – 10 stout setae. Segment III dorsal face without stout setae. Segment III 2.56 – 3.33 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 69 F). Thorax. Dorsal margin of fore femur with few stout, short, pointed setae, and a double submarginal row of long, stout, pointed setae (Fig. 70 A); dorsal margin of mid- and hind femur with a row of long, thin, pointed setae; a submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin (Fig. 70 B); ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae, except on hind leg where those on ventral margin are shorter; central area of upper surface with numerous short, stout, blunt setae. Fore tibia with two or three rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with one row of short, simple, pointed setae, and a submarginal row of long, simple, pointed stout setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with a row of simple long, stout setae on ventral margin, outer margin with short pointed stout setae and hair-like setae. Tarsal claw hooked with 6 – 8 denticles progressively larger apically, followed by two tiny protuberances (Fig. 70 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size. Gill I with ~ 10 filaments on each lamella, gills II – VI with ~ 15 filaments on each lamella (Fig. 70 F), gill VII minute with 5 – 7 filaments on each lamella. Filamentous projections on apex. Posterior margin of terga IX with minute denticles, or with larger denticles (Fig. 70 D). Posterior margin of terga VIII with minute denticles (Fig. 70 E). Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9C67D26A68C35566A388366F02EE5DE4.taxon	etymology	Derivatio nominis. The specific name is to acknowledge the Kalam tribe of the remote area of Simbai, where the specimens described here come from.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B808E33780E5539CB1EC90D3B0EA8C7F.taxon	description	Figs 34 J, 61 C, 71, 72	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B808E33780E5539CB1EC90D3B0EA8C7F.taxon	diagnosis	Diagnosis. Labrum trapezoid; dorsal surface of labrum with distal row simple; tarsal claw hooked with seven teeth; abdominal tergites predominantly medium to dark brown; central area of upper surface of femora with numerous long, stout, and blunt setae, shorter on hind femur; posterior margin of abdominal segment VII with minute denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B808E33780E5539CB1EC90D3B0EA8C7F.taxon	description	Description. Head. Labrum trapezoid (Fig. 71 A). Ratio labrum width / clypeus width 1.28. Ratio labrum width / insertion width 1.71. Medial emargination shallow with flat denticles. Proximal row of setae simple, shifted to anterior margin. Number of setae on proximal row ~ 120. Distal row of setae multiple, shifted to anterior margin. Median part of labrum in ventral view with scattered thin setae (Fig. 71 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with 5 setae below the mola (Fig. 71 C). Maxillae. Posterior margin of cardo with ~ 10 stout pointed and long setae. Proximo-lateral margin of stipes with a bunch of ~ 12 medium size and short stout setae (Fig. 34 J). Apical-ventral row with 45 pectinate setae. Maxillary palp segment II 0.86 as long as segment I. Maxillary palp segment III 0.67 as long as segment I (Fig. 71 D). Stout setae on outer margin of segment I absent. Segment II inner margin without stout setae. Segment II outer margin without stout setae. Segment III inner margin with six stout setae, dorsal surface with few setae at apex. Segment III 1.87 × as long as base width. Labium. Labial palp segment II 0.75 as long as segment I. Labial palp segment III 0.59 as long as segment I (Fig. 71 E). Segment I inner margin with ten stout setae. Segment I outer margin with 17 stout setae. Segment II outer margin with ten stout setae. Segment III dorsal face with two stout setae. Segment III 2.71 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 71 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; one submarginal row of pointed setae as long as those on the dorsal margin (Fig. 72 A), ventral margin with short, stout setae and a submarginal row of pointed stout setae, except on hind leg where those on ventral margin are shorter; central area of upper surface with numerous long, stout, blunt setae, shorter on hind femur. Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with a row of short, pointed setae and a submarginal row of long, pointed stout setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with a row of short, pointed setae and a submarginal row of long, pointed simple stout setae on ventral margin, lower surface covered with simple, thin setae, outer margin with short and long pointed stout setae and numerous hair-like setae. Tarsal claw hooked with seven subequal denticles, except median much larger (Fig. 72 B – C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size, or with ventral lamella smaller. Gill I with ~ 15 filaments on each lamella, gills II – VI with ~ 15 filaments on upper lamella, and ~ 10 on lower lamella, gill VII with ~ 15 filaments on each lamella. Filamentous projections on apex, or along entire margin. Posterior margin of terga IX with denticles. Posterior margin of terga VII – VIII with minute denticles. Posterior margin of terga V – VI without denticles. Posterolateral projections present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B808E33780E5539CB1EC90D3B0EA8C7F.taxon	etymology	Derivatio nominis. This species is dedicated to Pauline Sartori, daughter of the first author.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
DF5F44A602E75406A87B76F073855BFF.taxon	description	Figs 61 B, 64 E, F, 73, 74	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
DF5F44A602E75406A87B76F073855BFF.taxon	diagnosis	Diagnosis. Labrum cordiform; median part of labrum in ventral view with a row of stout and short setae; abdominal tergites predominantly orange with dark lateral marks; gills II – VI with ~ 25 filaments on each lamella; posterior margin of abdominal segment VII with minute denticles; posterolateral expansions present on abdominal segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
DF5F44A602E75406A87B76F073855BFF.taxon	description	Description. Head. Labrum cordiform (Fig. 73 A). Ratio labrum width / clypeus width 0.96 – 1.02. Ratio labrum width / insertion width 1.19 – 1.26. Medial emargination narrow with pointed denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 88 – 96. Distal row of setae multiple. Median part of labrum in ventral view with stout setae in row (Fig. 73 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 12 – 15 setae below the mola (Fig. 73 C). Maxillae. Posterior margin of cardo with many long, stout and pointed setae together with some hair-like setae. Proximo-lateral margin of stipes with a bunch of ~ 10 long hair-like setae. Apical-ventral row with 12 – 15 pectinate setae. Maxillary palp segment II 0.86 – 1.00 as long as segment I (Fig. 73 D). Maxillary palp segment III 0.67 – 0.78 as long as segment I. Stout setae on outer margin of segment I absent. Segment II inner margin with one or two stout setae, outer margin with 3 – 5 stout setae. Segment III inner margin with 3 – 5 stout setae, dorsal surface half covered with setae. Segment III 1.53 – 2.40 × as long as base width. Labium. Labial palp segment II 0.75 – 0.85 as long as segment I. Labial palp segment III 0.67 – 0.83 as long as segment I (Fig. 73 E). Segment I inner margin with 26 – 30 stout setae, outer margin with 24 – 30 stout setae. Segment II outer margin with 12 – 22 stout setae. Segment III dorsal face with 5 – 10 stout setae. Segment III 2.56 – 3.00 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 73 F). Thorax. Dorsal margin of femur with a row of very long, thin, pointed setae; irregular submarginal rows of pointed setae as long as or a little bit shorter than those on the dorsal margin, ventral margin with short, stout setae and submarginal rows of long, pointed stout setae, except on hind leg where those on ventral margin are shorter (Fig. 74 A – B); central area of upper surface with numerous long, stout, pointed setae, shorter on hind femur. Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with several rows of long, simple, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with several rows of simple and feathered long, stout setae on ventral margin, lower surface covered with simple and feathered setae, outer margin with short and long pointed stout setae and hair-like setae. Tarsal claw hooked, with eight to ten denticles progressively larger apically (Fig. 74 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size. Gill I with ~ 10 filaments on each lamella (Fig. 74 G), gills II – VI with 20 – 25 filaments on each lamella (Fig. 74 H), gill VII with 12 – 15 filaments on each lamella. Filamentous projections on apex, or along entire margin. Posterior margin of terga IX with denticles (Fig. 74 D). Posterior margin of terga VIII with minute denticles, or with larger denticles (Fig. 74 E). Posterior margin of terga VII with minute denticles (Fig. 74 F). Posterior margin of terga V – VI without denticles. Posterolateral projections present on segment VI – IX. Eggs. General shape elongated. Size 205 – 210 μm × 90 – 95 μm. Chorionic surface smooth with longitudinal ridges well marked having the appearance of being segmented. Attachment structures at each pole composed of four anchors with long stem, reaching the middle of the egg (Fig. 64 E – F). Micropyle not visible.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
DF5F44A602E75406A87B76F073855BFF.taxon	etymology	Derivatio nominis. This species is dedicated to Sabrina Perret Sartori, daughter in law of the first author.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2A3E61887BB85BBEB47E6423FE6BC107.taxon	diagnosis	Diagnosis. Gill I similar in shape to the following ones. Ventral lamella of gills smaller but not highly reduced, with ≥ 9 processes; claws with 6 – 10 teeth; posterior margin of abdominal segment VIII with small denticles; posterolateral expansions on abdominal segments VII – IX, VI – IX or even V – IX. Eggs fusiform with hook-like attachment structures at each pole.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2A3E61887BB85BBEB47E6423FE6BC107.taxon	etymology	Derivatio nominis. From the greek κόσμημα meaning jewel and νύμφη meaning nymph, to recognise the peculiar beauty of the species in this genus. The gender is feminine.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2A3E61887BB85BBEB47E6423FE6BC107.taxon	type_taxon	Type species. Kosminympha sabrinae sp. nov. by present designation.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2A3E61887BB85BBEB47E6423FE6BC107.taxon	description	Description. Size: 10.0 – 11.0 mm for male mature nymphs; 10.5 – 12.5 mm for mature female nymphs. Head. Prognathous; labrum cordiform or trapezoid, anterior emargination either shallow with flat denticles or narrow with pointed denticles; proximal row of setae long, with 80 – 140 setae; distal row of setae multiple; median part of labrum in ventral view with scattered thin and long setae or with stout and short setae in a row. Outer margin of mandibles with a row of setae. Proximo-lateral margin of stipes with a bunch of setae, either long and thin or short and stout. Apex of superlingua of hypopharynx either truncate or emarginate. Thorax. Central area of upper face of femora with numerous short or long, pointed, or blunt setae; tarsal claw hooked, with 6 – 10 teeth. Abdomen. Gill I similar in shape to the following ones; gill I composed of dorsal and ventral lamellae, each with 9 – 15 filaments; gills II – VI composed of dorsal and ventral lamellae, each with 15 – 25 filaments; gill VII composed of dorsal and ventral lamellae, each with 5 – 15 filaments, depending on species. Posterior margin of abdominal terga II – VIII without or with minute denticles; posterolateral expansions of the abdomen present on segments V – IX, VI – IX or VII – IX. Egg. Elongated, with attachments structures fixed at each pole, composed of a long branch ending by a double fishhook.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
2A3E61887BB85BBEB47E6423FE6BC107.taxon	distribution	Distribution. The genus Kosminympha is only known from Papua New Guinea and has been found on the eastern part of the island; the genus has yet not been recorded from Indonesia Papua.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7B61E37782015D8EB4CB50B7A852B291.taxon	description	Figs 36 B, 37, 38	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7B61E37782015D8EB4CB50B7A852B291.taxon	diagnosis	Diagnosis. Femora uniformly dark brown, without transverse bands; tarsal claw with 6 – 8 teeth; gill I – II uni-lamellate, without ventral lamella; gills III – VI with 5 – 8 filaments on dorsal lamella and four or five filaments on ventral lamella, gill VII with four or five filaments on dorsal lamella and one or two filaments on ventral lamella.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7B61E37782015D8EB4CB50B7A852B291.taxon	description	Description. Head. Labrum cordiform (Fig. 37 A). Ratio labrum width / clypeus width 0.99 – 1.10. Ratio labrum width / insertion width 1.35 – 1.53. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 45 – 61. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 37 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 6 – 10 setae below the mola (Fig. 37 C). Maxillae. Posterior margin of cardo with few hair-like and stout setae, together with one long and stout seta in submarginal position. Proximo-lateral margin of stipes with a bunch of six stout setae increasing in size distally. Apical-ventral row with 21 – 24 pectinate setae. Maxillary palp segment II 0.83 – 0.91 as long as segment I. Maxillary palp segment III 0.67 – 0.82 as long as segment I (Fig. 37 D). Stout setae on outer margin of segment I absent. Segment II inner margin with 2 – 4 stout setae. Segment II outer margin with 1 – 4 stout setae. Segment III inner margin with 4 – 6 stout setae, dorsal surface with few setae at apex. Segment III 1.83 – 2.00 × as long as base width. Labium. Labial palp segment II 0.72 – 0.80 as long as segment I. Labial palp segment III 0.78 – 0.88 as long as segment I (Fig. 37 E). Segment I inner margin with 10 – 15 stout setae, outer margin with 9 – 12 stout setae. Segment II outer margin with four or five stout setae. Segment III dorsal face with 3 – 5 stout setae. Segment III 3.00 – 3.86 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 37 F). Thorax. Dorsal margin of femur with a row of very long, stout, pointed setae; an irregular submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin on fore femur, more regular on mid- and hind femur; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae; central area of upper surface with few long and short, stout, pointed setae (Fig. 38 A – B). Fore tibia with two rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, outer margin with numerous hair-like setae; hind tibia with two rows of simple and feathered long, stout setae on ventral margin, outer margin with short and long pointed stout setae and numerous hair-like setae. Tarsal claw slightly hooked with 6 – 8 denticles progressively larger distally (Fig. 38 C). Abdomen. Gills I and II with single lamella (Fig. 38 E – F). Gills III – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with three filaments on dorsal lamella, without ventral lamella, gill II with four or five filaments on dorsal lamella, without ventral lamella, gills III – VI with 5 – 8 filaments on dorsal lamella and four or five on ventral lamella (Fig. 38 G), gill VII with four or five filaments on dorsal lamella and one or two filaments on ventral lamella (Fig. 38 H). Posterior margin of terga IX with minute denticles (Fig. 38 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7B61E37782015D8EB4CB50B7A852B291.taxon	etymology	Derivatio nominis. This species is dedicated to Prof. Nadir Alvarez, current director of Naturéum and instigator of the Cozzarolo et al. paper who revealed the diversity of New Guinean Thraulini.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E532969E6CC750D2BDC38D5AE062D693.taxon	description	Figs 35 A, 39, 40	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E532969E6CC750D2BDC38D5AE062D693.taxon	diagnosis	Diagnosis. Labrum cordiform; dorsal face of labrum with proximal and distal rows multiple; femora pale brown with two transverse dark brown bands in middle and at apex; gill I with ventral lamella composed of a single filament; gill II – VI with six or seven filaments on dorsal lamella and two or three on ventral lamella; gill VII with four or five filaments on dorsal lamella and a single one on ventral lamella.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E532969E6CC750D2BDC38D5AE062D693.taxon	description	Description. Head. Labrum cordiform (Fig. 39 A). Ratio labrum width / clypeus width 1.05. Ratio labrum width / insertion width 1.36. Medial emargination narrow with flat denticles. Proximal row of setae present, multiple. Number of setae on proximal row ~ 104. Distal row of setae multiple. Median part of labrum in ventral view with scattered thin setae (Fig. 39 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with seven setae below the mola (Fig. 39 C). Maxillae. Posterior margin of cardo with many long hair-like setae. Proximo-lateral margin of stipes twisted. Apical-ventral row with 23 pectinate setae. Maxillary palp segment II 0.88 as long as segment I. Maxillary palp segment III 0.68 as long as segment I (Fig. 39 D). Stout setae on outer margin of segment I absent. Segment III dorsal surface half covered with setae. Segment III 1.55 × as long as base width. Labium. Labial palp segment II 0.82 as long as segment I. Labial palp segment III 0.71 as long as segment I (Fig. 39 E). Segment I inner margin with nine stout setae, outer margin with 14 stout setae. Segment II outer margin with four stout setae. Segment III dorsal face with four stout setae. Segment III 2.67 × as long as base width. Hypopharynx. Apex of superlingua truncate. Thorax. Dorsal margin of femur with a row of very long, thin, pointed setae; regular submarginal row of pointed setae as long as those on the dorsal margin; ventral margin with relatively long, pointed setae and an irregular submarginal row of long, pointed thin setae (Fig. 40 A – B); central area of upper surface with very long pointed setae, as long as those on the dorsal margin. Fore tibia with two or three rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, outer margin with numerous hair-like setae; hind tibia with a marginal row of simple short, thin setae and a submarginal row of long to very long setae on ventral margin, outer margin with long to very long pointed setae and hair-like setae (Fig. 40 C). Tarsal claw hooked with three or four denticles progressively larger, except distal much larger. Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with three filaments on dorsal lamella, ventral lamella with a single filament (Fig. 40 E); gill II – VI with six or seven filaments on dorsal lamella and two or three on ventral lamella (Fig. 40 F); gill VII with four or five filaments on dorsal lamella and a single one on ventral lamella (Fig. 40 G). Posterior margin of terga IX with minute denticles (Fig. 40 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E532969E6CC750D2BDC38D5AE062D693.taxon	etymology	Derivatio nominis. The specific name is to recognize the Anga people who live in the area where the new species has been found and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
635CB6BF9057584597488555C7AF9BC6.taxon	description	Figs 36 A, 41, 42	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
635CB6BF9057584597488555C7AF9BC6.taxon	diagnosis	Diagnosis. Femora uniformly dark brown, without transverse bands; tarsal claw with 6 – 8 teeth; gill I and II uni-lamellate, without ventral lamella; gills III – VI with five or six filaments on dorsal lamella and a single or two filaments on ventral lamella, gill VII with three filaments on dorsal lamella and a single filament on ventral lamella.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
635CB6BF9057584597488555C7AF9BC6.taxon	description	Description. Head. Labrum cordiform (Fig. 41 A). Ratio labrum width / clypeus width 0.99 – 1.04. Ratio labrum width / insertion width 1.36 – 1.45. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 42 – 69. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 41 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with eight or nine setae below the mola (Fig. 41 C). Maxillae. Posterior margin of cardo with few hair-like setae, together with two stout and long setae in submarginal position. Proximo-lateral margin of stipes with a bunch of five stout setae of irregular length. Apical-ventral row with 22 – 24 pectinate setae. Maxillary palp segment II 0.86 – 1.00 as long as segment I. Maxillary palp segment III 0.64 – 0.95 as long as segment I (Fig. 41 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two or three stout setae. Segment II outer margin without stout setae. Segment III inner margin with four stout setae, dorsal surface with few setae at apex. Segment III 1.67 – 2.00 × as long as base width. Labium. Labial palp segment II 0.70 – 0.80 as long as segment I. Labial palp segment III 0.74 – 0.80 as long as segment I (Fig. 41 E). Segment I inner margin with 13 – 15 stout setae, outer margin with 12 – 15 stout setae. Segment II outer margin with three or four stout setae. Segment III dorsal face with 3 – 5 stout setae. Segment III 2.91 – 3.30 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 41 F). Thorax. Dorsal margin of femur with few long, stout, pointed setae; an irregular submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae; central area of upper surface with numerous long and short, stout, pointed setae (Fig. 42 A). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, dorsal margin with few hair-like setae; hind tibia with two rows of simple and feathered long, stout setae on ventral margin, dorsal margin with short and long pointed stout setae and few hair-like setae. Tarsal claw slightly hooked with seven or eight denticles progressively larger apically (Fig. 42 B). Abdomen. Gill I – II with single lamella. Gills III – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I uni-lamellate, with two or three filaments (Fig. 42 D), gill II uni-lamellate with four filaments (Fig. 42 E), gills III – VI with five or six filaments on dorsal lamella and a single or two filaments on ventral lamella (Fig. 42 F), gill VII with three filaments on dorsal lamella and a single filament on ventral lamella (Fig. 42 G). Posterior margin of terga IX with minute denticles (Fig. 42 C). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
635CB6BF9057584597488555C7AF9BC6.taxon	etymology	Derivatio nominis. This species is dedicated to Prof. Boonsatien Boonsoong (Kasetsart University, Bangkok, Thailand) for his outstanding contributions to our knowledge of Southeast Asian Ephemeroptera.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7E5B29C393835B07AF5F0BB0881451AC.taxon	description	Figs 35 C, 43, 44	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7E5B29C393835B07AF5F0BB0881451AC.taxon	diagnosis	Diagnosis. Femora uniformly dark brown, without transverse bands; gill I with ventral lamella composed of one or two filaments; gills II – VI with four to six filaments on dorsal lamella and three or four on ventral lamella, gill VII with 2 – 4 filaments on each lamella; posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7E5B29C393835B07AF5F0BB0881451AC.taxon	description	Description. Head. Labrum cordiform (Fig. 43 A). Ratio labrum width / clypeus width 1.06 – 1.13. Ratio labrum width / insertion width 1.46 – 1.56. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 33 – 58. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae (Fig. 43 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 7 – 10 setae below the mola (Fig. 43 C). Maxillae. Posterior margin of cardo with few hair-like and some stout setae, together with one long and stout seta in submarginal position. Proximo-lateral margin of stipes with a bunch of ten stout medium size and short setae. Apical-ventral row with 22 – 29 pectinate setae. Maxillary palp segment II 0.80 – 1.00 as long as segment I. Maxillary palp segment III 0.68 – 0.86 as long as segment I (Fig. 43 D). Stout setae on outer margin of segment I absent. Segment II inner margin with four or five stout setae. Segment II outer margin without or with one stout seta. Segment III inner margin with 3 – 5 stout setae, dorsal surface with few setae at apex. Segment III 1.57 – 1.83 × as long as base width. Labium. Labial palp segment II 0.71 – 0.94 as long as segment I. Labial palp segment III 0.73 – 0.81 as long as segment I (Fig. 43 E). Segment I inner margin with 10 – 12 stout setae, outer margin with 10 – 12 stout setae. Segment II outer margin with 3 – 5 stout setae, dorsal face with 3 – 5 stout setae. Segment III 2.90 – 3.40 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 43 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; a regular submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae; central area of upper surface with numerous long and short, stout, pointed setae (Fig. 44 A – B). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, dorsal margin with numerous hair-like setae; hind tibia with two rows of simple long, stout setae on ventral margin, dorsal margin with short and long pointed stout setae and numerous hair-like setae. Tarsal claw slightly hooked with 4 – 6 denticles progressively larger apically (Fig. 44 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with three or four filaments on dorsal lamella, ventral lamella with a single or two filaments (Fig. 44 E), gills II – VI with 4 – 6 filaments on dorsal lamella and three or four on ventral lamella (Fig. 44 F), gill VII with 2 – 4 filaments on each lamella (Fig. 44 G). Posterior margin of terga IX with minute denticles (Fig. 44 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
7E5B29C393835B07AF5F0BB0881451AC.taxon	etymology	Derivatio nominis. This species is dedicated to Dr Camille-Sophie Cozzarolo, who performed the first genetic work which revealed the diversity of New Guinean Thraulini.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B6062997E0F35C1A8C1319CB3E686F78.taxon	description	Figs 34 C, 35 B, 45, 46, 64 C, D	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B6062997E0F35C1A8C1319CB3E686F78.taxon	diagnosis	Diagnosis. Femora uniformly dark brown, without transverse bands; gill I uni-lamellate, without ventral lamella; gills II – VI with four or five filaments on dorsal lamella and two filaments on ventral lamella, gill VII with three or four filaments on dorsal lamella and a single one on ventral lamella.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B6062997E0F35C1A8C1319CB3E686F78.taxon	description	Description. Head. Labrum cordiform (Fig. 45 A). Ratio labrum width / clypeus width 1.04 – 1.10. Ratio labrum width / insertion width 1.45 – 1.53. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 36 – 37. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 45 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 7 – 9 setae below the mola (Fig. 45 C). Maxillae. Posterior margin of cardo with few hair-like and some stout setae (Fig. 34 C). Proximo-lateral margin of stipes with a bunch of six stout medium size and long setae. Apical-ventral row with 23 – 27 pectinate setae. Maxillary palp segment II 0.79 – 0.80 as long as segment I. Maxillary palp segment III 0.79 – 0.80 as long as segment I (Fig. 45 D). Stout setae on outer margin of segment I absent. Segment II inner margin with three or four stout setae. Segment II outer margin without stout setae. Segment III inner margin with five stout setae, dorsal surface with few setae at apex. Segment III 1.69 – 2.10 × as long as base width. Labium. Labial palp segment II 0.84 – 0.86 as long as segment I. Labial palp segment III 0.81 – 0.84 as long as segment I (Fig. 45 E). Segment I inner margin with 11 – 14 stout setae, outer margin with 14 stout setae. Segment II outer margin with four or five stout setae. Segment III dorsal face with three or four stout setae. Segment III 3.20 – 3.40 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 45 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; a regular submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin, shorter on hind femur; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae; central area of upper surface with numerous long and short, stout, pointed setae (Fig. 46 A). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, dorsal margin with numerous hair-like setae; hind tibia with two rows of simple and feathered long, stout setae on ventral margin, dorsal margin with short and long pointed stout setae and numerous hair-like setae. Tarsal claw slightly hooked with four or five denticles progressively larger apically (Fig. 46 B). Abdomen. Gill I with single lamella. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I uni-lamellate with three or four filaments (Fig. 46 D), gills II – VI with four or five filaments on dorsal lamella and two filaments on ventral lamella (Fig. 46 E), gill VII with three or four filaments on dorsal lamella and a single one on ventral lamella (Fig. 46 F). Posterior margin of terga IX with minute denticles (Fig. 46 C). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX. Eggs. General shape rounded. Size 130 μm × 80 μm. Chorionic surface smooth without visible attachment structures (Fig. 64 C). Micropyle in equatorial position (Fig. 64 D).	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B6062997E0F35C1A8C1319CB3E686F78.taxon	etymology	Derivatio nominis. The specific name honours the tribes of the Fuyug (Fuyughe) people, who live where the new species has been found.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E35DB7C1E5375F9E86ED4551AF3E2EEC.taxon	diagnosis	Diagnosis. Ventral lamella of gills highly reduced or absent, ventral lamella of first gill with <8 processes, even absent in some species; claw with generally 3 – 6 teeth, rarely up to 8; posterior margin of abdominal segment VIII generally without denticles; posterolateral expansions on abdominal segments VIII – IX, rarely VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E35DB7C1E5375F9E86ED4551AF3E2EEC.taxon	distribution	Distribution. Species of the genus Nonnullidens are restricted to New Guinea; N. billhilli is also reported from New Britain (Grant and Peters 1993).	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
33D473FC81D45A31AFA635E7F1E0115E.taxon	description	Figs 47, 48	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
33D473FC81D45A31AFA635E7F1E0115E.taxon	diagnosis	Diagnosis. Nymph. Labrum emargination narrow and very deep with flat denticles; femora pale brown with two transverse dark brown bands in middle and at apex; central area of upper surface of femora with very few long and pointed setae; posterior margin of abdominal segments IX with minute denticles; posterolateral expansions on the abdomen present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
33D473FC81D45A31AFA635E7F1E0115E.taxon	description	Description. Head. Labrum cordiform (Fig. 47 A). Ratio labrum width / clypeus width 1.03 – 1.07. Ratio labrum width / insertion width 1.40 – 1.42. Medial emargination deep, narrow with flat denticles. Proximal row of setae present, simple, shifted to distal margin. Number of setae on proximal row ~ 69 – 75. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 47 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with 4 – 6 setae below the mola (Fig. 47 C). Maxillae. Posterior margin of cardo with few hair-like setae, together with one stout and long seta in submarginal position. Proximo-lateral margin of stipes with a bunch of six stout setae increasing in size distally, most distal much longer than the others. Apical-ventral row with 23 – 26 pectinate setae. Maxillary palp segment II 0.79 – 0.84 as long as segment I. Maxillary palp segment III 0.61 as long as segment I (Fig. 47 D). Stout setae on outer margin of segment I absent. Segment II inner margin with one stout seta. Segment II outer margin without or with up to two stout setae. Segment III inner margin without or with up to three stout setae, dorsal surface half covered with setae. Segment III 1.25 – 1.50 × as long as base width. Labium. Labial palp segment II 0.68 – 0.86 as long as segment I. Labial palp segment III 0.58 – 0.66 as long as segment I (Fig. 47 E). Segment I inner margin with 6 – 10 stout setae outer margin with 9 – 13 stout setae. Segment II outer margin with two or three stout setae, dorsal face with two or three stout setae. Segment III 2.71 – 2.75 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 47 F). Thorax. Dorsal margin of femur with a row of long, pointed setae; irregular submarginal row of long, pointed setae as long as those on the dorsal margin; ventral margin with short, pointed setae and an irregular submarginal row of longer and blunt setae (Fig. 48 A – B); central area of upper surface with very few long, pointed setae (Fig. 48 C). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with one row of short, simple, pointed setae, and an irregular submarginal row of long, pointed setae on ventral margin, dorsal margin with numerous hair-like setae; hind tibia with a row of short simple setae on ventral margin and a submarginal row of long, simple, pointed setae, outer margin with long and short pointed setae and numerous hair-like setae. Tarsal claw hooked with three denticles progressively larger apically (Fig. 48 D). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with five filaments on dorsal lamella and a single filament on ventral lamella (Fig. 48 E), gills II – VI with seven or eight filaments on dorsal lamella and three or four filaments on ventral lamella (Fig. 48 F), gill VII with seven filaments on dorsal lamella and two filaments on ventral lamella (Fig. 48 G). Posterior margin of terga IX with minute denticles. Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
33D473FC81D45A31AFA635E7F1E0115E.taxon	etymology	Derivatio nominis. This species is dedicated to Dr Thomas Kaltenbach (Naturéum, Lausanne) who revealed the megadiversity of the mayfly genus Labiobaetis in New Guinea.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FF9A8430E3A95ADC88374C60CFF258A7.taxon	description	Figs 49, 50	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FF9A8430E3A95ADC88374C60CFF258A7.taxon	diagnosis	Diagnosis. Labrum emargination narrow without denticles; central area of upper surface of femora with very few short blunt setae; femora pale brown with two transverse dark brown bands in middle and at apex; posterior margin of abdominal segments IX with well-developed denticles; posterolateral expansions on the abdomen present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FF9A8430E3A95ADC88374C60CFF258A7.taxon	description	Description. Head. Labrum rounded (Fig. 49 A). Ratio labrum width / clypeus width 0.97 – 1.00. Ratio labrum width / insertion width 1.19 – 1.24. Medial emargination narrow without denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 51 – 75. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 49 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with 8 – 12 setae below the mola (Fig. 49 C). Maxillae. Posterior margin of cardo with few hair-like setae, together with one stout and long seta in submarginal position. Proximo-lateral margin of stipes with a single stout and long seta. Apical-ventral row with 23 – 25 pectinate setae. Maxillary palp segment II 0.76 – 0.91 as long as segment I. Maxillary palp segment III 0.62 – 0.73 as long as segment I (Fig. 49 D). Stout setae on outer margin of segment I absent. Segment II inner margin with three stout setae, outer margin without stout setae. Segment III inner margin with three or four stout setae, dorsal surface with few setae at apex, or half covered with setae. Segment III 1.78 – 2.05 × as long as base width. Labium. Labial palp segment II 0.75 as long as segment I. Labial palp segment III 0.56 as long as segment I (Fig. 49 E). Segment I inner margin with ten stout setae, outer margin with 13 stout setae. Segment II outer margin without stout setae dorsal face with three stout setae. Segment III 2.71 × as long as base width. Hypopharynx. Apex of superlingua somewhat pointed (Fig. 49 F). Thorax. Dorsal margin of femur with a row of few (<12) long, pointed setae; irregular submarginal row of blunt setae shorter than those on the dorsal margin; ventral margin with short, pointed setae and an irregular submarginal row of longer and blunt setae; central area of upper surface with very few short blunt setae (Fig. 50 A). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with two rows of long, simple, pointed setae on ventral margin, dorsal margin with few hair-like setae; hind tibia with two rows of long and short feathered and simple setae on ventral margin, outer margin with short pointed setae and few hair-like setae. Tarsal claw straight with 2 – 4 denticles progressively larger apically (Fig. 50 B). Abdomen. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I broken; gill II – VI with six or seven filaments on dorsal lamella and four on ventral lamella; gill VII with five filaments on dorsal lamella and three or four on ventral lamella. Posterior margin of terga IX with denticles. Posterior margin of terga VIII without or with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FF9A8430E3A95ADC88374C60CFF258A7.taxon	etymology	Derivatio nominis. This species is dedicated to Dr Marcela Miranda de Lima (Viçosa, Brazil).	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5D86B46012A55F69A56F8E182E52E81D.taxon	description	Fig. 51	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5D86B46012A55F69A56F8E182E52E81D.taxon	diagnosis	Diagnosis. Medial emargination of labrum narrow with rounded denticles; ventral face of labrum with scattered thin setae; femora pale brown with two transverse dark brown bands in middle and at apex; posterior margin of abdominal segments V – VIII with well-developed denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5D86B46012A55F69A56F8E182E52E81D.taxon	description	Description. Head. Labrum cordiform. Ratio labrum width / clypeus width 1.13. Ratio labrum width / insertion width 1.35. Medial emargination narrow with rounded denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 95 – 100. Distal row of setae simple. Median part of labrum in ventral view with scattered thin setae. Mandibles. Outer margin with row of setae on entire margin. Ending of the row near incisor. Right mandible with 7 – 9 setae below the mola. Maxillae. Posterior margin of cardo with numerous thin and stout setae. Proximo-lateral margin of stipes twisted. Apical-ventral row with 25 – 27 pectinate setae. Maxillary palp segment II 0.88 – 0.93 as long as segment I. Maxillary palp segment III 0.68 – 0.79 as long as segment I (Fig. 51 B). Stout setae on outer margin of segment I present. Segment II inner margin without stout setae, outer margin without stout setae. Segment III inner margin without stout setae, dorsal surface fully covered with setae. Segment III 1.67 – 1.80 × as long as base width. Labium. Labial palp segment II 0.80 – 0.81 as long as segment I. Labial palp segment III 0.62 – 0.63 as long as segment I. Segment I inner margin with 17 – 20 stout setae, outer margin with 14 – 16 stout setae. Segment II outer margin with 17 – 20 stout setae. Segment III dorsal face with four stout setae. Segment III 2.97 – 3.02 × as long as base width. Hypopharynx. Apex of superlingua truncate. Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; irregular submarginal rows of shorter and pointed setae on fore femur, regular row of shorter, pointed setae on hind femur; ventral margin with short, stout setae and a submarginal row of long, pointed stout setae, except on hind leg where those of the submarginal row are short, pointed; central area of upper surface with long, pointed setae, shorter on hind femur. Fore tibia with several rows of long, simple setae on ventral margin, outer margin with few hair-like setae; middle tibia with two rows of long, simple pointed setae on ventral margin, outer margin with few hair-like setae; hind tibia with a row of simple long, thin setae on ventral margin, upper and lower surface covered with simple and feathered setae, outer margin with short and long pointed stout setae and hair-like setae. Tarsal claw hooked, with seven or eight denticles progressively larger apically (Fig. 51 C). Abdomen. Gills I – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with ~ eight filaments on upper lamella and five on lower lamella, gills II – V with ~ 8 – 10 filaments on each lamella, gill VI with 5 – 7 filaments on each lamella, gill VII with four to five filaments on each lamella. Posterior margin of terga V – IX with denticles (Fig. 51 D – G). Posterolateral projections present on segments VII – IX. Eggs. General shape rounded. Size 140 – 145 μm × 85 – 90 μm (Fig. 51 H). Chorionic surface smooth. No attachment structures visible. Micropyle not visible.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
897C3C11098151748E246AB829BA0BA0.taxon	description	Figs 34 I, 36 C, 52, 53	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
897C3C11098151748E246AB829BA0BA0.taxon	diagnosis	Diagnosis. Femora uniformly dark brown, without transverse bands; tarsal claw with only three teeth; all gills uni-lamellate.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
897C3C11098151748E246AB829BA0BA0.taxon	description	Description. Head. Labrum cordiform (Fig. 52 A). Ratio labrum width / clypeus width 1.02. Ratio labrum width / insertion width 1.45. Medial emargination shallow without denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 30 – 40. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 52 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with six setae below the mola (Fig. 52 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with one medium size and two short stout setae (Fig. 34 I). Apical-ventral row with 20 – 25 pectinate setae. Maxillary palp segment II 0.76 – 1.0 as long as segment I. Maxillary palp segment III 0.76 – 0.9 as long as segment I (Fig. 52 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two or three stout setae. Segment II outer margin with two stout setae. Segment III inner margin with three or four stout setae, dorsal surface with few setae at apex. Segment III 1.86 – 2.07 × as long as base width. Labium. Labial palp segment II 0.73 – 0.75 as long as segment I. Labial palp segment III 0.67 – 0.80 as long as segment I (Fig. 52 E). Segment I inner margin with 7 – 9 stout setae, outer margin with 6 – 9 stout setae. Segment II, outer margin with 2 – 4 stout setae. Segment III, dorsal face with two or three stout setae. Segment III 2.88 – 3.33 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 52 F). Thorax. Dorsal margin of femur with a row of few (<12) long and pointed setae; irregular submarginal row of blunt, pointed setae shorter or as long as those on the dorsal margin; ventral margin with short, pointed setae and an irregular submarginal row of longer, blunt setae; central area of upper surface with very few short pointed and blunt setae (Fig. 53 A – B). Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with one row of short, simple, pointed setae, and an irregular submarginal row of short, pointed setae on ventral margin, dorsal margin with numerous hair-like setae; hind tibia with two rows of long and short feathered and simple setae on ventral margin, dorsal margin with long pointed setae and numerous hair-like setae. Tarsal claw slightly hooked with three denticles progressively larger apically (Fig. 53 C). Abdomen. All gills uni-lamellate, ventral lamella lacking; gill I with two filaments (Fig. 53 D), gills II – VI with four or five filaments (Fig. 53 E), gill VII with two filaments (Fig. 53 F). Posterior margin of terga IX with minute denticles. Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
897C3C11098151748E246AB829BA0BA0.taxon	etymology	Derivatio nominis. The species name is to recognise the Moi people, a native tribe of Sorong, who fight to keep their intact environment and their way of living.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
6C62AFD745D85534A43CF1E34726C655.taxon	description	Figs 54, 55	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
6C62AFD745D85534A43CF1E34726C655.taxon	diagnosis	Diagnosis. Median emargination of labrum shallow with flat denticles; ventral surface of labrum with stout setae in row; femora pale brown with two transverse dark brown bands in middle and at apex; posterolateral expansions present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
6C62AFD745D85534A43CF1E34726C655.taxon	description	Description. Head. Labrum cordiform (Fig. 54 A). Ratio labrum width / clypeus width 0.95. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 22. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 54 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with eight setae below the mola (Fig. 54 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with one long and stout seta. Apical-ventral row with 16 pectinate setae. Maxillary palp segment II 0.84 as long as segment I. Maxillary palp segment III 0.67 as long as segment I (Fig. 54 D). Stout setae on outer margin of segment I absent. Segment II inner margin with four stout setae, outer margin with four stout setae. Segment III inner margin with four stout setae, dorsal surface with few setae at apex. Segment III 1.64 × as long as base width. Labium. Labial palp segment II 0.94 as long as segment I. Labial palp segment III 0.82 as long as segment I (Fig. 54 E). Segment I inner margin with 14 stout setae, outer margin with 13 stout setae. Segment II outer margin with seven stout setae. Segment III dorsal face with six stout setae. Segment III 3.11 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 54 F). Thorax. Dorsal margin of fore femur with long, stout, pointed setae, and a submarginal row of long, stout, pointed setae; dorsal margin of mid- and hind femur with a row of long, stout, pointed setae; a submarginal row of pointed setae as long as or a little bit shorter than those on the dorsal margin; ventral margin with short, stout setae and an irregular submarginal row of long, pointed stout setae, except on hind leg where those on ventral margin are shorter; central area of upper surface with few short or long stout and pointed setae (Fig. 55 A – B). Fore tibia with two or three rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with two rows of long, simple, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with two rows of simple and feathered long, stout setae on ventral margin, outer margin with a row of long and short simple stout setae, with numerous hair-like setae. Tarsal claw hooked with 4 – 6 denticles progressively larger apically (Fig. 55 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae of the same size. Gills very small, gill I with five or six filaments on each lamella, gills II – VI with 7 – 10 filaments on each lamella (Fig. 55 D). Posterior margin of terga IX with denticles. Posterior margin of terga VIII without or with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
6C62AFD745D85534A43CF1E34726C655.taxon	etymology	Derivatio nominis. The species name is a Latin word composed of silva (forest) and pumilo (dwarf) to recognise the Melanesian Dwarf tribe of Aiome, and the land where the specimens of this new species come from.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B34BFFD0095A5567AE1EE16808A35CD1.taxon	description	Figs 34 H, 56, 57	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B34BFFD0095A5567AE1EE16808A35CD1.taxon	diagnosis	Diagnosis. Nymph. Labrum trapezoidal; dorsal face of labrum with simple row in distal position; femora pale brown with two transverse dark brown bands in middle and at apex.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B34BFFD0095A5567AE1EE16808A35CD1.taxon	description	Description. Head. Labrum trapezoid (Fig. 56 A). Ratio labrum width / clypeus width 1.11. Ratio labrum width / insertion width 1.56. Medial emargination narrow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 46. Distal row of setae present, simple. Median part of labrum in ventral view with scattered thin setae (Fig. 56 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row before incisor. Right mandible with seven setae below the mola (Fig. 56 C). Maxillae. Posterior margin of cardo with few hair-like setae, together with one stout and long seta in submarginal position. Proximo-lateral margin of stipes with a bunch of ~ six stout setae increasing in size distally (Fig. 34 H). Apical-ventral row with 32 pectinate setae. Maxillary palp segment II 0.73 – 0.78 as long as segment I. Maxillary palp segment III 0.79 – 0.81 as long as segment I (Fig. 56 D). Stout setae on outer margin of segment I absent. Segment II inner margin with four stout setae, outer margin with four stout setae. Segment III inner margin with six stout setae, dorsal surface with few setae at apex. Segment III 2.36 × as long as base width. Labium. Labial palp segment II 0.79 as long as segment I. Labial palp segment III 0.67 as long as segment I (Fig. 56 E). Segment I inner margin with nine stout setae, outer margin with ten stout setae. Segment II outer margin with five stout setae. Segment III dorsal face with three stout setae. Segment III 3.00 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 56 F). Thorax. All legs much longer than in any other species of the Thraulus lineage; fore femur 3 × longer than wide (Fig. 57 A), hind femur 4 × longer than wide (Fig. 57 B). Dorsal margin of femur with a row of long, pointed setae; regular submarginal row of blunt setae shorter than those on the dorsal margin; ventral margin with short, pointed setae and an irregular submarginal row of longer, pointed setae; central area of upper surface with few long and short blunt setae. Fore tibia with several rows of long simple or feathered setae on ventral margin, dorsal margin with few hair-like setae; middle tibia with two rows of short, simple, pointed setae on ventral margin, dorsal margin with numerous hair-like setae; hind tibia with two or three rows of long and short feathered and simple setae on ventral margin, outer margin with short, pointed setae and numerous hair-like setae. Tarsal claw hooked with four or five denticles progressively larger apically (Fig. 57 C – D). Abdomen. Most gills missing. Gill IV very long, uni-lamellate, with 5 – 6 filaments (Fig. 57 E). Filamentous projections on apex. Posterior margin of terga IX with minute denticles. Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FA1D57D131905024BA8D062294E6395A.taxon	description	Figs 58, 59	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FA1D57D131905024BA8D062294E6395A.taxon	diagnosis	Diagnosis. Nymph. Labrum trapezoidal; dorsal face of labrum with multiple rows in distal position; femora pale brown with two transverse dark brown bands in middle and at apex.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FA1D57D131905024BA8D062294E6395A.taxon	description	Description. Head. Labrum trapezoid (Fig. 58 A). Ratio labrum width / clypeus width 1.17. Ratio labrum width / insertion width 1.46. Medial emargination narrow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 106. Distal row of setae multiple. Median part of labrum in ventral view with scattered thin setae (Fig. 58 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with nine setae below the mola (Fig. 58 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with a bunch of ~ ten stout short and medium size setae. Apical-ventral row with 30 pectinate setae. Maxillary palp segment II 0.76 as long as segment I. Maxillary palp segment III 0.71 as long as segment I (Fig. 58 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two stout setae, outer margin without stout setae. Segment III inner margin with one stout seta, dorsal surface half covered with setae. Segment III 1.50 × as long as base width. Labium. Labial palp segment II 0.79 as long as segment I. Labial palp segment III 0.65 as long as segment I (Fig. 58 E). Segment I inner margin with 14 stout setae, outer margin with 15 stout setae. Segment II outer margin with nine stout setae. Segment III dorsal face with two stout setae. Segment III 2.73 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 58 F). Thorax. Dorsal margin of femur with a row of very long, thin, pointed setae; regular submarginal row of pointed setae as long as or shorter than those on the dorsal margin, multiple submarginal rows on fore femur (Fig. 59 A); ventral margin with short, pointed setae and an irregular submarginal row of long, pointed setae, except on hind femur where they are short and stout; central area of upper surface with very long pointed setae on fore femur, long and short on middle femur and short on hind femur (Fig. 59 B). Fore tibia with two or three rows of long simple or feathered setae on ventral margin, dorsal margin with numerous hair-like setae; middle tibia with a row of short, simple, pointed setae on ventral margin and a submarginal row of long, simple setae, outer margin with numerous hair-like setae; hind tibia with a marginal row of simple short, thin setae and a submarginal row of long to very long simple setae on ventral margin, outer margin with long to very long pointed setae and numerous hair-like setae. Tarsal claw slightly hooked, with five or six denticles progressively larger, except distal much larger (Fig. 59 C). Abdomen. Gill I with both lamellae. Gills II – VII with dorsal and ventral lamellae, ventral lamella smaller. Gill I with nine or ten filaments on dorsal lamella and five or six on ventral lamella, gills II – VI with 11 – 12 filaments on dorsal lamella and six or seven on ventral lamella (Fig. 59 E), gill VII with nine or ten filaments on dorsal lamella and four or five on ventral lamella. Posterior margin of terga IX with denticles (Fig. 59 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E7EE67EF018455C2903B39CE0943E58E.taxon	description	Figs 24 B, 25, 26, 34 B	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E7EE67EF018455C2903B39CE0943E58E.taxon	diagnosis	Diagnosis. Labrum width smaller than clypeus width; proximal row of setae on dorsal face of labrum with <30 setae; ventral side of labrum with stout setae in a row; maxillary palp segment III 1.7 – 2.2 × as long as base width; posterior margin of abdominal segment VIII with well-developed denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E7EE67EF018455C2903B39CE0943E58E.taxon	description	Description. Head. Labrum cordiform (Fig. 25 A). Ratio labrum width / clypeus width 0.86 – 0.97. Ratio labrum width / insertion width 1.26 – 1.30. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row 19 – 23. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 25 B). Mandibles. Outer margin with middle and distal tuft of setae. Right mandible with 9 – 11 setae below the mola (Fig. 25 C). Maxillae. Posterior margin of cardo with many hair-like setae and one stout and long seta in submarginal position (Fig. 34 B). Proximo-lateral margin of stipes with a single stout and medium size seta. Apical-ventral row with 23 – 25 pectinate setae. Maxillary palp segment II 0.82 – 0.86 as long as segment I. Maxillary palp segment III 0.71 – 0.77 as long as segment I (Fig. 25 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two to five stout setae. Segment II outer margin with three or four stout setae. Segment III inner margin with 4 – 6 stout setae, dorsal surface with few setae at apex. Segment III 1.82 – 2.44 × as long as base width. Labium. Labial palp segment II 0.67 – 0.80 as long as segment I. Labial palp segment III 0.79 – 0.90 as long as segment I (Fig. 25 E). Segment I inner margin with 5 – 9 stout setae, outer margin with 10 – 12 stout setae. Segment II outer margin with one or two stout setae. Segment III dorsal face with 5 – 7 stout setae. Segment III 3.50 – 3.80 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 25 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; submarginal row with shorter setae; ventral margin with short, blunt, stout setae and a submarginal row of feathered setae; central area of upper surface without setae (Fig. 26 A – B). Outer margin of fore tibia without setae; mid tibia with a row of stout, pointed setae on ventral margin; outer margin with only some hair-like setae; hind tibia covered with numerous simple and feathered setae, outer margin with a dense row of stout, pointed setae. Tarsal claw hooked, with 10 – 11 subequal denticles (Fig. 26 C). Abdomen. Gill I dorsal lamella narrow and lanceolate, ventral lamella shaped as other gills. Gills II – VII with dorsal and ventral lamellae of the same size. Gill II to VI with ~ 40 filaments on each lamella (Fig. 26 G); gill VII with ~ 25 filaments on each lamella. Filamentous projections along entire margin. Posterior margin of terga VIII and IX with denticles (Fig. 26 D – E). Posterior margin of terga VII with minute denticles (Fig. 26 F). Posterior margin of terga V – VI without denticles. Posterolateral projections present on segment VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
E7EE67EF018455C2903B39CE0943E58E.taxon	etymology	Derivatio nominis. This species is named after the Latin word iter, meaning path, to remember the new species has been collected along the trail from Port Moresby to Kokoda.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
A2A45575057955AA94A177CB092F8CAB.taxon	description	Figs 27, 28, 34 F	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
A2A45575057955AA94A177CB092F8CAB.taxon	diagnosis	Diagnosis. Labrum width equal to clypeus width; proximal row of setae on dorsal face of labrum with> 35 setae; maxillary palp segment III <1.5 × as long as base width; tarsal claw with four teeth, distal one much larger than the others; posterior margin of abdominal segment VII – VIII without denticles; posterolateral projections of the abdomen present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
A2A45575057955AA94A177CB092F8CAB.taxon	description	Description. Head. Labrum cordiform (Fig. 27 A). Ratio labrum width / clypeus width 1.00. Ratio labrum width / insertion width 1.40. Medial emargination shallow with flat denticles. Proximal row of setae present simple. Number of setae on proximal row 42. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae (Fig. 27 B). Mandibles. Outer margin with row of setae on distal 1 / 2. Ending of the row near incisor. Right mandible with nine setae below the mola (Fig. 27 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with two long and two short stout setae (Fig. 34 F). Apical-ventral row with 24 pectinate setae. Maxillary palp segment II 0.81 as long as segment I. Maxillary palp segment III 0.80 as long as segment I (Fig. 27 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two stout setae. Segment II outer margin without stout setae. Segment III inner margin with three stout setae, dorsal surface with few setae at apex. Segment III 1.45 × as long as base width. Labium. Labial palp segment II 0.69 as long as segment I. Labial palp segment III 0.88 as long as segment I (Fig. 27 E). Segment I inner margin with seven stout setae, outer margin with eight stout setae. Segment II outer margin with three stout setae. Segment III dorsal face with three stout setae. Segment III 3.0 × as long as base width. Hypopharynx. Apex of superlingua rounded (Fig. 27 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; submarginal row with setae of irregular length, as long or half the length of dorsal margin; ventral margin with short, blunt, stout setae and a submarginal irregular row of simple setae (Fig. 28 A); central area of upper surface with few stout, pointed setae. Fore tibia covered with numerous simple and feathered setae on ventral margin; outer margin of fore tibia without setae; mid- and hind tibia with a row of stout, pointed setae on ventral margin, outer margin without setae on mid tibia, with a sparse row of stout and pointed setae on hind tibia. Tarsal claw hooked with 4 denticles progressively larger, except distal much larger (Fig. 28 B). Abdomen. Gill I dorsal lamella narrow and lanceolate, ventral lamella shaped as other gills. Gills II – VII with dorsal and ventral lamellae of the same size. Gill I with ~ 20 filaments on the ventral lamella. Gill II – VI with 25 – 30 filaments on each lamella. Posterior margin of terga IX with denticles. Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segment VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
A2A45575057955AA94A177CB092F8CAB.taxon	etymology	Derivatio nominis. This species is named after our colleague and friend Dr Jean-Luc Gattolliat (Naturéum, Lausanne), in recognition of his outstanding contributions to the Baetidae in general and to those of New Guinea in peculiar.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
061A328CB70C5699B1B3DE24EB9F867F.taxon	description	Figs 24 C, 29, 30, 34 G	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
061A328CB70C5699B1B3DE24EB9F867F.taxon	diagnosis	Diagnosis. Apical ventral row of maxilla composed of 25 – 35 pectinate setae; outer margin of maxillary palp II with five stout setae; outer margin of labial palp segment II with five stout setae; tarsal claw with six or seven teeth progressively larger; posterior margin of abdominal segment IX with well-developed denticles; posterolateral projections of the abdomen present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
061A328CB70C5699B1B3DE24EB9F867F.taxon	description	Description. Head. Labrum cordiform (Fig. 29 A). Ratio labrum width / clypeus width 0.93. Ratio labrum width / insertion width 1.29. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 31. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 29 B). Mandibles. Outer margin with middle and distal tufts of setae. Ending of the row near incisor. Right mandible with ten setae below the mola (Fig. 29 C). Maxillae. Posterior margin of cardo with numerous hair-like setae and few stout setae, with a long and stout seta in submarginal position. Proximo-lateral margin of stipes with a single stout and long seta (Fig. 34 G). Apical-ventral row with 20 pectinate setae. Maxillary palp segment II 0.85 as long as segment I. Maxillary palp segment III 0.69 as long as segment I (Fig. 29 D). Stout setae on outer margin of segment I absent. Segment II inner margin with five stout setae. Segment II outer margin with five stout setae. Segment III inner margin with four stout setae, dorsal surface with few setae at apex. Segment III 1.73 × as long as base width. Labium. Labial palp segment II 0.84 as long as segment I. Labial palp segment III 0.74 as long as segment I (Fig. 29 E). Segment I inner margin with nine stout setae, outer margin with ten stout setae. Segment II outer margin with five stout setae. Segment III dorsal face with four stout setae. Segment III 3.11 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 29 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; submarginal row with setae of irregular length, row as long or half the length of dorsal margin; ventral margin with short, blunt, stout setae and a submarginal irregular row of simple setae (Fig. 30 A – B); central area of upper surface with few long, pointed setae. Fore tibia with three rows of numerous simple and feathered setae on ventral margin; outer margin of fore tibia with few hair-like setae; middle tibia with two rows of long, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with a row of long, thin setae on ventral margin, with a sublateral row of simple and feathered setae, ventral surface covered with simple and feathered setae, outer margin with long, pointed stout setae. Tarsal claw hooked with six or seven denticles progressively larger apically (Fig. 30 C). Abdomen. Gill I dorsal lamella narrow and lanceolate, ventral lamella shaped as other gills. Gills II – VII with dorsal and ventral lamellae of the same size. Gill II – VI with ~ 25 filaments on each lamella, gill VII with ~ 15 filaments. Posterior margin of terga IX with denticles. Posterior margin of terga VIII with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
061A328CB70C5699B1B3DE24EB9F867F.taxon	etymology	Derivatio nominis. This species is dedicated to Dr Pascale Derleth Sartori, wife of the first author.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9558DEE90AD553EE9C26385D5308F8F0.taxon	diagnosis	Diagnosis. Identical to the generic diagnosis, except that gill I is always composed of one dorsal lanceolate lamella and a large oval fimbriate ventral lamella.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9558DEE90AD553EE9C26385D5308F8F0.taxon	type_taxon	Type species. Masharikella fasciatus (Kimmins, 1956) by original designation.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9558DEE90AD553EE9C26385D5308F8F0.taxon	type_taxon	Included species. Thraulus (Masharikella) fasciatus (Kimmins, 1956), comb. nov. [Uganda, Zimbabwe], T. (M.) gopalani Grant & Sivaramakrishnan, 1985, comb. nov. [southern India], Th. (M.) iteris sp. nov. [Papua New Guinea], Th. (M.) johannisluci sp. nov. [Papua New Guinea], T. (M.) malabarensis Vasanth, Subramanian & Selvakumar, 2022, comb. nov. [southern India], T. (M.) opifer Grant, 2024, comb. nov. [northern Australia], Th. (M.) pascalae sp. nov. [Indonesia, Papua], Th. (M.) samueli sp. nov. [Papua New Guinea], T. (M.) thiagarajani Balasubramanian & Muthukatturaja, 2019, comb. nov. [southern India], and T. (M.) torrentis (Gillies, 1964), comb. nov. [Tanzania, Angola].	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9558DEE90AD553EE9C26385D5308F8F0.taxon	distribution	Distribution. The subgenus exhibits a disjunct distribution in Africa, southern India, New Guinea, and northern Australia. Currently, it is surprisingly not recorded from Oriental southeast Asia, suggesting the species found in New Guinea originate from tropical Australia. This may indicate a Gondwanan origin for the subgenus. Its presence in Madagascar is expected but yet not proved.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B1B9E9DD110953B1BCCF3C4E9BA8A96F.taxon	description	Figs 24 A, 31, 32, 64 A, B	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B1B9E9DD110953B1BCCF3C4E9BA8A96F.taxon	diagnosis	Diagnosis. Tarsal claw straight, without teeth; apical ventral row of maxilla composed of 16 – 20 pectinate setae; posterior margin of abdominal segment IX with minute denticles; posterolateral projections of the abdomen present on segments VIII – IX only.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B1B9E9DD110953B1BCCF3C4E9BA8A96F.taxon	description	Description. Head. Labrum cordiform (Fig. 31 A). Ratio labrum width / clypeus width 0.89 – 0.98. Ratio labrum width / insertion width 1.22 – 1.24. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 16 – 37. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 31 B). Mandibles. Outer margin with middle and distal tuft of setae. Right mandible with 6 – 11 setae below mola (Fig. 31 C). Maxillae. Posterior margin of cardo with few hair-like setae and a long and stout seta in submarginal position. Proximo-lateral margin of stipes with a single stout and medium size seta. Apical-ventral row with 16 – 20 pectinate setae. Maxillary palp segment II 0.79 – 0.92 as long as segment I. Maxillary palp segment III 0.62 – 0.68 as long as segment I (Fig. 31 D). Stout setae on outer margin of segment I absent. Segment II inner margin with two or three stout setae. Segment II outer margin without stout setae. Segment III inner margin with 2 – 4 stout setae, dorsal surface with few setae at apex. Segment III 1.5 – 2.0 × as long as base width. Labium. Labial palp segment II 0.72 – 0.76 as long as segment I. Labial palp segment III 0.79 – 0.92 as long as segment I. Segment I inner margin with 7 – 12 stout setae, outer margin with 8 – 12 stout setae. Segment II outer margin without or with one or two stout setae. Segment III dorsal face with five or six stout setae. Segment III 3.0 – 3.67 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 31 E). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; incomplete submarginal row, sometimes with scattered long, pointed setae; ventral margin with short, blunt, stout setae; central area of upper surface without setae (Fig. 32 A – B). Inner margin of fore- and mid- tibia covered with several rows of simple and feathered setae; inner margin of hind tibia with several rows of simple and feathered setae; outer margin of fore- and mid- tibia naked, with a row of stout, pointed setae on hind tibia, especially developed in the distal half. Tarsal claw straight. Denticles absent (Fig. 32 C). Abdomen. Gill I dorsal lamella narrow and lanceolate, ventral lamella shaped as other gills (Fig. 32 E). Gills II – VII with dorsal and ventral lamellae of the same size. Ventral lamella of gill I with ~ 25 long filaments; dorsal and ventral lamellae of gills II – VI with ~ 40 very long filaments each (Fig. 32 F); gill VII with ~ 20 filaments on each lamella. Posterior margin of terga IX with small denticles (Fig. 32 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections on segments VIII – IX. Eggs. General shape elongated. Size ca 185 μm × 80 μm. Chorionic surface with irregular longitudinal ridges connected transversally, forming rectangular smooth areas in-between (Fig. 64 A). On each pole arise basally six equally distant pairs of long, apically hooked attachment structure (Fig. 64 B). Micropyle not visible.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
B1B9E9DD110953B1BCCF3C4E9BA8A96F.taxon	etymology	Derivatio nominis. This species is dedicated to Samuel Sartori, grandson of the first author.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9FE71F2E93C95D74955B84C4FDCECCC7.taxon	description	Fig. 33	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9FE71F2E93C95D74955B84C4FDCECCC7.taxon	diagnosis	Diagnosis. Nymph. Labrum width smaller than clypeus width; proximal row of setae on dorsal face of labrum with <30 setae; ventral side of labrum with scattered stout setae; maxillary palp segment III ~ 1.7 × as long as base width; posterior margin of abdominal segment VIII with minute denticles; posterolateral projections on the abdomen present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
9FE71F2E93C95D74955B84C4FDCECCC7.taxon	description	Description. Head. Labrum cordiform (Fig. 33 A). Ratio labrum width / clypeus width 0.95. Ratio labrum width / insertion width 1.33. Medial emargination shallow with flat denticles. Proximal row of setae present simple. Number of setae on proximal row 23. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae (Fig. 33 B). Mandibles. Outer margin with middle and distal tuft of setae. Right mandible with 11 setae below the mola (Fig. 33 C). Maxillae. Posterior margin of cardo with few hair-like setae. Proximo-lateral margin of stipes with two long and two short stout setae. Apical-ventral row with 24 pectinate setae. Maxillary palp segment II 0.78 as long as segment I. Maxillary palp segment III 0.74 as long as segment I (Fig. 33 D). Stout setae on outer margin of segment I absent. Segment II inner margin with three stout setae. Segment II outer margin with four stout setae. Segment III inner margin with three stout setae, dorsal surface with few setae at apex. Segment III 1.70 × as long as base width. Labium. Labial palp segment II 0.71 as long as segment I. Labial palp segment III 0.90 as long as segment I (Fig. 33 E). Segment I inner margin with seven stout setae, outer margin with ten stout setae. Segment II outer margin with one stout seta. Segment III dorsal face with five stout setae. Segment III 3.00 × as long as base width. Hypopharynx. Apex of superlingua emarginated (Fig. 33 F). Abdomen. Gill I dorsal lamella narrow and lanceolate, ventral lamella shaped as other gills. Gill I with ~ 18 filaments. Posterior margin of terga IX with denticles. Posterior margin of terga VII – VIII with minute denticles. Posterior margin of terga V – VI without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5F268960C10D5150B425F2F43AD55B2A.taxon	diagnosis	Diagnosis. Identical to the generic diagnosis, except that gill I is always composed of one or two lanceolate and long lamellae.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5F268960C10D5150B425F2F43AD55B2A.taxon	type_taxon	Type species. Thraulus bellus Eaton, 1881 by original designation.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
5F268960C10D5150B425F2F43AD55B2A.taxon	distribution	Distribution. Mainly in the Oriental and Palaearctic regions, with only two species known from the Afrotropical area in the islands of Comoros and Madagascar; the subgenus seems absent from continental Africa south of the Sahara, as well as from Australia.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
673A0BA7DB475123AC428D352F7DF545.taxon	description	Figs 2 A, 3, 4, 34 A	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
673A0BA7DB475123AC428D352F7DF545.taxon	diagnosis	Diagnosis. Labrum rectangular, with emargination shallow; proximo-lateral margin of stipes with a single long and stout seta; maxillary palp greatly elongated; outer margin of maxillary palp segment II without stout setae; inner margin of labial palp segment I with 16 – 18 stout setae; apex of superlingua truncate. Gills II – VI completely fringed with filaments, with ~ 30 filaments on dorsal lamella and ~ 40 on ventral lamella; posterior margin of abdominal segment VIII without denticles; posterolateral projections of the abdomen present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
673A0BA7DB475123AC428D352F7DF545.taxon	description	Description. Head. Labrum rectangular (Fig. 3 A). Ratio labrum width / clypeus width 0.87 – 0.90. Ratio labrum width / insertion width 1.08 – 1.10. Medial emargination shallow with rounded denticles (Fig. 3 B). Proximal row of setae present, simple. Number of setae on proximal row ~ 75 – 100. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 3 B). Mandibles. Outer margin with row of setae on distal 2 / 3. Ending of the row before incisor. Right mandible with 10 or 11 setae below the mola (Fig. 3 C). Maxillae. Posterior margin of cardo with few long hair-like setae, together with some long and stout setae (Fig. 34 A). Proximo-lateral margin of stipes with a single long and stout seta. Apical-ventral row with 15 – 18 pectinate setae. Maxillary palp 1.5 × longer than maxilla; maxillary palp segment II 1.17 – 1.39 × as long as segment I. Maxillary palp segment III 0.77 – 0.94 as long as segment I (Fig. 3 D). Few stout setae on outer margin of segment I. Segment II inner margin with one stout seta. Segment II outer margin without stout setae. Segment III inner margin with three or four stout setae, dorsal surface fully covered with setae, 2.57 – 2.64 × as long as base width. Labium. Labial palp segment II 0.59 – 0.71 as long as segment I. Labial palp segment III 0.64 – 0.71 as long as segment I (Fig. 3 E). Segment I inner margin with 16 – 18 stout setae, outer margin with 12 – 15 stout setae. Segment II outer margin with two or three stout setae. Segment III dorsal face with six or seven stout setae (Fig. 3 F), 2.00 – 2.43 × as long as base width. Hypopharynx apex of superlingua truncate. Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; submarginal row with setae as long as those of dorsal margin (Fig. 4 A); ventral margin with short, blunt, stout setae and a submarginal row of long, pointed, stout setae, except on hind leg where they are short and blunt (Fig. 4 B); central area of upper surface with few long pointed setae. Fore tibia with several rows of numerous simple feathered setae on ventral margin; outer margin with few hair-like setae; middle tibia with two rows of long pointed setae on ventral margin, outer margin with few hair-like setae; hind tibia with a row of long thin setae on ventral margin, upper and lower surfaces covered with simple and some feathered setae, outer margin with short and long, pointed, stout setae. Tarsal claw straight with 5 – 8 minutes denticles subequal in length (Fig. 4 C). Abdomen. Gill I with single bifid lamella. Gills II to VII with dorsal and ventral lamellae, ventral lamella smaller. Gills II – VI with upper lamella lacking filaments on basal half of outer margin, with ~ 25 – 30 short filaments, lower lamella with ~ 35 – 40 short filaments (Fig. 4 E); on both lamellae apical filament much longer than the others; gill VII small, with narrow lamellae ending with a single filament (Fig. 4 F). Posterior margin of terga IX with denticles (Fig. 4 D). Posterior margin of terga V – VIII without denticles. Posterolateral projections present on segments VIII and IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
673A0BA7DB475123AC428D352F7DF545.taxon	etymology	Derivatio nominis. This species is dedicated to Eloïse Sartori, daughter of the first author.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
4C6BE53923B65C4F9C1A7BAB34A52F25.taxon	description	Figs 2 B, 5, 6	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
4C6BE53923B65C4F9C1A7BAB34A52F25.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow with flat denticles; dorsal face of labrum with a simple row in distal position; median part of labrum in ventral view with stout setae in row; apical ventral row of maxilla composed of 19 – 23 pectinate setae; tarsal claw with 7 – 10 teeth; gill lamellae on segments II – VI with> 25 filaments each; posterolateral projections of the abdomen present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
4C6BE53923B65C4F9C1A7BAB34A52F25.taxon	description	Description. Head. Labrum cordiform (Fig. 5 A). Ratio labrum width / clypeus width 0.90 – 0.95. Ratio labrum width / insertion width 1.23 – 1.24. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 26 – 40. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 5 B). Mandibles. Outer margin with row of setae on distal 2 / 3. Ending of the row near incisor. Right mandible with 10 – 17 setae below the mola (Fig. 5 C). Maxillae. Posterior margin of cardo with few hair-like setae and three or four stout and long setae in submarginal position. Proximo-lateral margin of stipes with a single long and stout seta. Apical-ventral row with 19 – 23 pectinate setae. Maxillary palp segment II 0.89 – 1.00 as long as segment I. Maxillary palp segment III 0.67 – 0.80 as long as segment I (Fig. 5 D). Stout setae on outer margin of segment I absent. Segment II inner margin with 3 – 7 stout setae, outer margin with 3 – 5 stout setae. Segment III inner margin with four or five stout setae, dorsal surface with few setae at apex, 1.60 – 2.10 × as long as base width. Labium. Labial palp segment II 0.69 – 0.74 as long as segment I. Labial palp segment III 0.96 – 0.86 as long as segment I (Fig. 5 E). Segment I inner margin with 9 – 15 stout setae, outer margin with 15 – 20 stout setae. Segment II outer margin with two to four stout setae. Segment III dorsal face with five or six setae, 3.00 – 3.38 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 5 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; irregular submarginal row with setae as long as or half the size of those of dorsal margin (Fig. 6 A); ventral margin with shorter, stout setae and a submarginal row of long and pointed stout setae, except on hind leg where those on ventral margin are short and blunt (Fig. 6 B); central area of upper surface with few long, pointed setae somewhat arranged in a longitudinal row. Fore tibia with several rows of numerous simple, feathered setae on ventral margin; outer margin of fore tibia with few hair-like setae; middle tibia with two rows of long, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with a row of long thin setae on ventral margin, upper and lower surface covered with simple and some feathered setae, outer margin with short and long, pointed, stout setae. Tarsal claw straight with 7 – 10 denticles progressively larger apically (Fig. 6 C). Abdomen. Gill I with single bifid lamella. Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gill II – VI with upper and lower lamella entirely bordered with ~ 30 short filaments (Fig. 6 F); gill VII with ~ 15 filaments and ~ 10 filaments on upper and lower lamellae, respectively (Fig. 6 G). Posterior margin of terga IX with minute denticles (Fig. 6 D), or with small denticles. Posterior margin of terga VIII with minute denticles (Fig. 6 E). Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
4C6BE53923B65C4F9C1A7BAB34A52F25.taxon	etymology	Derivatio nominis. This species is dedicated to Prof. Peter Grant (SWOSU, Oklahoma, USA) for his outstanding contribution to the study of the genus Thraulus.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FCCCD12E5C33593FB5CC41CF983A4D9E.taxon	description	Figs 7, 8	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FCCCD12E5C33593FB5CC41CF983A4D9E.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow with flat denticles; dorsal face of labrum with a simple row in distal position; median part of labrum in ventral view with stout setae in row; apical ventral row of maxilla composed of 17 pectinate setae; tarsal claw with six teeth; gill lamellae on segments II – VI with> 25 filaments each; posterolateral projections of the abdomen present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FCCCD12E5C33593FB5CC41CF983A4D9E.taxon	description	Description. Head. Labrum cordiform (Fig. 7 A). Ratio labrum width / clypeus width 0.91. Ratio labrum width / insertion width 1.21. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 24. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 7 B). Mandibles. Outer margin with middle and distal tuft of setae. Right mandible with 12 setae below the mola (Fig. 7 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with a single long and stout seta. Apical-ventral row with 17 pectinate setae. Maxillary palp segment II 0.82 as long as segment I. Maxillary palp segment III 0.75 as long as segment I (Fig. 7 D). Stout setae on outer margin of segment I absent. Segment II inner margin with three stout setae. Segment II outer margin without stout setae. Segment III inner margin with four stout setae. Segment III dorsal surface with few setae at apex. Segment III 1.60 × as long as base width. Labium. Labial palp segment II 0.68 as long as segment I. Labial palp segment III 0.76 as long as segment I (Fig. 7 E). Segment I inner margin with nine stout setae, outer margin with eight stout setae. Segment II outer margin with four stout setae. Segment III dorsal face with four stout setae. Segment III 3.11 × as long as base width. Hypopharynx. Apex of superlingua pointed (Fig. 7 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; irregular submarginal row of shorter pointed setae; ventral margin with short stout setae and a submarginal row of long, pointed, stout setae, except on hind leg where those on ventral margin are very short and blunt (Fig. 8 A); central area of upper surface with minute pointed setae. Middle tibia with two rows of long, simple, feathered, or pointed setae on ventral margin, outer margin with few hair-like setae; hind tibia with row of simple long and thin setae on ventral margin, upper and lower surface covered with simple and feathered setae, outer margin with short and long pointed, stout setae and hair-like setae. Tarsal claw straight with six denticles progressively larger apically (Fig. 8 B). Abdomen. Gill I with single bifid lamella. Gills II – VII with dorsal and ventral lamellae of the same size. Gill II – VI with upper lamella lacking filaments on basal half of outer margin, with ~ 25 – 30 filaments on each lamella (Fig. 8 C); gill VII with ~ 15 filaments on each lamella. Posterior margin of terga IX with denticles. Posterior margin of terga VIII with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
FCCCD12E5C33593FB5CC41CF983A4D9E.taxon	etymology	Derivatio nominis. The Latin adjective longinquus means distant or lost, as an indication of the remote place where the species was found.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
09A3A1A7479950379F4FE58CE6DAA3B6.taxon	description	Figs 9, 10	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
09A3A1A7479950379F4FE58CE6DAA3B6.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow with flat denticles; median part of labrum in ventral view with stout setae in a row; proximal row of setae on dorsal face of labrum with 19 – 23 setae; apical ventral row of maxilla composed of 15 – 17 pectinate setae; tarsal claw with eight or nine teeth, distal one much larger; gill lamellae on segments II – VI fringed only on distal half, with <20 filaments on each.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
09A3A1A7479950379F4FE58CE6DAA3B6.taxon	description	Description. Head. Labrum cordiform (Fig. 9 A). Ratio labrum width / clypeus width 0.81 – 0.90. Ratio labrum width / insertion width 1.15 – 1.26. Medial emargination shallow with flat denticles. Proximal row of setae present simple. Number of setae on proximal row ~ 19 – 23. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in row (Fig. 9 B). Mandibles. Outer margin with middle and distal tuft of setae. Right mandible with eight to nine setae below the mola (Fig. 9 C). Maxillae. Posterior margin of cardo with few hair-like setae and one or two stout and long setae in submarginal position. Proximo-lateral margin of stipes with a single long and stout seta. Apical-ventral row with 15 – 17 pectinate setae. Maxillary palp segment II 0.88 – 0.90 as long as segment I. Maxillary palp segment III 0.76 – 0.85 as long as segment I (Fig. 9 D). Stout setae on outer margin of segment I absent. Segment II inner margin with three or four stout setae. Segment II outer margin with 3 – 5 stout setae. Segment III inner margin with 2 – 4 stout setae. Segment III 1.86 – 2.00 × as long as base width. Labium. Labial palp segment II 0.78 – 0.81 as long as segment I. Labial palp segment III 0.69 – 0.74 as long as segment I (Fig. 9 E). Segment I inner margin with 8 – 13 stout setae, outer margin with nine stout setae. Segment II outer margin with three or four stout setae. Segment III dorsal face with four or five stout setae. Segment III 2.86 – 3.29 × as long as base width. Hypopharynx. Apex of superlingua somewhat pointed (Fig. 9 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae; submarginal row with long setae decreasing in size towards apex (Fig. 10 A); ventral margin with short and stout setae and a submarginal row of long and pointed stout setae, except on hind leg where those on ventral margin are short and blunt (Fig. 10 B); central area of upper surface with few long and pointed setae. Fore tibia with several rows of numerous mostly feathered setae on ventral margin; outer margin of fore tibia with few hair-like setae; middle tibia with two rows of long and simple or feathered, pointed setae on ventral margin, outer margin with numerous hair-like setae; hind tibia with a row of simple short thin setae on ventral margin, upper and lower surfaces covered with simple and feathered setae, outer margin with short and long, pointed, stout setae. Tarsal claw hooked with eight or nine denticles progressively larger, except distal one much larger (Fig. 10 C). Abdomen. Gill I with single bifid lamella. Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gill II – VI with upper lamella lacking filaments on basal half of outer margin, with <20 short filaments, lower lamella with <20 short filaments (Fig. 10 D). Posterior margin of terga IX with denticles. Posterior margin of terga VIII without denticles, or with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
09A3A1A7479950379F4FE58CE6DAA3B6.taxon	etymology	Derivatio nominis. This species is named after the district of Nabire where it can be found and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
F817B76E72485635A839B78534428E89.taxon	description	Figs 11 A, 12, 13, 20 A	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
F817B76E72485635A839B78534428E89.taxon	diagnosis	Diagnosis. Labrum rectangular, with emargination narrow; proximo-lateral margin of stipes with one long and stout seta together with two small and stout setae; maxillary palp greatly elongated; outer margin of maxillary palp segment II with 1 – 3 stout setae; inner margin of labial palp segment I with 20 – 26 stout setae; apex of superlingua rounded. Gills II – VI without filaments on basal half, with ~ 15 filaments on dorsal lamella and ~ 20 on ventral lamella; posterior margin of abdominal segment VIII with minute denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
F817B76E72485635A839B78534428E89.taxon	description	Description. Head. Labrum rectangular (Fig. 12 A). Ratio labrum width / clypeus width 0.88 – 0.92. Ratio labrum width / insertion width 1.12 – 1.13. Medial emargination narrow with rounded denticles. Proximal row of setae present, simple. Number of setae on proximal row 69 – 87. Distal row of setae present, simple. Median part of labrum in ventral view with stout setae in a row (Fig. 12 B). Mandibles. Outer margin with row of setae on distal 2 / 3, row ending before incisor. Right mandible with 7 – 10 setae below the mola (Fig. 12 C). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with one long stout seta together with two small stout setae. Apical-ventral row with 15 – 17 pectinate setae. Maxillary palp extremely long, almost 2 × longer than maxilla; segment II 1.3 – 1.8 × as long as segment I. Maxillary palp segment III 0.77 – 0.99 as long as segment 1 (Fig. 12 D). Stout setae on outer margin of segment I present. Segment II inner margin with two stout setae, outer margin with 1 – 3 stout setae. Segment III inner margin with three or four stout setae, dorsal surface with few setae at apex. Segment III 2.46 – 3.00 × as long as base width. Labium. Labial palp segment II 0.67 – 0.71 as long as segment 1. Labial palp segment III 0.63 – 0.70 as long as segment 1 (Fig. 12 E). Segment I inner margin with 20 – 26 stout setae, outer margin with 12 – 15 stout setae. Segment II outer margin with two or three stout setae. Segment III dorsal face with six stout setae. Segment III 2.15 – 2.29 × as long as base width. Hypopharynx. Apex of superlingua rounded (Fig. 12 F). Thorax. Dorsal margin of femur with a row of long stout setae, and a submarginal row of same setae; ventral margin with shorter and stouter setae (Fig. 13 A, B); upper surface with few long and pointed but stout setae. Tarsal claw straight with 4 – 7 subequal denticles (Fig. 13 C). Abdomen. Gill I with single bifid lamella (Fig. 13 F). Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gills II – VI with ~ 15 filaments on upper lamella and ~ 20 on the ventral one, filamentous projections lacking on proximal half (Fig. 13 G); gill VII with three filaments on each lamella, one much longer than the other two (Fig. 13 H). Posterior margin of terga IX with denticles (Fig. 13 D). Posterior margin of terga VIII with minute denticles (Fig. 13 E). Posterior margin of terga V – VII without denticles. Posterolateral projections present only on segment VIII – IX. Eggs. General shape elongated. Size 190 μm × 75 μm. Chorionic surface smooth, with longitudinal ridges weakly elevated, interrupted partially in the equatorial area. No attachment structures. Micropyle in equatorial position (Fig. 20 A).	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
F817B76E72485635A839B78534428E89.taxon	etymology	Derivatio nominis. This species is dedicated to Noe Sartori, grandson of the first author, and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
820127E1D2DE59DE99F215EE7EBEADB7.taxon	description	Figs 11 B, 14, 15, 20 B – D	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
820127E1D2DE59DE99F215EE7EBEADB7.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow with flat denticles; median part of labrum in ventral view with stout setae in a row; outer margin of mandibles with tuft of setae in medium and distal positions; posterior margin of cardo with few hair-like setae; gill lamellae on segments II – VI without filaments on basal half, with <20 filaments on each; posterolateral projections of the abdomen present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
820127E1D2DE59DE99F215EE7EBEADB7.taxon	description	Description. Head. Labrum cordiform (Fig. 14 A). Ratio labrum width / clypeus width 0.93 – 0.94. Ratio labrum width / insertion width 1.29 – 1.30. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row 35 – 47. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae (Fig. 14 B). Mandibles. Outer margin with middle tuft of setae, and with distal tuft of setae. Right mandible with nine or ten setae below the mola (Fig. 14 C). Maxillae. Posterior margin of cardo with few hair-like setae. Proximo-lateral margin of stipes with a single stout and medium-sized seta. Apical-ventral row with 25 – 29 pectinate setae. Maxillary palp segment II 0.77 – 1.05 as long as segment I. Maxillary palp segment III 0.69 – 0.73 as long as segment I (Fig. 14 D). Stout setae on outer margin of segment I absent. Segment II inner margin with five stout setae, outer margin with five or six stout setae. Segment III inner margin with three stout setae, dorsal surface with few setae at apex. Segment III 1.89 – 2.57 × as long as base width. Labium. Labial palp segment II 0.81 – 0.82 as long as segment I. Labial palp segment III 0.71 – 0.81 as long as segment I (Fig. 14 E). Segment I inner margin with 8 – 11 stout setae, outer margin with 12 or 13 stout setae. Segment II outer margin with four or five stout setae. Segment III, dorsal face with three or four stout setae. Segment III 3.0 – 3.57 as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 14 F). Thorax. Dorsal margin of femur with a row of stout and long setae, and a submarginal row with same setae; ventral margin with shorter and stouter setae; upper surface with few long and pointed stout setae (Fig. 15 A). Tarsal claw hooked with 4 – 6 denticles progressively larger apically. Abdomen. Gill I with single bifid lamella. Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gill II to VI with seven to ten filaments on upper lamella and 12 – 14 on the ventral one, filamentous projections lacking on proximal half; gill VII with three or four filaments on each lamella. Posterior margin of terga IX with denticles. Posterior margin of terga IX with denticles (Fig. 15 B). Posterior margin of terga VIII with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VII – IX. Eggs. General shape elongated. Size 190 μm × 75 μm. Chorionic surface smooth and uplifted compared to attachment structures (Fig. 20 B). Attachment structures composed of nine long slender filaments attached to each pole (Fig. 20 C); presence of two transverse filaments at the micropyle opening (Fig. 20 D). Micropyle in equatorial position.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
820127E1D2DE59DE99F215EE7EBEADB7.taxon	etymology	Derivatio nominis. This species is named after the ethnic group Ogea, who live close to the Gogol River, and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
3AB9A5D350E455A392654BA483FF654D.taxon	description	Figs 16, 17, 34 E	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
3AB9A5D350E455A392654BA483FF654D.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow without denticles; dorsal face of labrum with multiple rows in distal position; tarsal claw hooked with 15 teeth and a palisade of four other teeth in distal position; gill lamellae on segments II – VI with> 25 filaments each; posterolateral projections of the abdomen present on segments VIII and IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
3AB9A5D350E455A392654BA483FF654D.taxon	description	Description. Head. Labrum cordiform (Fig. 16 A). Ratio labrum width / clypeus width 0.95. Ratio labrum width / insertion width 1.39. Medial emargination shallow without denticles. Proximal row of setae present, simple. Number of setae on proximal row ~ 29. Distal row of setae multiple (Fig. 16 C). Median part of labrum in ventral view with stout setae in a row (Fig. 16 B). Mandibles. Outer margin with middle tuft of setae. Right mandible with ten setae below mola (Fig. 16 D). Maxillae. Posterior margin of cardo broken. Proximo-lateral margin of stipes with single long stout seta (Fig. 34 E). Apical-ventral row with 22 pectinate setae. Maxillary palp segment II 0.82 as long as segment I. Maxillary palp segment III 0.78 as long as segment I (Fig. 16 E). Stout setae on outer margin of segment I absent. Segment II inner margin with five stout setae, outer margin without stout setae. Segment III inner margin with five stout setae, dorsal surface with few setae at apex. Segment III 1.67 × as long as base width. Labium. Labial palp segment II 0.76 as long as segment I. Labial palp segment III 0.76 as long as segment I (Fig. 16 F). Segment I inner margin with two stout setae, outer margin with 12 stout setae. Segment II outer margin with ten stout setae. Segment III dorsal face with five stout setae. Segment III 2.90 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 16 F). Thorax. Dorsal margin of femur with a row of long, stout and pointed setae; regular submarginal row with setae as long as those of dorsal margin (Fig. 17 A); ventral margin with short and pointed setae and a submarginal row of short and blunt but stout setae, except on hind leg where those on ventral margin are short and blunt (Fig. 17 B); central area of upper surface with few long pointed setae. Middle tibia with two rows of long pointed setae on ventral margin, outer margin with few hair-like setae; hind tibia with a row of long thin setae on ventral margin, upper and lower surfaces covered with simple and some feathered setae, outer margin with long and short pointed stout setae. Tarsal claw hooked with 15 denticles, together with a palisade of four small denticles at apex (Fig. 17 C). Abdomen. Gill I with single bifid lamella. Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gills II – VI with upper lamella entirely bordered with ~ 35 very long filaments, lower lamella with ~ 30 very long filaments (Fig. 17 D); gill VII smaller but with the same number of long filaments. Posterior margin of terga IX with denticles. Posterior margin of terga VIII with minute denticles. Posterior margin of terga V – VII without denticles. Posterolateral projections present on segments VIII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
3AB9A5D350E455A392654BA483FF654D.taxon	etymology	Derivatio nominis. This species is named after the island from where it comes. Although we have a single nymph in hand, we think important to formally named it, since it possesses distinct characters; moreover, it is the first Leptophlebiidae described from this island and only the second mayfly species (Kaltenbach and Gattolliat 2017).	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8D89F12EC98C5C0393F55C596E757A7B.taxon	description	Figs 21, 22, 23	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8D89F12EC98C5C0393F55C596E757A7B.taxon	diagnosis	Diagnosis. Nymph. Labrum trapezoidal; ventral side of the labrum with only scattered stout setae; labial palp with segment III> 4 × as long as base width. Posterior margin of abdominal segment VI – VII with well-developed denticles.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
8D89F12EC98C5C0393F55C596E757A7B.taxon	description	Description. Head. Labrum trapezoid (Fig. 21 A). Ratio labrum width / clypeus width 1.07. Ratio labrum width / insertion width 1.52. Medial emargination shallow with flat denticles. Proximal row of setae simple, shifted to distal margin (Fig. 21 B). Number of setae on proximal row ~ 30. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae. Mandibles. Outer margin with row of setae on distal 2 / 3. Ending of the row before incisor. Right mandible with eight setae below the mola (Fig. 21 C). Maxillae. Posterior margin of cardo with very few hair-like setae and a long and stout seta in submarginal position. Proximo-lateral margin of stipes with a single long and stout seta. Apical-ventral row with 26 pectinate setae. Maxillary palps lost (Fig. 21 D). Labium. Labial palp segment II 0.87 as long as segment I. Labial palp segment III 0.89 as long as segment I (Fig. 21 E). Segment I inner margin with five stout setae, outer margin with eight stout setae. Segment II outer margin with two stout setae. Segment III dorsal face with eight stout setae. Segment III 4.40 × as long as width base. Hypopharynx. Apex of superlingua truncate (Fig. 21 F). Thorax. Dorsal margin of femur with a row of long, stout, pointed setae (Fig. 22 A); irregular submarginal row on fore femur, regular on mid femur (Fig. 22 B), absent on hind femur; ventral margin with short, stout setae and a submarginal row of long, pointed stout setae, except on hind leg where those on ventral margin are short and blunt; central area of upper surface with few long, pointed setae. Fore tibia with several rows of numerous mostly feathered setae on ventral margin; outer margin of fore tibia with few hair-like setae; middle tibia with two rows of long, simple or feathered, pointed setae on ventral margin, outer margin with few hair-like setae; hind tibia with a row of simple short, thin setae on ventral margin, upper and lower surface covered with simple and feathered setae, outer margin with short and long pointed stout setae. Tarsal claw straight with eight or nine denticles progressively larger, with a palisade of four small denticles at apex (Fig. 22 C). Abdomen. Gills lost. Posterior margin of terga VI – IX with denticles (Fig. 23 A – D). Posterior margin of terga V with minute denticles (Fig. 23 E). Posterolateral projections present on segments VII – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
0082BB0CEE8F526CB02C82AC58D055F4.taxon	description	Figs 11 C, 18, 19, 20 E, F	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
0082BB0CEE8F526CB02C82AC58D055F4.taxon	diagnosis	Diagnosis. Labrum cordiform, medial emargination shallow with flat denticles; median part of labrum in ventral view with stout setae in a row; outer margin of mandibles with row of setae on distal 2 / 3; posterior margin of cardo with many hair-like setae and one stout long seta in submarginal position; gill lamellae on segments II – VI with <20 filaments, without filaments on basal half; posterolateral projections of the abdomen present on segments VI – IX.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
0082BB0CEE8F526CB02C82AC58D055F4.taxon	description	Description. Head. Labrum cordiform (Fig. 18 A). Ratio labrum width / clypeus width 0.96 – 1.03. Ratio labrum width / insertion width 1.31 – 1.39. Medial emargination shallow with flat denticles. Proximal row of setae present, simple. Number of setae on proximal row 33 – 45. Distal row of setae present, simple. Median part of labrum in ventral view with only scattered stout setae (Fig. 18 B). Mandibles. Outer margin with row of setae on distal 2 / 3 ending near incisor. Right mandible with 7 – 11 setae below the mola (Fig. 18 C). Maxillae. Posterior margin of cardo with many hair-like setae and one stout long seta in submarginal position. Proximo-lateral margin of stipes with a single stout long seta. Apical-ventral row with 18 – 23 pectinate setae. Maxillary palp segment II 0.70 – 1.00 × as long as segment I. Maxillary palp segment III 0.67 – 0.82 as long as segment 1 (Fig. 18 D). Stout setae on outer margin of segment I absent. Segment II inner margin with four or five stout setae, outer margin with 4 – 6 stout setae. Segment III inner margin with 4 – 6 stout setae, dorsal surface with few setae at apex. Segment III 1.8 – 2.22 × as long as base width. Labium. Labial palp segment II 0.72 – 0.90 as long as segment I. Labial palp segment III 0.67 – 0.78 as long as segment I (Fig. 18 E). Segment I inner margin with 9 – 13 stout setae, outer margin with 11 or 12 stout setae. Segment II outer margin with 3 – 5 stout setae. Segment III dorsal face with 3 – 5 stout setae. Segment III 3.20 – 3.57 × as long as base width. Hypopharynx. Apex of superlingua truncate (Fig. 18 F). Thorax. Dorsal margin of femur with a row of stout, long setae, and a submarginal row with same setae; ventral margin with shorter and stouter setae; upper surface with few long, pointed stout setae (Fig. 19 A, B). Tarsal claw hooked with 4 – 7 denticles progressively larger apically (Fig. 19 C). Abdomen. Gill I with single bifid lamella (Fig. 19 F). Gills II – VII with dorsal and ventral lamellae, dorsal lamella smaller. Gills II – VI with <10 filaments on each lamella (Fig. 19 G); gill VII with two or three filaments on each lamella (Fig. 19 H). Posterior margin of terga IX with denticles (Fig. 19 D). Posterior margin of terga VII – VIII with minute denticles (Fig. 19 E). Posterior margin of terga V – VI without denticles. Posterolateral projections present on segment VI – IX. Eggs. General shape elongated. Size 190 μm x 75 μm. Chorionic surface smooth and uplifted compared to attachment structures. Attachment structures composed of 12 long and slender filaments attached to each pole and able to inflate (Fig. 20 E). At the micropyle opening, presence of two transverse short filaments, easily broken (Fig. 20 F). Micropyle in equatorial position.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
0082BB0CEE8F526CB02C82AC58D055F4.taxon	etymology	Derivatio nominis. This species is named to honour the ancient ethnic group Wemale, who still inhabits the inland of Seram where the specimens of this species were collected and is a noun in apposition.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
0823F45AA6A9592196EF62F8F6B54369.taxon	diagnosis	Diagnosis. Nymphs of the genus Thraulus are adequately characterised by Grant (2024): two dorsal rows of setae on the dorsal face of labrum; width of labrum subequal to width of clypeus (range: 0.81 – 1.07); lateral margins of clypeus parallel; posterolateral projections on abdominal segments VI – IX, VII – IX or VIII – IX; gills I – VII dissimilar; gill I composed of one or two lanceolate and long lamellae, or a dorsal lanceolate lamella and a ventral oval and fringed lamella; gills II – VII composed of two fringed lamellae along the entire margin, with some species having the basal half of lateral margin bare. Contrary to Grant (2024) the ventral side of labrum can bear not only a row of short stout setae (e. g., Fig. 12 B) but in some species scattered stout or thin setae (e. g., Fig. 18 B). The hypopharynx does not always bear a “ small, rounded, posterolateral projection on arms of the superlingua ”, compare for instance Fig. 21 F and Fig. 29 F.	en	Sartori, Michel, Salles, Frederico F. (2025): The Leptophlebiidae of New Guinea (Ephemeroptera, Choroterpinae, Thraulini). ZooKeys 1245: 41-175, DOI: 10.3897/zookeys.1245.141723
