taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
87504AD5384F5E30B65F0D33A3F4A28E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331564	https://doi.org/10.3897/zookeys.1238.124514.figure9	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
B1A3FFE360C25789816FFB4478FC02C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331564	https://doi.org/10.3897/zookeys.1238.124514.figure9	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C88F050665775F36952D6743E62AD719.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331564	https://doi.org/10.3897/zookeys.1238.124514.figure9	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
CF44E66CF77E5DDE8B109B9166E8AB2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331564	https://doi.org/10.3897/zookeys.1238.124514.figure9	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C4A7B25B00AC56B19C75F9056A205276.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331565	https://doi.org/10.3897/zookeys.1238.124514.figure10	Figure 10. Habitus of adults and leaf mines of four Argopus species A – D A. balyi E – H A. clypeatus. Host plants A – D Clematis stans at Fukuji-onsen, Gifu Pref. E – H Clematis terniflora (E – G at Tsushima Nagasaki Pref., H at Uri-tôge, Shizuoka Pref.).	Figure 10. Habitus of adults and leaf mines of four Argopus species A – D A. balyi E – H A. clypeatus. Host plants A – D Clematis stans at Fukuji-onsen, Gifu Pref. E – H Clematis terniflora (E – G at Tsushima Nagasaki Pref., H at Uri-tôge, Shizuoka Pref.).	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
2213D4CCF5115003B3B1BF5BD107650F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331565	https://doi.org/10.3897/zookeys.1238.124514.figure10	Figure 10. Habitus of adults and leaf mines of four Argopus species A – D A. balyi E – H A. clypeatus. Host plants A – D Clematis stans at Fukuji-onsen, Gifu Pref. E – H Clematis terniflora (E – G at Tsushima Nagasaki Pref., H at Uri-tôge, Shizuoka Pref.).	Figure 10. Habitus of adults and leaf mines of four Argopus species A – D A. balyi E – H A. clypeatus. Host plants A – D Clematis stans at Fukuji-onsen, Gifu Pref. E – H Clematis terniflora (E – G at Tsushima Nagasaki Pref., H at Uri-tôge, Shizuoka Pref.).	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
60A8DB8745D95D18BB59CC4E033F9804.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331566	https://doi.org/10.3897/zookeys.1238.124514.figure11	Figure 11. Adult morphology and leaf mines of Argopus punctipennis A – H on Asarum I – N on Aconitum O – S on Cirsium. Host plants A – F Asarum heterotropoides at Samani, Hidaka, Hokkaido G – H Asarum megacalyx at Mt. Haguro, Tsuruoka, Yamagata Pref. I – M Aconitum japonicum subsp. subcuneatum at Shiriya, Aomori Pref. N Aconitum sachalinense at Hinoemata, Fukushima Pref. O – R Cirsium kamtschaticum at Aibetsu, Kamikawa, Hokkaido, S Cirsium japonicum at Mt. Mikusa, Hyogo Pref. A, I, O Habitus B, J, P frontal head C, D, K, L, Q, R male genitalia in ventral and lateral views E – G, L, M, R, S leaf mines.	Figure 11. Adult morphology and leaf mines of Argopus punctipennis A – H on Asarum I – N on Aconitum O – S on Cirsium. Host plants A – F Asarum heterotropoides at Samani, Hidaka, Hokkaido G – H Asarum megacalyx at Mt. Haguro, Tsuruoka, Yamagata Pref. I – M Aconitum japonicum subsp. subcuneatum at Shiriya, Aomori Pref. N Aconitum sachalinense at Hinoemata, Fukushima Pref. O – R Cirsium kamtschaticum at Aibetsu, Kamikawa, Hokkaido, S Cirsium japonicum at Mt. Mikusa, Hyogo Pref. A, I, O Habitus B, J, P frontal head C, D, K, L, Q, R male genitalia in ventral and lateral views E – G, L, M, R, S leaf mines.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C7C0D40A34205B64AD54F122E904E9B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331573	https://doi.org/10.3897/zookeys.1238.124514.figure18	Figure 18. Adult morphology, male genitalia, and leaf mines of two Dactylispa species A – H D. higoniae I – W D. adinae sp. nov. A – L dorsal (A, I), ventral (B, J), lateral (C, K), frontal (D, L) views of adults M lateral view of prothorax N dorsal view of prothorax O arrangement of spines on interstices I – VIII of left elytra (yellow circles denote spines on interstices, size of circle indicates length of spine) E, F, P – Q aedeagus in male genitalia (E, P ventral, F, Q lateral) R spermatheca and ductus spermatheca in female genitalia S ventral view of pupal exuvia G, H, T – W leaf mines. Host plants G, H Callicarpa mollis T – W Adina pilulifera at Inohae, Nichinan, Miyazaki Pref. An arrow in W indicates pupation chamber.	Figure 18. Adult morphology, male genitalia, and leaf mines of two Dactylispa species A – H D. higoniae I – W D. adinae sp. nov. A – L dorsal (A, I), ventral (B, J), lateral (C, K), frontal (D, L) views of adults M lateral view of prothorax N dorsal view of prothorax O arrangement of spines on interstices I – VIII of left elytra (yellow circles denote spines on interstices, size of circle indicates length of spine) E, F, P – Q aedeagus in male genitalia (E, P ventral, F, Q lateral) R spermatheca and ductus spermatheca in female genitalia S ventral view of pupal exuvia G, H, T – W leaf mines. Host plants G, H Callicarpa mollis T – W Adina pilulifera at Inohae, Nichinan, Miyazaki Pref. An arrow in W indicates pupation chamber.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
1578AC17D14650A3AF89DBF1AA12F4EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331572	https://doi.org/10.3897/zookeys.1238.124514.figure17	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
CA68E1103279574791EA37A251D65E30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331573	https://doi.org/10.3897/zookeys.1238.124514.figure18	Figure 18. Adult morphology, male genitalia, and leaf mines of two Dactylispa species A – H D. higoniae I – W D. adinae sp. nov. A – L dorsal (A, I), ventral (B, J), lateral (C, K), frontal (D, L) views of adults M lateral view of prothorax N dorsal view of prothorax O arrangement of spines on interstices I – VIII of left elytra (yellow circles denote spines on interstices, size of circle indicates length of spine) E, F, P – Q aedeagus in male genitalia (E, P ventral, F, Q lateral) R spermatheca and ductus spermatheca in female genitalia S ventral view of pupal exuvia G, H, T – W leaf mines. Host plants G, H Callicarpa mollis T – W Adina pilulifera at Inohae, Nichinan, Miyazaki Pref. An arrow in W indicates pupation chamber.	Figure 18. Adult morphology, male genitalia, and leaf mines of two Dactylispa species A – H D. higoniae I – W D. adinae sp. nov. A – L dorsal (A, I), ventral (B, J), lateral (C, K), frontal (D, L) views of adults M lateral view of prothorax N dorsal view of prothorax O arrangement of spines on interstices I – VIII of left elytra (yellow circles denote spines on interstices, size of circle indicates length of spine) E, F, P – Q aedeagus in male genitalia (E, P ventral, F, Q lateral) R spermatheca and ductus spermatheca in female genitalia S ventral view of pupal exuvia G, H, T – W leaf mines. Host plants G, H Callicarpa mollis T – W Adina pilulifera at Inohae, Nichinan, Miyazaki Pref. An arrow in W indicates pupation chamber.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
5EAA560121995B82AC89A7E0F92C760F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331574	https://doi.org/10.3897/zookeys.1238.124514.figure19	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C1B6D22CE9F256E5BA36AD3583FB7124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331572	https://doi.org/10.3897/zookeys.1238.124514.figure17	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
F54BE6A9D2145229AB89EF02AC07ECEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331572	https://doi.org/10.3897/zookeys.1238.124514.figure17	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	Figure 17. Habitus of adults and leaf mines of three Dactylispa species A – D D. subquadrata E – N D. masonii O – X D. angulosa. Host plants A, B Quercus serrata at Yashiro, Hyôgo Pref. C, D Quercus dentata at Nasu, Tochigi Pref. E – H Syneilesis palmata at Kawazu, Izu, Shizuoka Pref. N – S Parasenecio hastatus subsp. orientalis at Aikappu, Akkeshi, Hokkaido J – M Cirsium suzukaense at Mt. Ibuki, Shiga Pref. N Ligularia hodgsonii at Aikappu, Akkeshi, Hokkaido O – R Salvia ranzaniana at Sabushi-gawa, Niimi, Okayama Pref. S – U Salvia nipponica at Nasu, Tochigi Pref. V Clinopodium micranthum at Nagawado, Nagano Pref. W – X Lithospermum zollingeri at Taishaku-kyô, Okayama Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
B7DDF71DD6CA5577A75EE8481A7DCDE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331559	https://doi.org/10.3897/zookeys.1238.124514.figure4	Figure 4. Habitus of adults and leaf mines of two species belonging to two Alticini genera, Longitausus and Dibolia A – G Longitarsus holsaticus H – L Dibolia japonica. Host plants A – G Pennellianthus frutescens at Mt. Tarumae, Hokkaido H – L Stachys aspera var. hispidula at Lake Shoji-ko, Yamanashi Pref.	Figure 4. Habitus of adults and leaf mines of two species belonging to two Alticini genera, Longitausus and Dibolia A – G Longitarsus holsaticus H – L Dibolia japonica. Host plants A – G Pennellianthus frutescens at Mt. Tarumae, Hokkaido H – L Stachys aspera var. hispidula at Lake Shoji-ko, Yamanashi Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
3CF83FF4F4305FB9BE2F36897A39878B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331563	https://doi.org/10.3897/zookeys.1238.124514.figure8	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
2C950025D8EA50DCBE002B8553F881BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331563	https://doi.org/10.3897/zookeys.1238.124514.figure8	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C3DA3799AADD5FA79FA227B85AC4132A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331563	https://doi.org/10.3897/zookeys.1238.124514.figure8	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C3DA3799AADD5FA79FA227B85AC4132A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331562	https://doi.org/10.3897/zookeys.1238.124514.figure7	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
FF28D89241BB52E48F745A0090CD7F51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331563	https://doi.org/10.3897/zookeys.1238.124514.figure8	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	Figure 8. Habitus of adults and leaf mines of four Halticorcus species A – H H. kasuga I – L H. sauteri M – T H. hiranoi U – X H. duodecimmaculata. Host plants A, B Lepisorus thunbergianus at Mt. Shizuhata, Shizuoka Pref. C Lepisorus onoei at Shôjiko Lake, Yamanashi Pref. D – H L. miyoshianus at Kiso-fukushima, Nagano Pref. I – J Leptochilus neopothifolius at Kanyû, Kakeroma Is., Kagoshima Pref. K – L Cyrtomium falcatum at Imasato, Amami-ôshima Is., Kagoshima Pref. M – O Phymatosorus scolopendria at Tonaki Is., Okinawa Pref. P, Q Loxogramme salicifolia at Inohae, Kitago, Miyazaki Pref. R – T Pyrrosia lingua at Mt. Yuwan, Amami-ôshima Is., Kagoshima Pref. U, V Phymatosorus scolopendria at Iriomote Is., Okinawa Pref. W, X Crypsinus yakushimensis at Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
2A706F2C79885682A69F00361CF737C2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331561	https://doi.org/10.3897/zookeys.1238.124514.figure6	Figure 6. Habitus of adults, leaf mine, and larva of Hippuriphila babai on Equisetum arvense A habitus of adults B, C leaf-mine at Mt. Teine D – F larva at Yubari, Hokkaido.	Figure 6. Habitus of adults, leaf mine, and larva of Hippuriphila babai on Equisetum arvense A habitus of adults B, C leaf-mine at Mt. Teine D – F larva at Yubari, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
3803BDD86D855A85BC885C1EBEF24EEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331559	https://doi.org/10.3897/zookeys.1238.124514.figure4	Figure 4. Habitus of adults and leaf mines of two species belonging to two Alticini genera, Longitausus and Dibolia A – G Longitarsus holsaticus H – L Dibolia japonica. Host plants A – G Pennellianthus frutescens at Mt. Tarumae, Hokkaido H – L Stachys aspera var. hispidula at Lake Shoji-ko, Yamanashi Pref.	Figure 4. Habitus of adults and leaf mines of two species belonging to two Alticini genera, Longitausus and Dibolia A – G Longitarsus holsaticus H – L Dibolia japonica. Host plants A – G Pennellianthus frutescens at Mt. Tarumae, Hokkaido H – L Stachys aspera var. hispidula at Lake Shoji-ko, Yamanashi Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C74E1ADFC9225B719A3D0EC1B2443B1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331560	https://doi.org/10.3897/zookeys.1238.124514.figure5	Figure 5. Habitus of adults, habitat, and leaf mines of two Mantura species A – I M. clavareaui J – L M. fulvipes. Host plants A – F Rumex japonicus at Notojima Island, Ishikawa Pref. G – I Polygonum aviculare at Mihogaseki, Shimane Pref. J – L Oxalis corniculata at Tamanoura, Fukue Is., Nagasaki Pref.	Figure 5. Habitus of adults, habitat, and leaf mines of two Mantura species A – I M. clavareaui J – L M. fulvipes. Host plants A – F Rumex japonicus at Notojima Island, Ishikawa Pref. G – I Polygonum aviculare at Mihogaseki, Shimane Pref. J – L Oxalis corniculata at Tamanoura, Fukue Is., Nagasaki Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
66926E35CC8A5B9CA0A5C113DFDD4DCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331560	https://doi.org/10.3897/zookeys.1238.124514.figure5	Figure 5. Habitus of adults, habitat, and leaf mines of two Mantura species A – I M. clavareaui J – L M. fulvipes. Host plants A – F Rumex japonicus at Notojima Island, Ishikawa Pref. G – I Polygonum aviculare at Mihogaseki, Shimane Pref. J – L Oxalis corniculata at Tamanoura, Fukue Is., Nagasaki Pref.	Figure 5. Habitus of adults, habitat, and leaf mines of two Mantura species A – I M. clavareaui J – L M. fulvipes. Host plants A – F Rumex japonicus at Notojima Island, Ishikawa Pref. G – I Polygonum aviculare at Mihogaseki, Shimane Pref. J – L Oxalis corniculata at Tamanoura, Fukue Is., Nagasaki Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
3BCEB7F977B75ED0B9A456E58F390465.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331556	https://doi.org/10.3897/zookeys.1238.124514.figure1	Figure 1. Two cerambycid species associated with Cycas revouta: Mimectatina meridiana ohirai (A – F) and Sybra flavostriata flavostriata (G – I). A, G adult habitus B flowering cycad C flowering megasporophyll D cross section of mined megasporophyll E mine of megasporophyll F pupa in the mine H oviposition scars on leaf stalk I larva mining the leaf stalk.	Figure 1. Two cerambycid species associated with Cycas revouta: Mimectatina meridiana ohirai (A – F) and Sybra flavostriata flavostriata (G – I). A, G adult habitus B flowering cycad C flowering megasporophyll D cross section of mined megasporophyll E mine of megasporophyll F pupa in the mine H oviposition scars on leaf stalk I larva mining the leaf stalk.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
AE67ED2B92895B4CBA64EC2901FC6073.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331575	https://doi.org/10.3897/zookeys.1238.124514.figure20	Figure 20. Adult habitus and leaf mines of Notosacantha ihai on various host plants A – D Turpinia ternata at Mt. Yuwan, Amami-ôshima, Kagoshima Pref. E, F Symplocos glauca at Yona, Kunigami, Okinawa Pref. G Symplocos sonoharae at Mt. Yuwan, Amami-ôshima, Kagoshima Pref. H Gardneria nutans at Mt. Nishime, Kunigami, Okinawa Pref. I, J Eurya japonica at Mt. Nishime, Kunigami, Okinawa Pref. K Eurya japonica at Yona, Kunigami, Okinawa Pref. L Schima wallichii at Mt. Yuwan, Amami-ôhsima, Kagoshima Pref. M – P Adinandra ryukyuensis at Yona, Kunigami, Okinawa Pref. Q – T Helicia cochinchinensis at Yona, Kunigami, Okinawa Pref. U – W Bredia okinawensis at Mt. Nishime, Kunigami, Okinawa X, Y Bredia yaeyamensis at Komi, Iriomote Is., Okinawa Pref.	Figure 20. Adult habitus and leaf mines of Notosacantha ihai on various host plants A – D Turpinia ternata at Mt. Yuwan, Amami-ôshima, Kagoshima Pref. E, F Symplocos glauca at Yona, Kunigami, Okinawa Pref. G Symplocos sonoharae at Mt. Yuwan, Amami-ôshima, Kagoshima Pref. H Gardneria nutans at Mt. Nishime, Kunigami, Okinawa Pref. I, J Eurya japonica at Mt. Nishime, Kunigami, Okinawa Pref. K Eurya japonica at Yona, Kunigami, Okinawa Pref. L Schima wallichii at Mt. Yuwan, Amami-ôhsima, Kagoshima Pref. M – P Adinandra ryukyuensis at Yona, Kunigami, Okinawa Pref. Q – T Helicia cochinchinensis at Yona, Kunigami, Okinawa Pref. U – W Bredia okinawensis at Mt. Nishime, Kunigami, Okinawa X, Y Bredia yaeyamensis at Komi, Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
55255663A5AC558E89B73AB096CC8A56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331576	https://doi.org/10.3897/zookeys.1238.124514.figure21	Figure 21. Habitus and leaf mines of two Notosacantha species A – F Notosacantha loochooana G – O N. nishiyamai A, G, L dorsal B, H lateral C, I frontal view of adults M exuvia D, F, J, K, N, O leaf mines. Host plants A – D Gardenia jasminoides at Nagakumo-toge, Amami-ôshima, Kagoshima Pref. E, F Itea oldhamii at Higashinakama, Amami-ôshima Is., Kagoshima Pref. G – J Tarenna gracilipes at Nekumachiji, Ôgimi, Okinawa Pref. K Coptosapelta diffusa at Yona, Kunigami, Okinawa Pref. L – O Randia canthioides at Yona, Kunigami, Okinawa Pref.	Figure 21. Habitus and leaf mines of two Notosacantha species A – F Notosacantha loochooana G – O N. nishiyamai A, G, L dorsal B, H lateral C, I frontal view of adults M exuvia D, F, J, K, N, O leaf mines. Host plants A – D Gardenia jasminoides at Nagakumo-toge, Amami-ôshima, Kagoshima Pref. E, F Itea oldhamii at Higashinakama, Amami-ôshima Is., Kagoshima Pref. G – J Tarenna gracilipes at Nekumachiji, Ôgimi, Okinawa Pref. K Coptosapelta diffusa at Yona, Kunigami, Okinawa Pref. L – O Randia canthioides at Yona, Kunigami, Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
D62CE5437F555EACA2A6DA5944D53884.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331576	https://doi.org/10.3897/zookeys.1238.124514.figure21	Figure 21. Habitus and leaf mines of two Notosacantha species A – F Notosacantha loochooana G – O N. nishiyamai A, G, L dorsal B, H lateral C, I frontal view of adults M exuvia D, F, J, K, N, O leaf mines. Host plants A – D Gardenia jasminoides at Nagakumo-toge, Amami-ôshima, Kagoshima Pref. E, F Itea oldhamii at Higashinakama, Amami-ôshima Is., Kagoshima Pref. G – J Tarenna gracilipes at Nekumachiji, Ôgimi, Okinawa Pref. K Coptosapelta diffusa at Yona, Kunigami, Okinawa Pref. L – O Randia canthioides at Yona, Kunigami, Okinawa Pref.	Figure 21. Habitus and leaf mines of two Notosacantha species A – F Notosacantha loochooana G – O N. nishiyamai A, G, L dorsal B, H lateral C, I frontal view of adults M exuvia D, F, J, K, N, O leaf mines. Host plants A – D Gardenia jasminoides at Nagakumo-toge, Amami-ôshima, Kagoshima Pref. E, F Itea oldhamii at Higashinakama, Amami-ôshima Is., Kagoshima Pref. G – J Tarenna gracilipes at Nekumachiji, Ôgimi, Okinawa Pref. K Coptosapelta diffusa at Yona, Kunigami, Okinawa Pref. L – O Randia canthioides at Yona, Kunigami, Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
57C3B260E46B5028A2D69C919A668008.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331558	https://doi.org/10.3897/zookeys.1238.124514.figure3	Figure 3. Habitus of adults and leaf mines of two Phyllotreta species A – I P. ezoensis J – M P. shirahatai. Host plants A – I Draba nemorosa at Kiyosato, Yamanashi Pref. J – M Cardamine appendiculata at Eniwa (J, K) and Tobai (L, M), Hokkaido.	Figure 3. Habitus of adults and leaf mines of two Phyllotreta species A – I P. ezoensis J – M P. shirahatai. Host plants A – I Draba nemorosa at Kiyosato, Yamanashi Pref. J – M Cardamine appendiculata at Eniwa (J, K) and Tobai (L, M), Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
E47EC518EAA55834B80BA48FB3EDF243.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331558	https://doi.org/10.3897/zookeys.1238.124514.figure3	Figure 3. Habitus of adults and leaf mines of two Phyllotreta species A – I P. ezoensis J – M P. shirahatai. Host plants A – I Draba nemorosa at Kiyosato, Yamanashi Pref. J – M Cardamine appendiculata at Eniwa (J, K) and Tobai (L, M), Hokkaido.	Figure 3. Habitus of adults and leaf mines of two Phyllotreta species A – I P. ezoensis J – M P. shirahatai. Host plants A – I Draba nemorosa at Kiyosato, Yamanashi Pref. J – M Cardamine appendiculata at Eniwa (J, K) and Tobai (L, M), Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
6D753692D1AC548299C2319D014B8B4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331574	https://doi.org/10.3897/zookeys.1238.124514.figure19	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
1871083BACA45D329BB612C3A7B3024A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331562	https://doi.org/10.3897/zookeys.1238.124514.figure7	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
979B4F4163E75873A503C4D9A3674176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331562	https://doi.org/10.3897/zookeys.1238.124514.figure7	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	Figure 7. Habitus of adults and larvae, habitat, and leaf mines of three Psylliodes species A – J P. punctifrons K – P P. sasakii Q – T P. subrugosa. Host plants A – F Brassica juncea at Kizu-gawa, Kyôto G – I Raphanus sativus var. hortensis at Notojima Is., Ishikawa Pref. K – M Eutrema japonicum at Futamata, Oshamanbe, Hokkaido N – P Eutrema japonicum at Nyûkawa, Takayama, Gifu Pref. Q – T Arabis hirsuta at Shimo-suwa, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
9A15B03B5D6D55ED8A7925D0A42B0CE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331574	https://doi.org/10.3897/zookeys.1238.124514.figure19	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	Figure 19. Adult morphology and leaf mines of three Hispini species A – H Dactylispa issikii I – P Rhadinosa nigrocyanea Q – W Platypria melli A, I, Q dorsal B, J, R ventral C, K, S lateral D, L, T frontal views of adults E, M, U exuviae F, G, N – P, V, W leaf mines. Host plants G, H Pleioblastus chino at Tsuge, Kuma, Kumamoto Pref. B – O Miscanthus tinctorius at Hatabe, Kumamoto Pref. P Miscanthus tinctorius at Fukube-dani, Hakusan, Ishikawa Pref. V, W Hovenia dulcis at Kin, Tsushima Is., Nagasaki Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
7CB09DD1636E5BB4A6D1913F0AD888F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331569	https://doi.org/10.3897/zookeys.1238.124514.figure14	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
F90151A6EA305851A57AE69B16CE0006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331571	https://doi.org/10.3897/zookeys.1238.124514.figure16	Figure 16. Adult morphology, behavior, and leaf mines of Asteraceae - associated Sphaeroderma species A – J S. balyi K – Z S. komiana sp. nov. A, D, F adult habitus in dorsal view G, O frontal view of head H, X adult feeding behavior L – N ventral, lateral, frontal view P – Q male genitalia R female spermatheca S landscape of the habitat at type locality B, C, E, I, J, T – W, Y, Z leaf mines. Host plants A – C Petasites japonicus at Ukawa, Kyôtango, Kyôto Pref. D, E Parasenecio kamtschaticus at Tôbai, Nemuro, Hokkaido F – J Farfugium japonicum at Kobukei, Nichinan, Miyazaki Pref. T – X Ixeris japonica at Komi, Iriomote Island, Okinawa Pref. Y – Z Youngia japonica at Hoshidate, Iriomote Is., Okinawa Pref.	Figure 16. Adult morphology, behavior, and leaf mines of Asteraceae - associated Sphaeroderma species A – J S. balyi K – Z S. komiana sp. nov. A, D, F adult habitus in dorsal view G, O frontal view of head H, X adult feeding behavior L – N ventral, lateral, frontal view P – Q male genitalia R female spermatheca S landscape of the habitat at type locality B, C, E, I, J, T – W, Y, Z leaf mines. Host plants A – C Petasites japonicus at Ukawa, Kyôtango, Kyôto Pref. D, E Parasenecio kamtschaticus at Tôbai, Nemuro, Hokkaido F – J Farfugium japonicum at Kobukei, Nichinan, Miyazaki Pref. T – X Ixeris japonica at Komi, Iriomote Island, Okinawa Pref. Y – Z Youngia japonica at Hoshidate, Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
D7143DFDE8E25947B5F7A94FFA847433.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331569	https://doi.org/10.3897/zookeys.1238.124514.figure14	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
D46F349D326155688F158275E18C838F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331569	https://doi.org/10.3897/zookeys.1238.124514.figure14	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
93A02546038B5996B8CDD597C5387176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331568	https://doi.org/10.3897/zookeys.1238.124514.figure13	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
F579D9DAE5145EA186C7552BCB3E8B73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331571	https://doi.org/10.3897/zookeys.1238.124514.figure16	Figure 16. Adult morphology, behavior, and leaf mines of Asteraceae - associated Sphaeroderma species A – J S. balyi K – Z S. komiana sp. nov. A, D, F adult habitus in dorsal view G, O frontal view of head H, X adult feeding behavior L – N ventral, lateral, frontal view P – Q male genitalia R female spermatheca S landscape of the habitat at type locality B, C, E, I, J, T – W, Y, Z leaf mines. Host plants A – C Petasites japonicus at Ukawa, Kyôtango, Kyôto Pref. D, E Parasenecio kamtschaticus at Tôbai, Nemuro, Hokkaido F – J Farfugium japonicum at Kobukei, Nichinan, Miyazaki Pref. T – X Ixeris japonica at Komi, Iriomote Island, Okinawa Pref. Y – Z Youngia japonica at Hoshidate, Iriomote Is., Okinawa Pref.	Figure 16. Adult morphology, behavior, and leaf mines of Asteraceae - associated Sphaeroderma species A – J S. balyi K – Z S. komiana sp. nov. A, D, F adult habitus in dorsal view G, O frontal view of head H, X adult feeding behavior L – N ventral, lateral, frontal view P – Q male genitalia R female spermatheca S landscape of the habitat at type locality B, C, E, I, J, T – W, Y, Z leaf mines. Host plants A – C Petasites japonicus at Ukawa, Kyôtango, Kyôto Pref. D, E Parasenecio kamtschaticus at Tôbai, Nemuro, Hokkaido F – J Farfugium japonicum at Kobukei, Nichinan, Miyazaki Pref. T – X Ixeris japonica at Komi, Iriomote Island, Okinawa Pref. Y – Z Youngia japonica at Hoshidate, Iriomote Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
E3542A3CC80051BFBD268046AC0F4712.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331567	https://doi.org/10.3897/zookeys.1238.124514.figure12	Figure 12. Habitus of adults and leaf mines of host races of Sphaeroderma nigricolle. Host plants A, B Lilium auratum at Mt. Haguro-san, Yamagata Pref. C, D Tricyrtis macropoda at Tengu-kogen, Kôchi Pref. E – G Smilax stans at Mt. Torigata, Kôchi Pref. H Smilax stans at Mt. Ôtaki, Kagawa Pref. I – K Croomia heterosepala at Kikuchi-keikoku, Kumamoto Pref. L – O Croomia japonica at Naon, Amami-ôshima Is., Kagoshima Pref.	Figure 12. Habitus of adults and leaf mines of host races of Sphaeroderma nigricolle. Host plants A, B Lilium auratum at Mt. Haguro-san, Yamagata Pref. C, D Tricyrtis macropoda at Tengu-kogen, Kôchi Pref. E – G Smilax stans at Mt. Torigata, Kôchi Pref. H Smilax stans at Mt. Ôtaki, Kagawa Pref. I – K Croomia heterosepala at Kikuchi-keikoku, Kumamoto Pref. L – O Croomia japonica at Naon, Amami-ôshima Is., Kagoshima Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
E3542A3CC80051BFBD268046AC0F4712.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331564	https://doi.org/10.3897/zookeys.1238.124514.figure9	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	Figure 9. Habitus of adults and leaf mines of four Argopistes species A, B A. coccinelliformis C – H A. biplagiata I – N A. tsekooni O – Q A. ryukyuensis. Host plants A, B Ligustrum ovalifolium at Shirahama, Wakayama Pref. C, D Osmanthus × fortunei at Mizorogaike Lake, Kyôto Pref. E – H Osmanthus heterophyllus at Iwakura, Kyôto Pref. I – K Ligustrum obtusifolium at Mt Nabejiri, Shiga Pref. L Fraxinus sieboldiana at Uri-tôge, Hamamatsu, Shizuoka Pref. M, N Syringa reticulata at Tôro Lake, Kushiro, Hokkaido O – Q Ligustrum japonicum at Abu, Okinawa Is., Okinawa Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
5580FCC1C83258DABB2B0B6D2D19DD2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331570	https://doi.org/10.3897/zookeys.1238.124514.figure15	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
CC7B707BEFDD5CFD963976061E7E27E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331570	https://doi.org/10.3897/zookeys.1238.124514.figure15	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
0A2B5252B02952B8AC5EF3DDBD44C4A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331569	https://doi.org/10.3897/zookeys.1238.124514.figure14	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
B91FA7A49F9E5B8F9F759E8DE1B58AD9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331569	https://doi.org/10.3897/zookeys.1238.124514.figure14	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	Figure 14. Habitus of adults, habitats, and leaf mines of Akebia - associated and Clematis - associated Sphaeroderma species A – C S. akebia D S. inaizumii E – L S. quadrimaculatum M – Q S. flavonotatum R – V S. separatum. Host plants A – C Akebia trifoliata at Tsuruoka, Yamagata Pref. D Akebia trifoliata at Shôji Lake, Yamanashi Pref. E – G Clematis taiwaniana var. ryukiuensis at Hedo, Okinawa Is., Okinawa Pref. J – L Clematis taiwaniana var. ryukiuensis at Tonaki Is., Okinawa Pref. M – P Clematis tashiroi at Iriomote Is. Okinawa Pref. S – V Clematis japonica at Shirakawa-gô, Gifu Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
C6A481CE356052CF92CB38D28E7F7745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331568	https://doi.org/10.3897/zookeys.1238.124514.figure13	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
2043D58A4285579FBDE8A53696A667CE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331568	https://doi.org/10.3897/zookeys.1238.124514.figure13	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	Figure 13. Habitus of adults and leaf mines of monocot-associated Sphaeroderma species A – C S. japanum D – L S. tarsatum M – P S. seriatum. Host plants A – C Commelina communis at Makidô, Niimi, Okayama Pref. E – G Sasa nipponica at Akiu, Sendai, Miyagi Pref. H, I Sasa senanensis at Kamikôchi, Nagano Pref. J – L Stipa coreana var. japonica at Mt. Fuji, Shizuoka M – P Panicum bisulcatum at Yogo Lake, Shiga Pref. (M – O) and at Seryo, Kyôto Pref. (P).	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
6C1FB0F6B2955518A89CC7168BC8900D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331570	https://doi.org/10.3897/zookeys.1238.124514.figure15	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
E8BC6C9E138E585C8D5C1EF1359589DA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331570	https://doi.org/10.3897/zookeys.1238.124514.figure15	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	Figure 15. Adult morphology, behavior, and leaf mines of Clematis - associated and Cimicifuga - associated Sphaeroderma species A – G S. placidum H – L S. unicolor M – R S. uenoi S – Z S. ohkuboi A, E, H, J, M, S, Y adult habitus in dorsal view F, N ventral view B, F, I, O Frontal view of head K, P, T adult feeding behavior C, D, G, L, Q, R, U – X, Z leaf mines. Host plants A – D Clematis apiifolia at Shyôji Lake, Yamanashi Pref. E – G Clematis apiifolia at Ashiu, Kyôto Pref. H, I Clematis terniflora at Obama, Fukui Prefecture J – L Clematis terniflora at Iya, Tokushima Prefecture M – Q Clematis terniflora at Kizu-gawa, Kyôto Pref. R Clematis terniflora at Iwakura, Kyôto Pref. S – X Cimicifuga japonica at Mt. Nakatsu-myôjin, Kôchi Pref. Y, Z Cimicifuga simplex at Shirahone-onsen, Nagano Pref.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
1815AF3760B15689BB690C3626FB9ED5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331556	https://doi.org/10.3897/zookeys.1238.124514.figure1	Figure 1. Two cerambycid species associated with Cycas revouta: Mimectatina meridiana ohirai (A – F) and Sybra flavostriata flavostriata (G – I). A, G adult habitus B flowering cycad C flowering megasporophyll D cross section of mined megasporophyll E mine of megasporophyll F pupa in the mine H oviposition scars on leaf stalk I larva mining the leaf stalk.	Figure 1. Two cerambycid species associated with Cycas revouta: Mimectatina meridiana ohirai (A – F) and Sybra flavostriata flavostriata (G – I). A, G adult habitus B flowering cycad C flowering megasporophyll D cross section of mined megasporophyll E mine of megasporophyll F pupa in the mine H oviposition scars on leaf stalk I larva mining the leaf stalk.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
0D5C15D509E75E7583120BB791EBA2AD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
19F7DD9EBC3459009C1CEF331BE3772B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
BDE06BADF2C651C58BF1A386EF1984F9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
DBE888AD23FD5D62AF7F6C5C2F140703.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
9861A8A910485893BF21C99E21F2DF55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
B6C5DBBDBD755272B54E55022F53A3E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
0A80E343CF8C5414A7468F66C9AB7B5C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
B21F7B4145165A728F234E163359BA0E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1331557	https://doi.org/10.3897/zookeys.1238.124514.figure2	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	Figure 2. Habitus and feeding scars of adult beetles and leaf mines of eight Zeugophora species A, B Z. nigricolis C – E Z. unifasciata F – I Z. chujoi J – M Z. flavonotata N – Q Z. annulata R, S Z. varipes T – V Z. hozumii W – Z Z. japonica. Host plants A, B Euonymus sieboldianus at Hirayu, Gifu Pref. C – E E. sieboldianus at Sugadaira, Nagano Pref. F – I E. fortunei at Nagao-toge, Shizuoka Pref. J – L E. tanakae at Yutsun, Iriomote Is., Okinawa Pref. M E. japonicus at Hoshidate, Iriomote Is., Okinawa Pref. N, O E. alatus at Kiyosato, Yamanashi Pref. P E. melananthus at Mt. Tara, Nagasaki Pref. Q E. fortunei at Mt. Myôjô, Niigata Pref. R, S Symplocos sawafutagi at Iyari, Nagano Pref. T – V Salix cardiophylla var. urbaniana at Kitazawa-toge, Nagano Pref. W – Z Populus suaveolens at Sôunkyô, Hokkaido.	2025-05-16	Kato, Makoto;Imada, Yume		Zenodo	biologists	Kato, Makoto;Imada, Yume			
