taxonID	type	description	language	source
A8588792EA34FFADFD853BEBFE9926A5.taxon	description	urn: lsid: zoobank. org: act: 3 F 1 F 9 C 1 A- 4698 - 4 BFA- 9541 - 2 CFC 3 E 5 EECCB	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA34FFADFD853BEBFE9926A5.taxon	type_taxon	Type species Cubicostissus palaeocaeni sp. nov., designated here by monotypy.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA34FFADFD853BEBFE9926A5.taxon	diagnosis	Diagnosis Cubicostissus gen. nov. separates from other Hysteropterini genera by their forewing more than 2.5 as long as wide, wider before mid-length, with anterior margin slightly and regularly convex and posterior margin slightly concave at the end of the clavus. The postcostal cell forms a narrow band, slightly narrower than the radial cell band, and as wide as C 1 cell. ScP + RA and RP fork late with ScP + R more than 2.5 times as long as basal cell. MP forks in C 3 at the same level of Pcu and A 1 fusion and CuA remains single up to the apical level of the clavus, then forks in an open C 5. CuP is almost straight, connected by transverse veins to CuA. Pcu and A 1 are connected in the last quarter of the clavus.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA34FFADFD853BEBFE9926A5.taxon	etymology	Etymology Arbitrary combination referring to the forewing CuA vein (cubito costa) single and the generic name Issus.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA34FFADFD853BEBFE9926A5.taxon	discussion	Note Previously placed in Issinae sec. Wang et al. (2016), Hysteropterini were recently regarded deserving a subfamily rank (Zhao et al. 2019). They differ from Issinae Issini by their late forking of CuA after the end of the clavus while CuA forks well before in Issini genera. By its venation schema, Cubicostissus gen. nov. tegmen approaches species of Tshurtshurnella Kusnezov, 1927, but it appears more elongated: 2.6 times as long as wide (× 2 or less in Tshurtshurnella) and with a late fusion of Pcu and A 1 in the last quarter of the clavus (earlier in Tshurtshurnella). From Hysteropterum Amyot & Audinet-Serville, 1843, it differs by the elongated tegmen, the absence of identified apical cells, the presence of some transverse veins between CuA and CuP and, at mid-length, the costal area almost as wide as C 1 and the open radial cell. Its simpler venation also easily separates this new taxon from all the other issid fossils currently known.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA35FFABFE5D383FFC89209B.taxon	description	urn: lsid: zoobank. org: act: 3 E 61 A 3 B 6 - E 035 - 41 B 1 - 96 CD-B 450 B 0 F 71 F 93 Figs 1 – 2	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA35FFABFE5D383FFC89209B.taxon	diagnosis	Diagnosis Described from a single imprint from Menat (Fig. 1). Separated from all Hysteropterini species by its elongated forewing, the narrow postcostal band and the conformation of the venation.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA35FFABFE5D383FFC89209B.taxon	etymology	Etymology The specific epithet means ‘ from the Palaeocaenum (Paleocene) ’ and refers to the age of the deposit where the fossil was discovered.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA35FFABFE5D383FFC89209B.taxon	materials_examined	Material examined Holotype FRANCE • 1 specimen; Puy-de-Dôme, volcano-sedimentary maar of Menat; Paleocene, Selandian; Musée de la Paléontologie, Menat; NEL 3485.	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
A8588792EA35FFABFE5D383FFC89209B.taxon	description	Description Forewing (Figs 1 – 2) 2.6 times as long as wide, wider before mid-length: 4.04 mm long, 1.63 mm wide before mid-length at MP forking level. Basal cell (bc) short, 0.29 mm long. No hypocostal plate. Anterior margin slightly and regularly convex, maximum convexity at level of MP fork. Posterior margin slightly concave at the end of the clavus. Common stem ScP + R more than twice as long as basal cell (0.67 mm long). Postcostal cell forming a narrow band, slightly narrower than radial cell, itself narrower than basal cell. ScP + R forking late: ScP + R 2.7 times as long as basal cell. Radial cell open, as wide as C 1 cell before mid-length of tegmen, with ScP + RA and RP running parallel. Vein MP 1.33 mm long, forking late in cell C 3 at same level of fusion of Pcu and A 1; MP 1 + 2 in straight line with vein MP; MP 3 + 4 diverging posteriorly. CuA almost straight, single up to apical level of clavus, separated from MP and MP 3 + 4 by a wide area (open median cell) almost as wide as area width between MP + MP 3 + 4 and costal margin. C 5 cell open. CuA connected with parallel CuP before mid-length of tegmen by four transverse veins. Just before reaching clavus, apically forked with CuA 1 starts running parallel to the tegmen margin and CuA 2 joining tegmen margin. Pcu and A 1 connecting late in the last quarter of a long closed clavus. A 1 closely parallel to posterior margin. Traces of brownish coloration and small micro-granulations are visible in cubito-anal and subcostal areas (Fig. 1).	en	Bourgoin, Thierry, Wang, Menglin, Nel, André (2020): The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France). European Journal of Taxonomy 596: 1-8, DOI: 10.5852/ejt.2020.596
