taxonID	type	description	language	source
B44C6E19FFF1FFE7FC0BC25FFB25FD7C.taxon	description	(Figs. 1 A, 2, 7 A, B)	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF1FFE7FC0BC25FFB25FD7C.taxon	materials_examined	Material examined. 10 males (CL: 4.4 – 4.8 mm, CW: 5.9 – 6.5 mm), 7 females (CL: 2.9 – 4.3 mm, CW: 4.0 – 5.8 mm), LFSC. ZRC- 185, India, Goa, Chapora Estuary (15 ° 37 ’ 21 ” N 73 ° 44 ’ 45 ” E), coll. Mithila Bhat, 13 April 2016. Diagnosis. Carapace quadrangular, glabrous, 1.34 times wider than long; lateral margins slightly concave, subparallel posteriorly; dorsal surface covered with scattered pits, setal tufts and sparsely placed setae; protogastric and mesogastric regions delimited by inconspicuous grooves. External orbital tooth broadly triangular followed by rudimentary epibranchial tooth. Several oblique, low, setose granulated striations extending across branchial regions (Figs. 1 A, 2 A). Frontal margin wide, indistinctly bilobed, frontal lobes separated by shallow depression; 4 inconspicuous postfrontal lobes covered with setose patches, separated by longitudinal grooves (Fig. 2 A). Basal antennal article long (Fig. 2 B), reaching distal margin of first article. Third maxillipeds with distinct rhomboidal gap when closed; ischium subrectangular, slightly longer than wide, 1.27 times longer than merus, with oblique setose ridge; merus subovate, 1.01 times as wide as long, with oblique setose ridge. Exopod slender, flattened, with long flagellum (Fig. 2 C). Chelipeds subequal, massive; palm massive (Fig. 2 D), surfaces rugose, dorsal surface with two oblique pectinate crests and 4 or 5 oblique granular ridges (Fig. 2 E); outer surface pitted, covered with discreet granular striations on posterior half and scattered granules anteriorly, thin granular line on lower half commencing on pollex and extending posteriorly to carpal junction; inner surface sparsely granulated. Fingers (dactylus and pollex) massive, curved, with rounded spatulate tips, gaping when closed. Male cheliped dactylus short, with row of 13 or 14 small smooth transverse tubercles and proximal granules on dorsal surface; inner surface with proximal granular patch. Cutting edge of dactylus with 3 large and 2 small teeth, and large subdistal tooth, pollex with large subdistal tooth, followed by 3 small, 1 large median teeth and 4 proximal teeth (Fig. 2 D). P 2 – 5, compressed, shorter than chelipeds. P 2 short, P 3 and P 4 subequal, P 5 short. P 4 and P 5 meri flattened, wider distally. Carpi ventral surfaces with median longitudinal granular ridge. P 4 and P 5 carpi and propodi without dense setae. Propodi about twice as long as wide, posterior margins with distal spine and few long bristles, ventral surface with inconspicuous oblique groove. Dactyli smooth, tip corneous, slightly curved, dorsal margin with 2 pairs of slender subdistal spines and ventral margin with 1 pair of subdistal spines (Fig. 2 F). Male pleon narrow, bluntly triangular, surface pitted; lateral margins finely granulated, densely setose; telson 1.03 times as long as wide, 1.69 times as long as pleomere 6 (Fig. 2 G). Female pleon wide, rounded, pleomere 6 with concave distal margin; telson pentagonal, wider than long, partially sunken into distal margin of pleomere 6 (Fig. 2 H). G 1 long, straight, distal part corneous, flattened, slightly curved, medially notched, flanked by thick bristles, groove at base of tip extends along ventral surface to G 1 (Fig. 7 A, B). Remarks. The specimens examined in the present study resemble De Man’s (1888) description of his Sesarma andersoni, differing in the presence of a distinct subdistal spine on the anterior margin of P 2 – P 5 meri. Morphological comparison with other Indian congeneric species is provided in Tab. 1. Distribution. Nanosesarma andersoni was originally described from Mergui Archipelago (De Man, 1888; Alcock, 1900), and later reported from Malaysia, Singapore (Tweedie, 1950), Japan (Komai et al., 2004), and India (Trivedi et al., 2018). In India, it is known from Goa (Dev Roy and Bhadra, 2008; present study), Kerala (Devi et al., 2015), Tamil Nadu (Ravichandran and Kannupandi, 2007), and Andaman and Nicobar Islands (Dev Roy and Das, 2000).	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF4FFE5FCF1C706FC27F80F.taxon	description	(Figs. 1 B, 3, 7 C, D)	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF4FFE5FCF1C706FC27F80F.taxon	description	Soh, 1970: 394. Material examined. 1 male (CL: 4.8 mm, CW: 6.6 mm), GUMSIS 1, India, Goa, Verem (15 ° 29 ’ 74 ” N 73 ° 48 ’ 20 ” E), coll. Mithila Bhat, 9 April 2014. Description. Carapace quadrangular, 1.35 times wider than long, with slightly concave, converging lateral margins; dorsal surface covered with scattered pits, prominent setal tufts on postfrontal lobes, protogastric and anterior branchial regions, and sparsely placed setae on posterior half of carapace. External orbital tooth triangular. Several oblique, low, setose granulated striations extending across branchial regions (Figs. 1 B, 3 A). Frontal lobes separated by shallow depression; 4 inconspicuous post-frontal lobes covered with setose patches, separated by longitudinal grooves (Fig. 3 A). Third maxillipeds with distinct rhomboidal gap when closed; ischium subrectangular, 1.17 times longer than wide, 1.59 times longer than merus, with oblique setose ridge; merus subovate, 0.89 times longer than wide, with oblique setose ridge. Exopod slender, flattened, with long flagellum (Fig. 3 B). Right cheliped massive, longer than carapace; palm massive (Fig. 3 C), surfaces rugose, dorsal surface with 2 oblique pectinate crests and 5 or 6 oblique granular ridges (Fig. 3 D); outer surface smooth, slightly pitted, thin granular line on lower half commencing on pollex and extending posteriorly to carpal junction; inner surface sparsely granulated. Fingers massive, curved, with rounded spatulate tips, gaping when closed, outer surface with dense setose patch. Male dactylus with row of 12 prominent transverse ridges, proximal granules and scattered setae on dorsal surface; inner surface with proximal granular, setose patch. Cutting edges of dactylus and pollex with 4 subequal blunt teeth each (Fig. 3 C). P 2 – 5, compressed, shorter than chelipeds. P 2 and P 5 short, P 3 and P 4 subequal. P 2 and P 3 propodi and carpi densely setose on anterior surface, P 4 – P 5 propodi and carpi without dense setae. P 2 – P 5 propodi posterior margins with one distal spine, ventral surface with inconspicuous oblique groove. P 5 dactylus smooth, tip corneous, slightly curved, dorsal margin with six spines, ventral margin with three pairs of spines (Fig. 3 E). Male pleon bluntly triangular (AW / CW = 0.38), surface pitted, sutures with sparsely setose patches; lateral margins finely granulated, densely setose; telson 1.04 times as long as wide, 1.88 times as long as pleomere 6 (Fig. 3 F). G 1 long, straight, distal part corneous, slender, f lattened, slightly bent, lined by thick bristles, groove at base of tip extends along ventral surface to proximal portion of G 1 (Fig. 7 C, D). Remarks. The specimen examined in the present study agrees with the description by De Man (1890) but differs in having 6 oblique granular ridges on dorsal surface of male cheliped propodus (vs. 7 or 8 in the holotype), and 4 occlusal teeth on pollex (vs. 3 in the holotype). Morphological comparison with other Indian species is provided in Tab. 1. Geographical distribution. Nanosesarma batavicum was originally described from Batavia, Indonesia and later reported from Java, Malaysia, Singapore, (Tweedie, 1950) and India (Trivedi et al., 2018). In India, it is known from Goa (present study), Kerala (Devi et al., 2015), Tamil Nadu (Varadharajan and Soundarapandian, 2014), and Odisha (Dev Roy and Rath, 2017).	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF7FFEBFF7DC1BDFABFFA19.taxon	description	(Figs. 1 C, 4, 7 E, F)	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF7FFEBFF7DC1BDFABFFA19.taxon	materials_examined	Material examined. 10 males (CL: 2.4 – 4.6 mm, CW: 3.1 – 5.9 mm), 7 females (CL: 2.6 – 4.9 mm, CW: 3.3 – 6.2 mm), LFSC. ZRC- 186, India, Goa, Chapora Estuary (15 ° 37 ’ 21 ” N 73 ° 44 ’ 45 ” E), coll. Mithila Bhat, 13 May 2016. Diagnosis. Carapace quadrangular, 1.26 wider than long; lateral margins almost straight, gradually diverging posteriorly; dorsal surface, strongly rugose, slightly convex, covered with scattered pits and sparsely placed setae only on front, protogastric, mesogastric and branchial regions delimited by conspicuous depressions. External orbital tooth truncate and triangular, followed by sharp epibranchial tooth. Several oblique, low, setose granulated striations extending across branchial regions (Figs. 1 C, 4 A). Frontal margin bilobed, strongly declivous, thickly setose, at level of eyes, frontal lobes separated by shallow depression; 4 inconspicuous postfrontal lobes covered with setose patches, separated by longitudinal grooves. Ocular peduncle well-developed, slightly setose, cornea terminal (Fig. 4 A). Third maxillipeds with distinct rhomboidal gap when closed; ischium subrectangular, 1.38 ± 0.19 times longer than wide, 1.18 ± 0.10 times longer than merus, with oblique setose ridge; merus 1.01 ± 0.11 times longer than wide, with oblique setose ridge. Exopod slender, f lattened, with long f lagellum (Fig. 4 B). Male chelipeds subequal, massive, longer than carapace; palm massive, surfaces rugose, dorsal surface without oblique pectinated crest, 3 inconspicuous oblique granulated ridges; outer surface with densely setose patch extending anteriorly to proximal portion of fingers, pitted, prominent granular ridge on lower half commencing on pollex and extending posteriorly up to carpal junction, 4 rows of closely spaced granules above granular ridge; inner surface sparsely granulated. Fingers massive, curved, with rounded spatulate tips, gaping when closed. Cheliped dactylus of male short with proximal setose granular patch on dorsal surface, transverse ridges absent; inner and outer surfaces with proximal setose granular patch. Cutting edge of dactylus with large subdistal tooth, followed by 2 small median and 2 large proximal teeth, that of pollex with large subdistal tooth, followed by 2 small teeth, large median tooth and 3 or 4 small teeth (Fig. 4 C). Pereopods compressed, shorter than chelipeds. P 2 and P 5 short, P 3 and P 4 subequal. P 4 and P 5 meri f lattened, anterior margin terminating in blunt angle followed by subdistal spine; postero-distal margin with minute distal tooth-shaped granules followed by one spinule. P 2 – P 5 propodi and carpi with long bristles on anterior margins. Propodi about twice as long as wide, posterior margins with 2 distal spines and few long bristles. Dactyli smooth, tip corneous, slightly curved, with 4 slender subdistal spines (Fig. 4 D). Male pleon narrow (PW / CW = 0.40 ± 0.04), bluntly triangular, surface pitted; lateral margins finely granulated, densely setose; telson 1.17 times as long as wide, 2.00 times as long as pleomere 6 (Fig. 4 E). Female pleon wide (AW / CW = 0.72 ± 0.10), rounded, pleomere 6 withroundeddistalmargin, marginsdensely setose; telson wider than long, almost completely sunken into distal margin of pleomere 6 (Fig. 4 F). G 1 long, straight, corneous distal part curved, long scoop-shaped, flanked by stiff bristles, groove at base of tip extends along ventral surface to G 1 base (Fig. 7 E, F). Remarks. The morphology of the present specimens agree with that reported by De Man (1887), Rathbun (1910), Tweedie (1950), and Crosnier (1965). However, it differs in having the presence of stiff distal setae on the G 1 tip, which was probably omitted in the earlier reports. Morphological comparison with other Indian congeneric species is provided in Tab. 1. Distribution. Nanosesarma minutum was originally described from Edam Island near Jakarta, Indonesia and later reported from Malaysia, Singapore, Gulf of Thailand, South China Sea (Tweedie, 1936, 1950), Madagascar (Crosnier, 1965), Tanzania (Hartnoll, 1975), and India (Trivedi et al., 2018). In India, it is known from Gujarat (Saravanakumar et al., 2007), Maharashtra (Pati et al., 2012), Goa (present study), and Tamil Nadu (Kathirvel and Gokul, 2010).	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF8FFE9FCD6C1F8FAC4F9AF.taxon	description	(Figs. 1 D, 5, 7 G, H)	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFF8FFE9FCD6C1F8FAC4F9AF.taxon	materials_examined	Material examined. 3 males (CL: 4.6 – 5.6 mm, CW: 5.4 – 6.6 mm), LFSC. ZRC- 188, India, Gujarat, Shivrajpur (21 ° 19 ’ 55 ” N 68 ° 57 ’ 02 ” E), coll. Jigneshkumar Trivedi, 3 February 2016. Diagnosis. Carapace quadrangular, 1.13 – 1.17 times wider than long, widest at the level of second anterolateral teeth, evenly convex, sparsely plumose; lateral margins slightly concave, slightly converging posteriorly, dorsal surface covered with setal tufts and sparsely placed short and thin plumose setae, well-defined gastric region contained by deep groove, cardiac region with shallow groove. External orbital tooth sharply triangular, followed by smaller, slightly pointed epibranchial tooth. Dorsal surface finely granulated, 2 transverse granulated ridges originating from lateral margin, almost parallel to posterolateral margin, first ridge emerging just after epibranchial tooth, extending towards central region of carapace, second ridge originating near end of lateral margin, terminating at posterolateral edge, further 2 lessprominent, short ridges present, one emerging almost from the V-shaped notch between the anterolateral teeth, moving subparallel to first prominent ridge, other below first prominent one, nearly parallel to it (Figs. 1 D, 5 A). Basal antennal article long (Fig. 5 B), reaching to about half of second article, antennae flagellum short, reaching cornea. Third maxillipeds with distinct rhomboidal gap when closed, ischium subrectangular, 1.37 times longer than merus, internal margin crenulated, with long setae; merus subovate 1.18 times longer than wide, with longitudinal furrow on outer surface, margins crenulated, internal margin with long setae. Exopod slender, with long flagellum (Fig. 5 C). Chelipeds subequal; palm with entire outer surface covered with dense setae above lower row of granules, extending anteriorly to midlength of fingers (Fig. 5 D), 3 rows of granules, lowermost regularly granulated, extending to centre of pollex, middle one short, irregularly granulated, subparallel to lower one, uppermost short, irregularly granulated, upcurving, upper margin with crenulated longitudinal line, dorsal surface without oblique pectinate crest or granulated ridges. Dactylus (except tip) with dorsal surface completely tuberculated, densely set small tubercles on proximal part with 6 comparatively large granules, all covered by tufts of setae, cutting edge with small teeth. Pollex short, smooth, cutting edge with several small teeth and large prominent one medially, finger-tips spatulate (Fig. 5 E). Pereopods medium-sized, P 2 and P 5 short, P 3 and P 4 longest, anterior meral margins weakly crenulated, bearing pointed subdistal tooth, posterior margins crenulated, posterodistal lobe denticulate. Propodi usually longer, about twice as long as wide with oblique carina on posterior face, 2 rows of long setae along propodus on anterior and posterior margins, P 2 – P 4 with small brown setae on anterior margin of propodus, setae prominent and dense on P 2. Dactyli distinctly more than half as long as propodus, with 8 – 10 spines, in 2 rows on posterior margin, 2 or 3 spines on anterior margin of P 5, female P 5 without any spine on anterior margin (Fig. 5 F). Male pleon elongated, triangular, smooth, pleomere 6 almost twice as broad as long, telson markedly elongate, about 1.60 times as long as broad (Fig. 5 G). G 1 long, nearly straight, corneous distal part curved outwards with long stiff setae, groove at proximal part, with terminal opening on lateral side (Fig. 7 G – H). Remarks. The specimens examined in the present study agree with the holotype description of N. sarii provided by Naderloo and Türkay (2009). The morphological comparison of N. sarii with other species of Nanosesarma reported from India is provided in Tab. 1. Distribution. Nanosesarma sarii is previously known from Persian Gulf, Gulf of Oman, and Pakistan (Naderloo and Türkay, 2009). The present specimens represent the first record from India.	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFFAFFEFFCF1C269FB7BF894.taxon	description	(Figs. 1 E, 6, 7 I, J)	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
B44C6E19FFFAFFEFFCF1C269FB7BF894.taxon	materials_examined	Material examined. 5 males (CL: 2.71 – 4.82 mm, CW: 3.12 – 5.55 mm), 12 females (CL: 5.43 – 5.41 mm, CW: 4.08 – 6.24 mm), LFSC. ZRC- 187, India, Gujarat, Shivrajpur (21 ° 19 ’ 55 ” N 68 ° 57 ’ 02 ” E), coll. Jigneshkumar Trivedi, 3 February, 2016. Diagnosis. Carapace quadrangular, 1.1 – 1.15 times wider than long, lateral margins almost straight, converging posteriorly; dorsal surface smooth, covered with setal tufts and long setae sparsely set among shorter plumose setae, regions well defined, front slightly protruded, bilobed with a shallow depression, extending to gastric region, straight depression separating gastric and cardiac regions, cardiac region with 2 small elevations, separated from intestinal region by a shallow depression, weakly granulated oblique ridge on posterior surface of carapace, extending from lateral margin to posterior edge. External orbital tooth triangular, followed by small, blunt epibranchial tooth (Figs. 1 E, 6 A). Basal antennal article (Fig. 6 B) long, reaching to base of flagellum. Third maxilliped morphology identical to that of N. sarii; ischium 1.33 times longer than merus, internal margin crenulated, possess long setae, merus 1.26 times longer than wide, with crenulated longitudinal margins, internal margin with long setae. exopod slender, with long flagellum (Fig. 6 C). Chelipeds distinctly unequal, palm relatively large, dorsal surface without oblique pectinate crest or granulated ridges, outer surface bearing 3 rows of tubercles fully covered with dense setae extending anteriorly to proximal portion of fingers (Fig. 6 D), lower one with small tubercles, extending to basal third length of pollex, middle one shortest, oblique, with relatively large tubercles, upper row of granules short, nearly parallel to median one. Upper margin with sparse tubercles, some tubercles present on upper part. Fingers shorter than palm, with a large gap in proximal half, dactylus straight, dorsal surface completely smooth, small proximal granules with a large distinct tubercle on outer surface, distal half of cutting edge with 6 or 7 low tubercles, distalmost larger, pollex cutting edge with large prominent tubercle in middle part, tips of fingers spatulated (Fig. 6 E). Pereopods medium-sized, all articles except dactylus, densely covered with short plumose setae, long bristles between them. Meral posterior margins with distinct tooth-shaped granules, posterodistal margin with spiniform teeth, proximal one largest. Carpal anterior and posterior margins with longitudinal line of small tubercles, P 5 without such line on posterior surface of carpus. Propodi smooth, 1.5 times as long as wide, posterodistal end with 2 groups of 3 small spines. Dactylus ending in large spine; 6 spines in 2 rows along posterior margin (Fig. 6 F). Male pleon bluntly triangular, pleomere 6 about 3.5 times as broad as long, lateral margins gently converging distally, telson 0.97 times as long as wide, rounded distally (Fig. 6 G). Female pleon large, round, pleomere 6 longer than others, telson small, inserted into pleomere 6 (Fig. 6 H). G 1 long, straight, distal part long, gently bending outwards, covered with long setae, posterior surface concave, groove nearly straight, terminal opening in posterior apical part. Some plumose setae along lateral margin (Fig. 7 I – J). Remarks. The specimens examined in the present study agree with the descriptions and illustrations of N. jousseaumei and N. sarii as discussed in Naderloo and Türkay (2009). The morphological comparison of N. jousseaumei with other species of Nanosesarma reported from India is provided in Tab. 1. Distribution. Nanosesarma jousseaumei was originally described from Gulf of Aden (Nobili, 1906), and later reported from Madagascar, Dar es Salaam, Persian Gulf (Naderloo and Türkay, 2009), Mayotte (Poupin et al., 2018), and India (Trivedi et al., 2018). In India, it is known only from Gujarat (Ghosh, 2004; Dev Roy, 2013; present study).	en	Trivedi, Vinay P. Padate Krupal J Patel Chandrashekher U. Rivonker Jigneshkumar N. (2022): On Indian species of Nanosesarma Tweedie, 1950 (Decapoda: Brachyura: Sesarmidae). Nauplius (e 2022031) 30: 1-18, DOI: 10.1590/2358-2936e2022031, URL: https://doi.org/10.1590/2358-2936e2022031
