taxonID	type	description	language	source
AF77302C642F9F6EBBCCFA31FAFFFDE0.taxon	description	Specimens examined Type materials of this species have been collected from Chile, Valdivia, Misituni. In Museum für Naturkunde Berlin two microslides (one female and one male) were collected from Chile: female specimen, ZMB Nr. 40148, Slide Number 4033, 12.01.1965; male specimen, ZMB Nr. 40147, Slide Number 4032, 10.24.1965. All other specimens designated as Hypoaspis eugenitalis deposited in Museum für Naturkunde Berlin; three alcoholic tubes with numbers of: ZMB Nr. 40135, Argentina, Alkoholpraparate (Karg-Nr.) 1183, 1965; ZMB Nr. 40136, Chile, Alkoholpraparate (Karg-Nr.) 1184, 1965 and ZMB Nr. 40137, Argentina, Alkoholpraparate (Karg-Nr.) 1185, 1961 contain 50, 2 and 2 mites respectively. The microslides: ZMB Nr. 40138, female, South America, Argentina, Slide Number 4018, 10.12.1961; ZMB Nr. 40139, female, South America, Argentina, Slide Number 4024, 10.30.1961; ZMB Nr. 40140, female, South America, Argentina, Slide Number 4025, 11.13.1961; ZMB Nr. 40141, male, South America, Argentina, Slide Number 4026, 11.13.1961; ZMB Nr. 40142, male, South America, Argentina, Slide Number 4027, 09.26.1961; ZMB Nr. 40143, male, South America, Argentina, Slide Number 4028, 10.12.1961; ZMB Nr. 40144, male, South America, Argentina, Slide Number 4029, 06.14.1961; ZMB Nr. 40145, female, South America, Argentina, Slide Number 4030, 07.18.1961; ZMB Nr. 40146, female, South America, Argentina, Slide Number 4031, 11.09.1961. The following slides based on their qualities have been used for drawings (the information presented here is based on the data on microslide labels): Hypoaspis eugenitalis Karg, female, Chel. No. 4018. Argentinien, Rio Negro, El Bolson, Votgeb. d. Piltriquitron, 460 m, Laubstreu, 12.10.61; Hypoaspis eugenitalis Karg, 1978, female, Chel. No. 4024. Argentinien, Rio Negro, El Bolson, Vorberg vom Mt. Piltriquitron, 400 m, Laubstreu, 30.10.61; Hypoaspis eugenitalis Karg, 1978; Chel. B. II, 4026; [ZMB Kat. Nr. 40141]; Male, Argentinien, Rio Negro, El Bolson, Berg, Piltriquitron, 1170 m, Moosiges, Gras vom Rande eines Nothofagus-Waldes, 13.11.61; Hypoaspis eugenitalis Karg, 1978, Male, No. 4029. Argentinien, Rio Negro, El Bolson, Piltriquitron, 480 m, Frisches Moos von Fels, 14.04.61. Diagnosis Setae r 6 located on soft cuticle, Zx 3 present, dorsal setae Z 4 - 5 and S 4 - 5 are more elongate than the others; epistome with fair denticles in lateral and anterior margins; bearing a supralabral process as a slender and pointed style; genitiventral shield reticulated throughout with pore-like structures on surface; the gd 6 pustule like; male holoventral shield bears several pore-like structures on the surface. Description (Female) Dorsal idiosoma (Fig. 1 A) – Dorsal idiosoma oval-shaped, (878 – 889) long, (623 – 650) wide, with scanty ornamentation throughout, holotrichous both in podonotal and opisthonotal parts; with 23 pairs of setae on podonotal region including of one pair of extra paired setae (zx) between z 2 – z 3, dorsal setae designated as j 1 – 6; z 1 – 6 and one pair of zx; s 1 – 6 and r 2 – 5. Opisthonotal area of shield with 16 pairs of setae designated J 1 – 5; Z 1 – 5; S 1 – 5 and one pair of Zx 3. setae z 1 is the shortest (Fig. 2 A), and S 5 the longest (Fig. 1 A). Most dorsal shield setae moderate or elongate. The sizes of dorsal shield setae are as follows: j 1: 60 – 63; j 2 – 5: 93 – 102; j 6: 70 – 73; z 1: 23 – 25; z 2 – 6: 89 - 93; s 1 – 6: 99 – 112; r 2: 77 – 80; r 3 – 5: 99 – 113; J 1 – 3: 70 – 74; J 4 -- 5: 102 – 120; Z 1: 129 – 132; Z 2: 80 – 84; Z 3: 120 – 124; Z 4: 150 – 168; Z 5: 155 – 162; S 1: 90 – 95; S 2 – 3: 100 – 109; S 4: 145 – 156; S 5: 185 – 197; Zx 3: 84 – 91. Dorsal shield with 19 pairs of pore-like structures, an enlarged pore like structure (pustule-like in appearance) which probably accompanied by gd 6 and is 1 at dorso-marginal side of opisthonotal region (bent down in ventral side of Fig. 1 B) present near the setae Z 1. Ventral idiosoma (Fig. 1 B) – Tritosternum with columnar base 65 – 72 long, 20 – 24 wide, and pilose laciniae 182 – 190 long. Pre-sternal plates well sclerotized with 68 – 72 long and 25 – 28 wide with fair striae on surface. Sternal shield (Figs. 1 B, 2 B, C) 141 – 145 long, 178 – 183 wide at level of setae st 2 and 352 – 358 at level of lateral projections between coxae II-III, ornamented throughout, except in medio-posterior area which is smooth, anterior and posterior margins of shield concave, lateral margins alongside coxae II-III fused with endopodal plates. Sternal shield with three simple acicular pairs of setae, st 1: 83 – 86, st 2: 105 – 108, st 3: 103 – 105; the distances between sternal setae: st 1 - st 1: 90 – 96, st 2 - st 2: 105 – 108, st 3 - st 3: 201 – 206, iv 1 – 2 slit-like, iv 1 located between st 1 setae, distance between iv 1 – iv 1 10 – 12, iv 2 located between st 2 – 3 almost at level of projection between coxae II-III, iv 2 - iv 2 191 – 199. Setae st 4 absent and lyrifissures iv 3 located on postero-corner of sternal shield at the tip of interior section of endopodal III-IV (Figs. 2 B, C). Genitiventral shield 462 – 475 long, 289 – 292 wide at level of genital setae, 451 – 460 wide at widest part, bearing three pairs of setae including genital setae, st 5: 100 – 103, Zv 1 – 2: 138 – 146, Jv 1 – 2: 69 – 92, paragenital lyrifissure (iv 5) located on the surface of podal plate at level of genital setae (Figs. 2 B, C). Genitiventral surface fairly reticulate and bears numerous of different sized pore-like structures. Metapodal plates only could be seen in some specimens and almost fused with lateral margins of genitiventral shield (Fig. 2 C). Anal shield subtriangular with anterior margin convex, 127 – 139 long, 224 – 231 wide; paranal setae acicular (43 – 49), post-anal seta (23 – 26), gland pores gv 3 on lateral margins of anal shield. Cribrum normally developed posteriorly and laterally around postanal seta. Stigma located at posterior level of coxa III, stigmatal plate ending in a poststigmatal extension of peritrematal shield (Fig. 1 B) bears two-three pore-like structures. Peritreme anteriorly extending to mid-level of coxa I, peritrematal shield at level of coxae II-III bearing gd 3 accompanied by id 3 and small shoulder shield present near coxa I. Podal, endo- and exopodal plates described in genus definition. Gnathosoma (Figs. 3 A – D) – Setae h 1 and h 2 (59 – 67), h 3 (69 – 73); palpcoxal seta (pc) 49 – 55 long. Deutosternal groove with seven multidenticulate rows. Corniculi horn-like, with 54 – 59 long, 6 – 10 wide at midlevel, internal malae complex, with two pairs of lobes, inner lobes slender and long (55 – 60), lateral lobes smaller (29 – 33), wider with short hairs, extending to the midlevel of curniculi (Fig. 3 A). Epistome (Fig. 3 B) acuminate and fairly denticulate, with prominent denticles at anterior and lateral margins. Supralabral process style like. Chelicerae (Fig. 3 C) – With setaceous dorsal seta (21 – 25), arthrodial process and lateral lyrifissure; movable digit 68 – 75 long medially from top to posterior margin, with two teeth, middle article (152 – 157) ending in fixed digit (70 – 79 long from top to level of dorsal seta), bearing two distal, one large tooth at midlevel and one smaller proximal tooth in addition to distal hook, small needle-like pilus dentilis alongside the larger cheliceral tooth. Palp segment lengths as follows: trochanter (55 – 61), femur (75 – 81), genu (47 – 55), tibia (39 – 44) and tarsus (23 – 27), palp chaetotaxy (Fig. 6 D) normal (sensu Evans & Till 1965), with smooth acicular setae, palp-tarsal claw two-tined (Fig. 3 D). Labrum (Fig. 3 A) slender and pubescence. Legs (Fig. 4, 6 A – D) – Tarsi I-IV with developed claws and ambulacra, female and male specimens with similar legs setae (Figs. 6 A, 6 C, D) except legs II (Figs. 4, 6 B). Leg I (Fig. 6 A) 761 – 802 long, coxa (88 – 95), trochanter (60 – 65), femur (161 – 164), genu (107 – 116), tibia (114 – 129), tarsus (231 – 233); leg II (Fig. 4) 535 – 669 long, coxa (51 – 61), trochanter (86 – 91), femur (95 – 116), genu (84 – 86), tibia (74 – 82), tarsus (145 – 166); leg III (Fig. 6 C) 529 – 588 long, coxa (53 – 55), trochanter (80 – 89), femur (110 – 113), genu (75 – 90), tibia (63 – 75), tarsus (148 – 166); leg IV (Fig. 6 D) 666 – 801 long, coxa (52 – 69), trochanter (95 - 106), femur (121 – 163), genu (81 – 103), tibia (95 – 109), tarsus (222 – 251). All leg setae smooth, some spur or spine-like as in Figures 4, 6 B. Chaetotaxy of legs is as same as in genus (see above). The situation of setae thickness in male and female is similar in all legs except leg II which separately figured in both sexes. Leg I (Fig. 6 A): In trochanter al and in femur pd 2 - 3 slightly thicker than the other setae on the segment. Leg II (Fig. 4): in tarsus setae mv, av 2, pv 2, al 1, and md spine-like and thicker than other setae on the segment; setae av 1 and pv 1 strong spine-like. Leg III (Fig. 6 C): in tarsus setae mv, av 1 - 2, pv 1, al 1 and pl 1 thicker than the others on the segment. Leg IV (Fig. 6 D): femur (ad 2 and pd thicker than the others on the segment) and genu (pl slightly thicker than other setae on the segment). Male (Figs. 5 – 6). Dorsal idiosoma – Dorsal idiosomal length 729 – 734 width 491 – 545. Dorsal shield chaetotaxy similar to female with slightly smaller setae. Ventral idiosoma (Fig. 5 A) – With one pair of presternal plates with striae on surface located at either sides of genital orifice at anterior middle part of holoventral shield. Holoventral shield with ornamentation observed in anterior part, 152 – 163 wide in area of st 2, expanded posterior to coxae IV and created the widest part (389 – 403 wide posterior to coxae IV) then tend to be irregularly pointed toward the anus. Numerous pore-like structures present on the surface of the shield at level of st 3 to area posterior to coxae IV (Fig. 5 A). The shield bearing 10 - 11 pairs of smooth acicular setae. Lyrifissures iv 1 – 2 slit-like located at base of st 1 and between st 2 - st 3 respectively, iv 3 slit-like between st 3 - st 4. Endopodals fused with holoventral shield from anterior level of coxae II to area adjacent to midlevel of coxae IV. Stigmatal opening surrounded by wide peritrematal shield. Posterior extension bear three pore-like structures [is 1, gp 3, ip 3 following Moraza and Lindquist (2018)]. The broad peritrematal plate with widest area at level of coxae II-III, bearing gp 2 and ip 2 following Moraza and Lindquist (2018). Peritreme long, extended to anterior margin of coxa I. Gnathosoma – Movable digit (170 – 175) of chelicera with one large tooth (Fig. 5 B), spermatodactyl (260 – 266 long: from anterior tip to the attachment place to movable digit) much longer than movable digit, bent apically over it, with a small pore-like structure at distal bend which connected to a longitudinal tube inside the spermadactyl extended to a small fossa basally at basal part of movable digit (Fig. 5 C). Fixed digit with two teeth. Dorsal seta, cornet-shape setae, lyrifissure and setaceous pilus dentilis are present. Legs (Figs. 6 A – D) – Tarsi I-IV with developed claws and ambulacra, legs setae of male specimens are similar to female except legs II (Figs. 6 B). Leg I (Fig. 6 A) 738 – 788 long, coxa (90 – 98), trochanter (50 – 60), femur (158 – 165), genu (110 – 113), tibia (122 – 134), tarsus (208 – 218); leg II (Fig. 6 B) 559 – 615 long, coxa (51 – 61), trochanter (87 – 90), femur (101 – 128), genu (85 – 89), tibia (85 – 92), tarsus (150 – 155); leg III (Fig. 6 C) 473 – 506 long, coxa (53 – 54), trochanter (66 – 68), femur (86 – 98), genu (64 – 70), tibia (60 – 69), tarsus (144 – 147); leg IV (Fig. 6 D) 651 – 725 long, coxa (56 – 61), trochanter (78 – 103), femur (128 – 145), genu (92 – 96), tibia (90 – 93), tarsus (207 – 227). All leg setae smooth, some spine-like as in Figs. 6 B, C. Chaetotaxy of legs is as same as in genus (see above). The chaetotaxy and setae thickness in male and female legs are similar except legs II. Seta pv 1 in male femur II is strong spur-like, av 1 and pv 1 in tarsus II are strong spine-like setae (Fig. 6 B).	en	Nemati, Alireza, Gwiazdowicz, Dariusz J., Riahi, Elham (2021): A new genus of Laelapidae (Acari: Mesostigmata) from South America. Persian Journal of Acarology 10 (2): 167-189, DOI: 10.22073/pja.v10i2.64420, URL: http://zoobank.org/738a6baa-9164-479a-9950-775a3eee4599
AF77302C64269F75BBC4FF39FEAFFEC3.taxon	description	Specimen examined	en	Nemati, Alireza, Gwiazdowicz, Dariusz J., Riahi, Elham (2021): A new genus of Laelapidae (Acari: Mesostigmata) from South America. Persian Journal of Acarology 10 (2): 167-189, DOI: 10.22073/pja.v10i2.64420, URL: http://zoobank.org/738a6baa-9164-479a-9950-775a3eee4599
AF77302C64269F75BBC4FF39FEAFFEC3.taxon	description	While the above-mentioned slide was the only one in Museum für Naturkunde Berlin but marked as a paratype. Moreira (2014) recorded the type depository of this species in Naturwissenschaftlichen Museum, Budapest, Hungary. On the other hand, Karg (1979) himself has mentioned some ranges for some of the traits measurements of this species in its original description, which probably indicates that this species has more than one slide. Diagnosis Setae Zx absent; dorsal setae not elongate (33 – 63); genitiventral shield well expanded posterior to coxae IV, reticulated at anterior part; epistome with fair denticles in anterior margin, supralabral process wider and similar to long style. Description (Female) Dorsal idiosoma (Fig. 7 A) – Dorsal shield oval-shaped, 753 long, width at level of setae r 3 556; avoid ornamentation throughout, shield with 38 pairs of thin and simple setae, 23 pairs on podonotal region including j 1 – 6; z 1 – 6; s 1 – 6; r 2 – 5 and one extra paired zx setae between z 3 - z 4; and 15 pairs on opisthonotal part encompasses of J 1 – 5, Z 1 – 5, S 1 – 5; setae Zx 2 – 3 absent, the distance of J 2 - J 2 is about twice of J 1 - J 1 distance. Dorsal setae varies in length: in podonotal the setae z 1 shortest (13), the others 37 – 63; in opisthonotal area 33 – 57. Cuticle between dorsal and ventral side of body bearing r 6 at ventral side. Podonotal and opisthonotal regions with 11 pairs of discernable pore-like structures, as shown in Fig. 7 A. It is worth mentioning that the dashed lines in Fig. 7 A only indicate the location of the folded parts in the representative specimen on the slide. Obviously, the setae and the related discernible pore and pore-like structures have been displayed on the dorsal surface. An enlarged pore like structure (pustule-like in appearance) which probably accompanied by gd 6 and is 1 present at dorso-ventral side of opisthonotal near the setae Z 1. Ventral idiosoma (Fig. 7 B) – Tritosternum bearing columnar base with 27 long, 12 wide at middle part and 28 basally, the laciniae was broken and not observed. Pre-sternal with a pair of slightly sclerotized plates with striae on surface. Sternal shield with reticulation except in posterior surface, 111 long, 168 wide at level of st 2 and 286 at level of projection between coxae II-III, slightly concave at anterior margin and deeply at posterior. Sternal setae smooth, st 1 (59), st 2 (74) and st 3 (83), the distances of st 1 - st 1 (78), st 2 - st 2 (123), st 3 - st 3 (164), st 1 - st 2 (59), st 2 - st 3 (53); iv 1 – 2 slitlike, located slightly behind st 1 and between st 2 - st 3 respectively. Metasternal setae absent; iv 3 ovoid, located on postero-corner of sternal shield. Genitiventral shield well expanded posterior to coxae IV, extending to area adjacent to anterior margin of anal plate, reticulated at anterior surface, bearing several pore-like structures on the surface (Fig. 8), with 368 long, 233 wide at level of st 5, 360 at level of Zv 1 in widest part. Genitiventral shield bearing three pairs of setae, including st 5 (85), Zv 1 (81) and Jv 1 (66), the other setae on opisthogastric area consist of Zv 2 (48) and Jv 2 – 5 (35 – 47). Paragenital pores on podal plate surface at posterior part of coxae IV, five other pore-like structures observed on opisthogastric area. Anal shield not reticulate, with semi-circular anterior margin, 103 long, 172 wide (at widest part), paranal setae (47) longer than post-anal seta (27). Cribrum with prominent spicules, extending laterally almost to level of post-anal seta. Suboval metapodal plate observed at left side of genitiventral shield. Stigmata located at area between coxae III-IV, surrounded by almost wide stigmatal plate. Peritremes wide and long, extending to middle part of coxa I, separated from exopodal shield. Poststigmatal part extending nearly from anterior part of coxa IV to its posterior level. Exopodal and endopodal plates (Fig. 9) similar to H. eugenitalis as explained above.	en	Nemati, Alireza, Gwiazdowicz, Dariusz J., Riahi, Elham (2021): A new genus of Laelapidae (Acari: Mesostigmata) from South America. Persian Journal of Acarology 10 (2): 167-189, DOI: 10.22073/pja.v10i2.64420, URL: http://zoobank.org/738a6baa-9164-479a-9950-775a3eee4599
AF77302C64269F75BBC4FF39FEAFFEC3.taxon	materials_examined	Gnathosoma – Hypostome (Fig. 10 A) with three pairs of setae; only h 1 (48) could be observed in specimen examined (see a note under material examined). Palpcoxal setae smooth with 61 long. Deutosternal groove with six rows of 5 – 10 denticles, the anteriormost is smooth, first row of denticles arc medially. Corniculi normal, horn-like; internal malae consists of two toothed external appendages and two long and thread-like internal appendages. Epistome cup-like, rounded and fairly denticulate medially while smooth laterally (Fig. 10 B), with slender style underside. Labrum slender and slightly pubescent. Chelicerae (Fig. 10 C) arthrodial processes developed, moveable digit (89) with two teeth, middle article (157: from cheliceral dorsal seta to its posterior margin) ending in fixed digit (80), bearing a big tooth proximally behind setaceous pilus dentilis and three smaller teeth distally in addition to distal hook. Palp (Fig. 10 D) – Chaetotaxy normal (sensu Evans & Till 1965), with simple and thin setae, palp-tarsal claw two-tined, basal tine smaller (not in normal view in Fig. 10 D probably due to preparation condition).	en	Nemati, Alireza, Gwiazdowicz, Dariusz J., Riahi, Elham (2021): A new genus of Laelapidae (Acari: Mesostigmata) from South America. Persian Journal of Acarology 10 (2): 167-189, DOI: 10.22073/pja.v10i2.64420, URL: http://zoobank.org/738a6baa-9164-479a-9950-775a3eee4599
AF77302C64269F75BBC4FF39FEAFFEC3.taxon	description	Legs – Tarsi I-IV with claws and ambulacra. Leg I 738 μm, coxa (93), trochanter (59), basifemur (36), telo-femur (120), genu (107), tibia (118), tarsus (205 μm); leg II 590 μm (excluding stalk and pretarsus), coxa (44), trochanter (87), femur (114), genu (102), tibia (90), basi-tarsus (29), telotarsus (124); leg III 497 μm (excluding stalk and pretarsus), coxa (51), trochanter (78), basi-femur (40), telo-femur (60), genu (65), tibia (64), basi-tarsus (22), telo-tarsus (117); leg IV 646 μm (excluding stalk and pretarsus), coxa (48), trochanter (87), basi-femur (36), telo-femur (89), genu (92), tibia (75), basi-tarsus (43), telo-tarsus (176). Legs I and IV longer than legs II and III. All leg setae smooth and pointed. Legs chaetotaxy is similar to genus condition. Leg I (Fig. 11 A); leg II (Fig. 11 B): tarsus (mv pointed and slightly thickened, av 2 and pv 2 thickened and spine-like, av 1 and pv 1 thickened and spur-like); leg III (Fig. 11 C): tarsus (mv, av 1 - 2, pv 1 - 2 slightly thickened); leg IV (Fig. 11 D). Insemination structures – Not seen. Male Unknown.	en	Nemati, Alireza, Gwiazdowicz, Dariusz J., Riahi, Elham (2021): A new genus of Laelapidae (Acari: Mesostigmata) from South America. Persian Journal of Acarology 10 (2): 167-189, DOI: 10.22073/pja.v10i2.64420, URL: http://zoobank.org/738a6baa-9164-479a-9950-775a3eee4599
