taxonID	type	description	language	source
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	materials_examined	Holotype. SPAIN: Segovia: Carbonero el Mayor, 2 km SW C. las Tejeras to Rio Eresma, 41.1082 ° N / 4.2927 ° W, 18.7.2021, ♀ (leg. T. Wood). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. BULGARIA: Blagoevgrad: Sandanski, 17.7.1966, 1 ♀ (leg. M. Kocourek). GREECE: Aegean Islands: Samos, Kerkis (Vigla), 1200 – 1440 m, 11.7.1994, 1 ♀ (leg. A. W. Ebmer); Thasos, Potos, 40.613 ° N / 24.6195 ° E, 17.6.2012, 1 ♀ (leg. M. de Courcy); Peloponnese: Mani, Kap Tenaro, 6.6.1996, 1 ♀ (leg. W. Arens). TURKEY: Antalya: Karamanbeli pass, 18 km S Korkuteli, 1290 m, 14.7.1994, 1 ♂ (leg. O. Berg). SPAIN: Segovia: Carbonero el Mayor, 2 km SW C. las Tejeras to Rio Eresma, 41.1082 ° N / 4.2927 ° W, 18.7.2021, 11 ♀ (leg. T. Wood). Paratypes deposited in the Entomological Collection of ETH Zurich and the private collection of T. Wood.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	diagnosis	Diagnosis. Among the Western Palaearctic Heriades (Michenerella) species, the females of Heriades clypeonitida (Fig. 1) are recognisable by the following combination of characters: i) punctation of clypeus very scattered with interspaces reaching the diameter of several punctures (Fig. 3); ii) head slightly longer than wide (Fig. 2); and iii) absence of short and dense pilosity on the apical two thirds of the mandible (Fig. 3). The males differ by the following combination of characters: i) head about as long as wide; ii) tarsal segments 2 – 3 of hind leg asymmetrical; iii) hair band at apical margin of sternum 3 narrow and extending only over about one fourth of sternal width; and iv) curved apical part of gonoforceps long, finger-like, more slender than adjacent more basal part and apically with tuft of few long white hairs (Fig. 5, 6).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	description	Description. FEMALE (Fig. 1): Body length 4.5 – 5.5 mm. Head: Head 1.02 – 1.04 × as long as wide (Fig. 2). Distance between posterior margin of lateral ocellus and preoccipital margin 1.7 – 2.0 × as long as ocellar diameter. Maximum width of genal area 0.8 – 1.0 × as long as maximum width of compound eye. Proboscis short, reaching in repose not beyond proboscidial fossa. Second segment of labial palpus 2.0 – 2.2 × as long as first segment and 0.46 – 0.49 × as long as compound eye. Mandible three-toothed and black except for preapical reddish zone, without short and dense pilosity on its apical two thirds (Fig. 3). Clypeus distinctly convex and in its apical fourth strongly bent towards labral base, its punctation fine, shallow and very scattered with interspaces reaching diameter of several punctures (Fig. 3). Paraocular area covered with dense white pilosity (Fig. 2). Punctation of supraclypeal area dense except for median zone, where interspaces may reach diameter of one to two punctures (Fig. 2). Punctation of frons very dense with interspaces rarely exceeding diameter of half a puncture, punctures slightly larger than those on supraclypeal area (Fig. 2). Antenna predominantly blackish, anterior side of flagellum more or less brownish. Antennal segments 3 – 5 (6) shorter than wide, segments (6) 7 – 11 as long as wide to slightly longer and segment 12 almost 2 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum, scutellum and mesepisternum very dense with interspaces rarely reaching diameter of one puncture, punctures about as large as those on frons. Axilla of scutellum laterally not spined. Metanotum medially projecting to weakly spined. Basal area of propodeum at its base with very narrow and longitudinally carinate horizontal zone, which is bordered by distinct transverse carina along posterior margin. Posterior surface of propodeum adjacent to basal area polished and medially unpunctured. Tarsal segments 2 – 4 of fore and middle leg about as long as wide to slightly wider than long. Tibial spurs of hind leg yellowish and straight except for their apex, which is slightly bent. Metasoma: Declivous part of tergum 1 distinctly separated from horizontal part by sharp and raised carina. Punctation of terga dense with interspaces rarely exceeding diameter of one puncture, punctures half as large as those on scutum. Marginal zones of terga 1 – 5 with short and uninterrupted white hair bands (Fig. 1). Scopa white (Fig. 1). MALE: Body length 5 mm. Head: Head as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.7 × as long as ocellar diameter. Maximum width of genal area 0.5 × as long as maximum width of compound eye. Proboscis short, reaching in repose not beyond proboscidial fossa. Mandible two-toothed, black in its basal half and predominantly reddish in its apical half. Lower two thirds of face covered with dense white pilosity. Hypostomal area and adjacent base of mandible covered with dense white pilosity composed of long and distinctly plumose hairs. Punctation of frons very dense with interspaces rarely exceeding diameter of half a puncture. Antenna brownish except for predominantly blackish scapus. Antennal segments 3 – 4 shorter than wide, segments 5 – 12 as long as wide to slightly longer and segment 13 about 1.75 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum, scutellum and mesepisternum very dense with interspaces rarely exceeding diameter of half a puncture, punctures about as large as those on frons. Axilla of scutellum laterally not spined. Metanotum medially weakly spined. Basal area of propodeum at its base with very narrow and longitudinally carinate horizontal zone, which is bordered by distinct transverse carina along posterior margin. Posterior surface of propodeum adjacent to basal area polished and medially unpunctured. Lower half of lateral mesepisternum and entire ventral mesepisternum rather densely covered with white pilosity. Tibia of fore and middle leg slightly widened. Posterior side of tarsal segments 1 – 4 of fore leg with long and plumose white hairs, which decrease in length from segment 1, where they are almost 3 × as long as tarsal width, to segment 4, where they are about as long as tarsal width (Fig. 4). Tarsal segments 2 – 3 (4) of fore, middle and hind leg asymmetrical and distinctly to slightly wider than long (Fig. 4). Ventral side of trochanter and femur of middle leg with dense brush of plumose white hairs. Basitarsus of hind leg with long white hairs, which form a rather dense brush on its ventral side. Tibial spurs of hind leg yellowish. Metasoma: Declivous part of tergum 1 distinctly separated from horizontal part by sharp and raised carina. Punctation of terga very dense with interspaces rarely reaching diameter of one puncture, punctures smaller than those on scutum. Preapical zone of tergum 6 medially with oval bump and laterally with impressed and sparsely punctured transversal groove. Tergum 7 weakly sclerotised and completely hidden under tergum 6, its apical margin medially with distinct triangular incision. Marginal zones of terga 1 – 3 (4) with short and uninterrupted white hair bands. Sternum 1 convex and slightly wider than long. Sternum 2 medially emarginate. Sternum 3 medioapically with dense and triangular patch of long yellowish hairs largely surpassing tergal margin and covering about one fourth of sternal width. Apical margin of sternum 4 mediolaterally with two rows of six brownish bristles each, which are separated by deep triangular incision, the inner margins of which are lined with long yellowish hairs. Gonoforceps very slender, distinctly surpassing apex of penis valve and curved inwards in its apical third (Fig. 5, 6). Curved apical part of gonoforceps long, finger-like, more slender than adjacent more basal part and apically with tuft of few long white hairs (Fig. 5, 6).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	distribution	Distribution. Spain and southeastern Mediterranean region (southwesternmost Bulgaria, Greece including Aegean Islands, western Turkey). The females from Spain are morphological identical with those from southeastern Europe and Turkey. However, as no males from Spain are known and females of closely related Heriades species of the subgenus Michenerella are often very similar in morphology, it cannot be excluded that the Spanish and southeastern Mediterranean populations belong to two different species. Pending the discovery of the male sex, the Spanish population is here tentatively assumed to be conspecific with the eastern populations.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	biology_ecology	Habitat. Uncultivated slope between cereals fields and the river Eresma covered with scrub including Retama (Fabaceae) on the more steeply sloping parts and with patches of Eryngium campestre on the more open parts, these swarming with H. clypeonitida. With 55 additional species recorded, a diverse bee fauna was present. Bee sampling at many Eryngium - rich sites across Ávila and Segovia provinces from 16. – 27.7.2021 did not reveal additional specimens of H. clypeonitida, suggesting that the distribution of this species might be extremely local or that Eryngium is exploited only in the absence of the normal host flowers. Pollen hosts. In Spain, the type specimens were all observed to suck nectar on Eryngium campestre (Apiaceae), but no pollen collection was directly recorded (T. Wood, personal observation).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5730447ED6A6FAE03FCD8FE3.taxon	etymology	Etymology. The species epithet refers to the diagnostic character of the female, i. e. the punctation of the clypeus, which is very scattered giving the clypeus a very shiny appearance (lat. “ nitidus ” = shiny).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	materials_examined	Holotype. SPAIN: Granada: Sierra Nevada, Jardin Botanico, Hoya Pedraza, 37.1130 ° N / 3.4378 ° W, 1900 m, 29.6.2021, ♂ (leg. T. Wood). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. SPAIN: Granada: Sierra Nevada, Jardin Botanico, Hoya Pedraza, 37.1130 ° N / 3.4378 ° W, 1900 m, 29.6.2021, 3 ♀, 2 ♂ (leg. T. Wood); Sierra Nevada, El Dornajo, 37.1323 ° N / 3.4392 ° W, 1700 m, 6.6.2021, 1 ♂. Paratypes deposited in the Entomological Collection of ETH Zurich and the private collection of T. Wood.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	diagnosis	Diagnosis. Among the European and North African Hoplitis species of the subgenus Anthocopa, the females of Hoplitis nevadensis (Fig. 8) can be recognised by the following combination of characters: i) body length 6 – 7 mm; ii) apical margin of clypeus with strongly rounded median projection (Fig. 10); iii) second segment of labial palpus almost 2.5 × as long as first segment and as long as compound eye; iv) pilosity on inner side of basitarsus of hind leg brownish; v) tibial spurs of hind leg and tegula predominantly blackish; and vi) metasomal scopa white and with numerous distinctly plumose hairs. (Fig. 8) The males (Fig. 9) can be identified by the following combination of characters: i) body length 6.5 – 7 mm; ii) second segment of labial palpus almost 2.5 × as long as long as first segment and as long as compound eye; iii) mandible, antenna, tibial spurs of hind leg and tegula predominantly blackish; iv) tergum 7 bidentate and laterally not projecting, its median emargination 1.5 – 2.5 × as wide as preapical width of adjacent teeth (Fig. 11); v) pilosity of sternum 5 very short, sparse, neither hiding cuticular surface nor forming apical hair band and of about same density over entire sternal surface (Fig. 12); and vi) sternum 6 mediopreapically with polished roundish impression, which is neither haired nor punctured (Fig. 12).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	description	Description. FEMALE (Fig. 8: Body length 6 – 7 mm. Head: Head about 0.9 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.9 – 2.1 × as long as ocellar diameter. Maximum width of genal area 0.6 – 0.7 × as long as maximum width of compound eye. Second segment of labial palpus 2.3 – 2.4 × as long as first segment and as long as compound eye. Mandible three-toothed and blackish except for preapical reddish zone. Clypeus distinctly convex, its punctation very dense with only linear interspaces except for narrow apical zone, which is not to very sparsely punctured (Fig. 10). Apical margin of clypeus medially with strongly rounded median projection (Fig. 10). Lower two thirds of face with whitish and predominantly erect pilosity, which is sparse on clypeus and rather dense on paraocular area, supraclypeal area and frons. Punctation of supraclypeal area denser and finer than that of clypeus except for medioapical zone, where punctation may be more scattered (Fig. 10). Punctation of frons dense with interspaces reaching diameter of half a puncture to one puncture, punctures of about same size as those on clypeus. Hypostomal area and ventral side of mandibles covered with long and inwardly curved yellowish-white bristles forming distinct basket on head underside. Antenna blackish. Antennal segments 3 and 12 about 1.3 × as long as wide and segments 4 – 11 shorter than wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum and mesepisternum very dense with interspaces rarely exceeding diameter of half a puncture, punctures of about same size as those on frons. Punctation of scutellum of similar size and density as that of scutum except for median zone, where punctures may be more scattered. Basal area of propodeum polished except for narrow transverse zone at its base, which is finely sculptured. Posterior surface of propodeum adjacent to basal area polished and sparsely punctured. Tegula predominantly dark brown. Tibial spur of fore leg yellowish, its apex extended into long and well stepped tip, which is more than 2 × as long as basally wide. Tibial spurs of middle and hind leg dark brown, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg brownish. Veins of fore and hind wing dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Terga 2 – 3 basally distinctly constricted. Punctation of tergal discs 1 – 5 moderately dense with interspaces varying in size between diameter of half a puncture on lateral tergal sides and diameter of up to two to three punctures on median parts of terga 2 – 3, punctures slightly smaller than those on scutum. Punctation of tergum 6 distinctly smaller than that of terga 1 – 5 and very dense with interspaces hardly exceeding diameter of half a puncture. Marginal zones of terga 1 – 5 with short and uninterrupted white hair bands (Fig. 8). Tergal discs 2 – 5 covered with very short and predominantly dark hairs. Pilosity of tergum 6 short, semierect, predominantly whitish and sparse, not hiding cuticular surface. Scopa white (Fig. 8), scopal hairs of sterna 2 – 4 distinctly plumose. MALE (Fig. 9): Body length 6.5 – 7 mm. Head: Head 0.8 – 0.9 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.9 – 2 × as long as ocellar diameter. Maximum width of genal area 0.6 – 0.8 × as long as maximum width of compound eye. Second segment of labial palpus 2.3 – 2.5 × as long as first segment and as long as compound eye. Mandible three-toothed and black except for narrow reddish preapical zone. Apical margin of clypeus almost straight and with five minute teeth. Frons, paraocular area, supraclypeal area, clypeus and hypostomal area covered with dense and predominantly erect to semierect white pilosity. Punctation of frons very dense with interspaces not exceeding diameter of one puncture. Punctation of clypeus, supraclypeal area and paraocular area extremely fine and dense with only linear interspaces. Antenna dark brown to black. Antennal segment 3 about as long as wide, segments 4 – 12 shorter than wide and segment 13 about 1.25 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum and mesepisternum very dense with interspaces rarely exceeding diameter of half a puncture, punctures of about same size as those on frons. Punctation of scutellum of similar size and density as that of scutum except for median zone, where punctures may be more scattered. Basal area of propodeum polished except for narrow transverse zone at its base, which is finely sculptured. Posterior surface of propodeum adjacent to basal area polished with sparse and fine punctation. Tegula dark brown in its anterior and reddish-brown in its posterior half. Mesepisternum covered with long and erect white pilosity that does not hide cuticular surface. Tibial spur of fore leg yellowish, its apex extended into long and well stepped tip, which is more than 2 × as long as basally wide. Tibial spurs of middle and hind leg brown to dark brown, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg brownish. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Terga 2 – 3 basally distinctly constricted. Punctation of tergal discs moderately dense with interspaces varying in size between diameter of half a puncture or less on lateral tergal sides and diameter of up to two to three punctures on median parts of terga 3 – 4, punctures slightly smaller than those on scutum. Marginal zones of terga 1 – 5 with short white hair bands, which may be interrupted on terga 1 – 2 in worn specimens (Fig. 9). Tergal discs (3) 4 – 5 covered with moderately long and predominantly dark pilosity with more or less intermixed whitish hairs. Tergum 6 with lateral short tooth and without apical white hair band. Tergum 7 bidentate and laterally not projecting, its median emargination 1.5 – 2.5 × as wide as preapical width of adjacent teeth (Fig. 11). Sternum 2 covered with rather long and erect white pilosity that does not hide cuticular surface, its apical margin evenly rounded and with very loose band of white hairs, which surpass tergal margin (Fig. 12). Apical margin of sternum 3 medially very shallowly emarginate, emargination beset with dense band of yellowish-white and slightly inwardly directed hairs, which are about as long as hairs along apical margin of sternum 2 (Fig. 12). Apical margin of sternum 4 widely and shallowly emarginate, emargination beset with dense band of yellowish-white and distinctly inwardly directed hairs, which are about twice as long as hairs along apical margin of sternum 3 (Fig. 12). Sternum 5 medioapically very shallowly emarginate, its pilosity whitish, very short, neither hiding cuticular surface nor forming apical hair band and of about same density over entire sternal surface (Fig. 12). Sternum 6 mediopreapically with more or less roundish impression, which is strongly polished, unhaired and unpunctured (Fig. 12). Gonoforceps apically widened and curved under penis valve (Fig. 13, 14).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	distribution	Distribution. Sierra Nevada in southern Spain.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	biology_ecology	Habitat. Subalpine grassland near the edge of the tree line (Fig. 7). Woody plants such as Crataegus (Rosaceae), Berberis (Berberidaceae) and low growing clumps of Vella spinosa (Brassicaceae) were in bloom together with a rich ground flora of Asteraceae, Fabaceae, and Lamiaceae. With 51 additional species recorded, a diverse bee fauna was present.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E5735447FD6A6FD0C3F8C8ABB.taxon	etymology	Etymology. The species epithet refers to the occurrence of the species in the Sierra Nevada.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	materials_examined	Holotype. SPAIN: Granada: Sierra Nevada, Jardin Botanico, Hoya Pedraza, 37.1130 ° N / 3.4378 ° W, 1900 m, 9.6.2021, ♂ (leg. T. Wood). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. SPAIN: Granada: Sierra Nevada, Jardin Botanico, Hoya Pedraza, 37.1130 ° N / 3.4378 ° W, 1900 m, 9.6.2021, 1 ♀, 1 ♂ (leg. T. Wood). Paratypes deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	diagnosis	Diagnosis. Hoplitis spinidorsa is the only representative of the subgenus Anthocopa, whose scutellar axilla is in both sexes lateroapically elongated into a long, pointed and inwardly directed spine (Fig. 17), a character that it shares with species of Heriades (Heriades) and Osmia (Hoplosmia). The females (Fig. 15) differ from those of the latter two taxa by the presence of a distinct hair basket on the underside of the head, which is formed by long and inwardly curved yellowish bristles arising from the ventral side of gena and mandible (Fig. 18), and by the strongly convex scutellum. The males (Fig. 16) differ by i) the quadripartite tergum 7 formed by two lateral teeth and two closely adjacent small median lobes (Fig. 19), ii) the triangular tuft of yellowish-white hairs medioapically projecting far beyond the margin of sternum 4 (Fig. 20), iii) two closely adjacent and nearly parallel longitudinal keels medioapically on sternum 6 (Fig. 20), and v) the dense tuft of long yellowish hairs arising from the very wide apical margin of the gonoforceps.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	description	Description. FEMALE (Fig. 15): Body length 6.5 mm. Head: Head about 0.9 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin about 2 × as long as ocellar diameter. Maximum width of genal area about 0.75 × as long as maximum width of compound eye (Fig. 18). Second segment of labial palpus about 2 × as long as first segment and about 0.6 × as long as compound eye. Mandible three-toothed and blackish except for preapical reddish zone. Clypeus medially distinctly bulged with foremost point in a distance of about one third of clypeal length from lower margin of supraclypeal area. Punctation of clypeus rather dense with interspaces reaching diameter of up to one puncture except for narrow apical zone, which is unpunctured, polished and distinctly surpassed by long yellowish bristles arising at base of unpunctured zone. Apical margin of clypeus sharp and straight. Pilosity of paraocular area and lower frons white, rather dense and suberect to erect. Pilosity of clypeal disc yellowish-white, sparse and erect. Supraclypeal area plane to very weakly impressed, its punctation almost as large as that of clypeus and medially scattered with interspaces reaching up to two, rarely more punctures. Punctation of frons rather dense with interspaces varying in size between diameter of half a puncture on median part to diameter of two punctures on lateral parts, punctures of about same size as those on clypeus to slightly smaller. Ventral side of gena and mandibles with long and inwardly curved yellowish bristles forming distinct basket on head underside (Fig. 18). Antenna blackish except for apical half of flagellum, which is partly (reddish-) brown on anterior and posterior side. Antennal segment 3 about 1.3 × as long as wide, segments 4 – 11 shorter than wide and segment 12 about 1.6 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum dense with interspaces hardly exceeding diameter of one puncture, punctures of about same size as those on clypeus. Punctation of mesepisternum moderately dense with interspaces reaching diameter of up to two punctures. Punctation of scutellum very dense with interspaces not exceeding diameter of half a puncture (Fig. 17). Scutellum near its base distinctly bulged. Axilla elongated into long, pointed and inwardly directed spine (Fig. 17). Basal area of propodeum polished except for very narrow transverse zone at its base, which is finely sculptured. Posterior surface of propodeum adjacent to basal area polished with dense and fine punctation. Tegula predominantly yellowish-red. Scutum sparsely covered with short and erect yellowish-white hairs. Posterior fourth of scutellum and metanotum with long and backwardly directed yellowish-red hairs (Fig. 17). Mesepisternum sparsely covered with suberect and moderately long yellowish-white hairs. Tibial spur of fore leg yellowish, its apex extended into short tip, which is about as long as basally wide. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg yellowish-red. Veins of fore and hind wing brown to dark brown. Stigma of fore wing dark brown. Metasoma: Punctation of tergal discs 1 – 5 rather scattered with interspaces varying in size between diameter of up to one puncture on terga 1 and 5 and distance of up to two, rarely more punctures on median parts of terga 2 – 4, punctures slightly smaller than those on scutum. Punctation of tergum 6 distinctly smaller than that of terga 1 – 5 and very dense with interspaces not reaching diameter of half a puncture. Marginal zones of terga 1 – 4 with short and uninterrupted white hair bands (Fig. 15). Marginal zone of tergum 5 reddish and with very short and inconspicuous white hair band. Tergal discs 2 – 5 covered with very short, erect and predominantly dark hairs. Pilosity of tergum 6 very short, appressed, whitish and sparse, not hiding cuticular surface. Scopa yellowish-red (Fig. 15), scopal hairs of sterna 2 – 4 apically partly beaded. MALE (Fig. 16): Body length 7 mm. Head: Head 0.8 – 0.9 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 2.0 – 2.1 × as long as ocellar diameter. Maximum width of genal area 0.6 – 0.7 × as long as maximum width of compound eye. Second segment of labial palpus 2.1 – 2.2 × as long as first segment and 0.62 – 0.63 × as long as compound eye. Mandible three-toothed and black except for narrow reddish preapical zone. Apical margin of clypeus medially with short triangular process, section between this process and lateral clypeal corner emarginate. Frons, paraocular area, supraclypeal area, clypeus and hypostomal area covered with dense and predominantly erect to semierect white pilosity. Punctation of frons very dense with interspaces varying in size between diameter of less than half a puncture on median part to diameter of one puncture on lateral parts. Antenna blackish except for apical half of flagellum, which is partly (reddish-) brown on anterior and posterior side. Antennal segment 3 about 1.5 × as long as wide, segments 4 – 12 slightly shorter than wide to almost as long as wide and segment 13 about 1.25 × as long as wide. Ventral side of antennal segments 3 – 4 with long whitish hairs. Mesosoma: Parapsidal line long-linear. Punctation of scutum and mesepisternum dense with interspaces rarely reaching or exceeding diameter of one puncture, punctures slightly larger than those on frons. Punctation of scutellum very dense with interspaces not exceeding diameter of half a puncture except medially, where punctures may be more scattered. Scutellum near its base bulged. Axilla elongated into long, pointed and inwardly directed spine. Basal area of propodeum polished except for narrow transverse zone at its base, which is finely sculptured. Posterior surface of propodeum adjacent to basal area polished with dense and rather fine punctation. Tegula predominantly yellowish-red. Scutum sparsely beset with whitish hairs. Margins of scutum, scutellum and metanotum covered with rather dense, long and yellowish-red pilosity. Mesepisternum covered with similarly long and dense but whitish pilosity. Tibial spur of fore leg yellowish, its apex extended into tip, which is longer than basally wide. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg yellowish-red. Veins of fore and hind wing brown to dark brown. Stigma of fore wing dark brown with slightly lighter centre. Metasoma: Punctation of tergal discs rather dense with interspaces varying in size between diameter of half a puncture on lateral tergal sides and diameter of up to two punctures on median parts of terga 3 – 4, punctures of about same size as those on scutum. Marginal zones of terga (1,2) 3 – 6 yellowish or reddish (Fig. 19) and of terga 1 – 4 with short white hair bands (Fig. 16). Tergal discs 3 – 5 covered with moderately long and predominantly dark hairs. Tergum 6 without lateral tooth. Tergum 7 quadripartite with two lateral teeth and two closely adjacent small median lobes, which are basally fused and crossed by a transverse swelling (Fig. 19). Apical margin of sterna 2 – 3 medially shallowly emarginate. Sternum 4 medioapically with triangular tuft of yellowish-white hairs projecting far beyond sternal margin (Fig. 20). Sternum 5 medially slightly impressed, its apical margin weakly emarginate (Fig. 20). Sternum 6 basally without pair of translucent membraneous flaps, medially with distinct roundish impression and medioapically with two closely adjacent and nearly parallel longitudinal keels (Fig. 20). Gonoforceps increasing in width towards its apex, which is very wide, diagonally truncate and beset with dense tuft of long yellowish hairs.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	distribution	Distribution. Sierra Nevada in southern Spain.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	biology_ecology	Habitat. As for H. nevadensis sp. nov. (see above), which was found at the very same site as H. spinidorsa (Fig. 7).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57374472D6A6FF4D3A9288FB.taxon	etymology	Etymology. The species epithet refers to the spined axillae on the dorsal side of the mesosoma (lat. “ spina ” = spine, lat. “ dorsum ” = back)	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	materials_examined	Holotype. GREECE: Crete: Ida mountains, 2100 m, 14.6.1976, ♀ (leg. K. Warncke). Holotype deposited in the Entomological Collection of ETH Zurich. Paratype. GREECE: Crete: Ida mountains, Ebriskos, 10 km S Anogia, 15.6.2003, 1 ♀ (leg. W. H. Liebig). Paratype deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	diagnosis	Diagnosis. Hoplitis cretensis is a member of the Hoplitis (Hoplitis) adunca species group as shown by the presence of a submarginal carina on sternum 6 in the female sex. Among the European and North African representatives of the H. adunca group, the females of H. cretensis (Fig. 21) can be recognized by the following combination of characters: i) body length 6.5 – 7 mm; ii) punctation of lateral part of clypeus with distinct polished interspaces (Fig. 22); iii) clypeus with unpunctured and usually uninterrupted longitudinal midline, which extends almost till clypeal apex and whose maximum median width reaches diameter of up to two or three punctures (Fig. 22); iv) punctation of supraclypeal area scattered with interspaces reaching diameter of up to three punctures medioapically (Fig. 22); v) proboscis not reaching trochanter of hind leg in repose, second segment of labial palpus about 1.5 – 1.6 × as long as first segment and distinctly shorter than maximum length of mesosoma measured in lateral view; vi) galea of proboscis and clypeus normally haired, without specialised and apically curved or wavy pollen-harvesting bristles; vii) apical tooth of tibial spur of fore leg apically more or less rounded to obtuse rather than sharply pointed; viii) longest erect hairs on median half of tergum 1 distinctly shorter than maximum length of lateral hair tuft when seen from behind; and ix) sternum 6 lateroapically with very strong submarginal carina, its apical margin medially elongated into distinct and well delimited tooth (Fig. 24). In the key to the European species of the Hoplitis adunca species group (Müller 2016), the female of H. cretensis keys out at couplet 9 together with H. fabrei Zanden, 1987 and H. pallicornis (Friese, 1895). In addition to the smaller size (character i), the usually longer length and larger width of the clypeal midline (character iii) and the more scattered punctation of the supraclypeal area (character iv), H. cretensis differs from these two related species by the shape of the clypeus, which is more strongly convex in lateral view and apically more abruptly bent towards the clypeal margin (Fig. 23). The male of H. cretensis is unknown.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	description	Description. FEMALE (Fig. 21): Body length 6.5 – 7 mm. Head: Head 1.0 – 1.1 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin about 1.9 × as long as ocellar diameter. Maximum width of genal area about 0.7 – 0.8 × as long as maximum width of compound eye. Second segment of labial palpus 1.5 – 1.6 × as long as first segment and 0.7 – 0.8 × as long as compound eye. Mandible three-toothed and blackish except for dark reddish mandibular teeth. Clypeus strongly convex, distinctly surpassing supraclypeal area in lateral view and abruptly bent towards clypeal margin in its apical third (Fig. 23). Punctation of lateral part of clypeus with interspaces reaching diameter of up to one puncture (Fig. 22). Clypeus with unpunctured longitudinal midline, which extends almost till clypeal apex and whose maximum median width reaches diameter of up to two or three punctures (Fig. 22). Punctation of supraclypeal area scattered with interspaces reaching diameter of up to three punctures medially, punctures slightly smaller than those on clypeus (Fig. 22). Apical margin of clypeus straight and with 7 – 8 minute teeth. Pilosity of paraocular area, supraclypeal area and lower frons white, moderately dense and weakly appressed to erect. Clypeal disc sparsely covered with long and erect white hairs. Clypeal margin ciliated with yellowish-white hairs, which are up to half as long as clypeal length (Fig. 22). Punctation of frons very dense with interspaces not exceeding diameter of half a puncture except for two small areas in front of lateral ocelli, where interspaces may reach diameter of up to one and a half to two punctures. Punctures on frons slightly smaller than those on clypeus. Antenna blackish except for anterior surface of segments (4) 5 – 11 (12), which is reddish-brown. Antennal segments 3 and 12 about 1.5 × as long as wide and segments 4 – 11 shorter than wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum and scutellum very dense with interspaces exceeding diameter of one puncture only medially on scutellum, punctures of about same size as those on frons. Punctation of mesepisternum slightly more scattered than that of scutum with interspaces varying in size between diameter of half a puncture to diameter of one and a half punctures. Basal area of propodeum shagreened throughout. Posterior surface of propodeum adjacent to basal area shagreened to almost polished with very sparse and fine punctation. Tegula yellowish to yellowish-brown in its posterior half. Tibial spur of fore leg yellowish, its apex extended into tip, which is slightly longer than basally wide and apically more or less rounded to obtuse rather than sharply pointed. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg yellowish. Veins of fore and hind wing light brown to dark brown. Stigma of fore wing dark brown with slightly lighter centre. Metasoma: Punctation of tergal discs 1 – 4 rather fine and very scattered with interspaces varying in size between diameter of up to one puncture on tergal sides and diameter of up to several punctures on median tergal parts, punctures much smaller than those on scutum. Punctation of tergum 5 denser than on preceding terga with interspaces reaching diameter of two to three punctures. Punctation of tergum 6 very dense with interspaces hardly exceeding diameter of half a puncture. Terga 1 – 3 (4) predominantly polished, terga (4) 5 – 6 shagreened. Longest erect hairs on median half of tergum 1 distinctly shorter than maximum length of lateral hair tuft when seen from behind. Marginal zones of terga 1 – 5 inconspicuously dark reddish and with short white hair bands (Fig. 21), which may be medially interrupted in worn specimens. Tergum 6 rather densely covered with appressed white pilosity largely hiding cuticular surface. Sternum 6 lateroapically with very strong submarginal carina, its apical margin medially elongated into distinct and well delimited tooth of narrowly triangular shape (Fig. 24). Scopa white, on sternum 2 basally greyish (Fig. 21).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	distribution	Distribution. Ida mountains on Crete.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	biology_ecology	Pollen hosts. Unknown. Nesting biology. Unknown.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57394473D6A6FA343F068A93.taxon	etymology	Etymology. The species epithet refers to the occurrence of the species on Crete.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573B4471D6A6FF4D3F358ABB.taxon	materials_examined	Holotype. SPAIN: Almería: Nijar, 36 ° 59 ' 08 " N / 2 ° 14 ' 06 ” W, 23.5.2018, ♂ (leg. J. Smit). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. SPAIN: Almería: Berja, 50 km W Almería, 21. – 28.6.2003, 1 ♂ (leg. J. Halada); Cabo de Gata, Frente al Ctr., Visit. Amoladeras, 10.5.2004, 1 ♂ (leg. A. Aguirre); Los Muertos, 36.950 ° N / 1.898 ° W, 30.4.2014, 1 ♂ (leg. M. Aubert). Paratypes deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573B4471D6A6FF4D3F358ABB.taxon	diagnosis	Diagnosis. Hoplitis andalusiae is morphologically very close to H. hispaniae sp. nov. in the female sex and no morphological characters are currently known to separate the females of the two species. Among the European and North African Hoplitis species of the subgenus Micreriades, the females of H. andalusiae and H. hispaniae (Fig. 25) can be recognised by the following combination of characters: i) head 1.05 – 1.15 × as long as wide; ii) inner margins of compound eyes slightly converging towards clypeus; iii) clypeus basally moderately bulged: in lateral view, transitional zone between bulging part and marginal zone of clypeus straight to very slightly convex; iv) frons between anterior ocellus and antennal sockets with numerous apically wavy to corkscrew-like pollen-collecting hairs (Fig. 28), which may be worn off in old individuals; v) punctation of scutum between parapsidal line and median longitudinal furrow rather scattered with interspaces often reaching the diameter of two, rarely more punctures (Fig. 27); vi) toothlike projection at apex of tibia of fore leg yellowish-brown; and vii) tibial spurs of middle and hind leg yellowish. The males of H. andalusiae (Fig. 26) are characterised by the following combination of characters: i) head very slightly shorter than wide; ii) antennal segment 3 as long as maximally wide; iii) antennal segment 13 cylindrical and apically not curved downwards; iv) punctation of scutum between parapsidal line and median longitudinal furrow rather scattered with interspaces reaching the diameter of up to two or more punctures; v) apical margin of sternum 3 with dense median patch of long whitish to yellowish hairs distinctly surpassing tergal margin and covering almost half of horizontal sternal width; vi) sternum 6 lateropreapically roundishly sloping towards sternal centre, but without distinct and well-stepped lateral callosity (Fig. 29); vii) sternum 6 medioapically with distinct impression and without keel-like longitudinal elevation, its apical margin evenly rounded (Fig. 29); and viii) gonoforceps apically not evenly tapering but with slightly stepped tip (Fig. 30).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573B4471D6A6FF4D3F358ABB.taxon	description	Description. FEMALE (Fig. 25): Body length 5 – 6.5 mm. Head: Head 1.05 – 1.15 × as long as wide. Inner margins of compound eyes slightly converging towards clypeus. Distance between posterior margin of lateral ocellus and preoccipital margin 1.7 – 1.8 × as long as ocellar diameter. Maximum width of genal area 0.65 – 0.75 × as long as maximum width of compound eye. Second segment of labial palpus 1.8 – 2.0 × as long as first segment and 0.65 – 0.75 × as long as compound eye. Mandible three-toothed and blackish except for preapical reddish zone. Clypeus medially moderately bulged with foremost point in a distance of about one third of clypeal length from lower margin of supraclypeal area. In lateral view, transitional zone between bulging part and marginal zone of clypeus straight to very slightly convex. Punctation of clypeus dense with interspaces reaching diameter of half a puncture to one, rarely one and a half punctures except for narrow apical zone, which is unpunctured and polished. Clypeus usually crossed medially by weak longitudinal carina. Apical margin of clypeus medially sharp and almost straight. Upper half of paraocular area with rather dense and predominantly appressed white pilosity. Clypeus densely covered with rather long and apically wavy to corkscrew-like pollen-collecting hairs. Punctation of supraclypeal area dense with interspaces hardly exceeding diameter of half a puncture on lateral parts and reaching up to one, rarely one and a half punctures on median part; punctures slightly smaller than those on clypeus. Punctation of frons dense with interspaces reaching diameter of half a puncture to one puncture; punctures of about same size as those on supraclypeal area. Frons between anterior ocellus and antennal sockets densely covered with rather long and apically wavy to corkscrew-like pollen-collecting hairs (Fig. 28). Antenna blackish except sometimes for anterior side of flagellum, which may be more or less brownish. Antennal segment 3 – 9 distinctly wider than long, segments 10 – 11 almost as long as wide and segment 12 1.6 – 1.8 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum between parapsidal line and median longitudinal furrow rather scattered with interspaces often reaching diameter of two, rarely more punctures (Fig. 27), punctures smaller than those on frons. Punctation of scutellum dense with interspaces rarely reaching diameter of one puncture. Punctation of mesepisternum moderately dense with interspaces reaching diameter of up to one to one and a half punctures on upper part and of two or more punctures on lower part. Basal area of propodeum predominantly polished except for narrow transverse zone at its base, which is weakly carinate and shagreened. Posterior surface of propodeum adjacent to basal area polished with scattered and fine punctation. Tegula predominantly yellowish-brown to brown. Tibial spur of fore leg yellowish, its apex extended into well stepped tip, which is as long as to slightly longer than basally wide. Toothlike projection at apex of tibia of fore leg yellowish-brown. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Pilosity on inner side of basitarsus of hind leg yellowish. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs dense with interspaces varying in size between diameter of less than half a puncture on lateral tergal sides and diameter of up to one or two punctures on median tergal parts, punctures of about same size as those on scutum. Marginal zones of terga 1 – 5 dark reddish. Marginal zones of terga 1 – 3 (4) with short white hair bands, which are medially interrupted on terga 1 – 2 and decrease in length and density from tergum 1 to tergum 4 (Fig. 25). Scopa white (Fig. 25). MALE (Fig. 26): Body length 5.5 – 6.5 mm. Head: Head 0.97 – 0.99 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.5 – 1.7 × as long as ocellar diameter. Maximum width of genal area 0.45 – 0.55 × as long as maximum width of compound eye. Second segment of labial palpus 2.15 – 2.3 × as long as first segment and about 0.75 × as long as compound eye. Mandible two-toothed and black except for reddish preapical zone. Clypeus very finely and densely punctured with only linear interspaces except for narrow marginal zone, which is polished and unpunctured. Apical margin of clypeus medially very shallowly emarginate and inconspicuously crenulate. Lower half of frons, paraocular area, supraclypeal area and lateral sides of clypeus covered with dense yellowish-white pilosity, which is (semi) erect on frons and supraclypeal area and predominantly appressed on paraocular area and lateral sides of clypeus; pilosity of clypeus medially sparser than laterally. Hypostomal area with moderately dense whitish pilosity. Punctation of frons moderately dense with interspaces reaching diameter of up to one or one and a half punctures. Antenna predominantly (dark) brown. Antennal segment 3 as long as wide and slightly longer than segment 4, which is slightly shorter than wide, segments 5 – 11 roughly as long as wide to slightly shorter, segment 12 slightly longer than wide and segment 13 about 1.75 × as long as wide. Antennal segment 13 cylindrical and apically not curved downwards. Posterior side of antennal segments 9 – 13 almost straight in top view. Mesosoma: Parapsidal line long-linear. Punctation of scutum between parapsidal line and median longitudinal furrow rather scattered with interspaces reaching diameter of up to two or more punctures, punctures smaller than those on frons. Punctation of scutellum rather dense with interspaces rarely reaching diameter of one puncture. Punctation of upper part of mesepisternum rather scattered with interspaces reaching diameter of one and a half to two punctures, of lower part strongly scattered with interspaces reaching diameter of several punctures. Basal area of propodeum polished except for narrow transverse zone at its base, which is weakly carinate and shagreened. Posterior surface of propodeum adjacent to basal area polished with strongly scattered and fine punctation. Tegula predominantly yellowish-brown. Tibial spur of fore leg yellowish, its apex extended into short tip, which is about as long as basally wide. Toothlike projection at apex of tibia of fore leg yellowish-brown. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs 1 – 5 dense with interspaces varying in size between diameter of less than half a puncture on lateral tergal sides and diameter of up to two punctures on median tergal parts, punctures of about same size as those on scutum. Marginal zones of terga 1 – 6 reddish. Marginal zones of terga 1 – 4 with short white hair bands, which are medially interrupted on terga 1 – 2 (Fig. 26). Tergum 6 with short lateral tooth, its marginal zone slightly raised. Tergum 7 with roundish median pit at its base and medioapically elongated into narrowly triangular tip. Sternum 1 convex. Sternum 2 mediobasally flat and with polished or indistinctly shagreened interspaces between punctures, its apical margin evenly rounded (sometimes with very shallow and narrow medioapical emargination) and surpassed by moderately long and sparse white pilosity, which is about as long as hair bands of terga 3 – 4. Apical margin of sternum 3 medially very shallowly emarginate and with dense patch of whitish to yellowish hairs distinctly surpassing tergal margin and covering almost half of horizontal sternal width. Sternum 4 medioapically more deeply and slightly more widely emarginate than sternum 3, emargination lined with yellowish-white hairs (Fig. 29), which are similarly long as hair patch of sternum 3. Sternum 5 on marginal zone with rather dense but short and (semi) erect pilosity, its apical margin almost straight to medially slightly emarginate (Fig. 29). Sternum 6 basally with pair of translucent membraneous flaps, medioapically with distinct, widely triangular, strongly polished and almost unpunctured impression, and lateropreapically roundishly sloping towards sternal centre, without distinct and well-stepped lateral callosity (Fig. 29). Apical margin of sternum 6 evenly rounded (Fig. 29). Gonoforceps apically with slightly stepped tip (Fig. 30).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573B4471D6A6FF4D3F358ABB.taxon	distribution	Distribution. Southeasternmost Andalusia.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573B4471D6A6FF4D3F358ABB.taxon	etymology	Etymology. The species epithet refers to the occurrence of the species in Andalusia.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573C4474D6A6FF4D3F598A63.taxon	materials_examined	Holotype. SPAIN: Madrid: Velilla de San Antonio, 40.35741 ° N / 3.48137 ° W, 2.6.2013, ♂ (leg. W. H. Liebig). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. SPAIN: Granada: N Lanjaron, Rio de Lanjaron, 1200 m, 24.4.1999, 1 ♂ (leg. B. Pohl); Sierra Nevada, Bco. las Mimbres (Monachil), 1700 m, 17.6.2007, 1 ♀ (leg. F. J. Ortiz); Jaén: Cambil, 23.6.1988, 1 ♀ (leg. M. Schwarz); Málaga: Andalusia, Colmenar, Carrasco, 36 ° 52 ' 27 " N / 4 ° 18 ' 22 " W, 9.6.2016, 1 ♀ (leg. J. Smit); Salamanca: Villar de Ciervo, Las Coronas, 40.740922 ° N / 6.743543 ° W, 680 m, 31.5. – 9.6.2003, 1 ♀ (leg. Tschorsnig); Sevilla: Aznalcazar, S Pinares de Aznalcazar, 37.2622 ° N / 6.2216 ° W, 21.5.2021, 1 ♀ (leg. T. Wood); Toledo: Hormigos, 22.5.2022, 1 ♂ (leg. J. I. Pascual). Paratypes deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573C4474D6A6FF4D3F598A63.taxon	diagnosis	Diagnosis. Among the European and North African Hoplitis species of the subgenus Micreriades, Hoplitis fuscospina is the only species, which has in both sexes (light) brown to dark brown rather than yellowish tibial spurs of the middle and hind leg (Fig. 33). The females (Fig. 31) are additionally characterised by the following characters, which however can be found also in other H. (Micreriades) species: i) head 1.11 – 1.13 × as long as wide; ii) frons between anterior ocellus and antennal sockets without or with at most single apically wavy to corkscrew-like pollen-collecting hairs (Fig. 34); iii) punctation of scutum between parapsidal line and median longitudinal furrow dense with interspaces not exceeding the diameter of one puncture; and iv) toothlike projection at apex of tibia of fore leg dark brown to black. The males (Fig. 32) additionally possess the following non-exclusive characters: i) antennal segment 3 about as long as maximally wide; ii) antennal segment 13 cylindrical and apically not distinctly curved downwards; iii) apical margin of sternum 3 with dense median patch of long whitish to yellowish hairs distinctly surpassing tergal margin and covering about two fifths of entire sternal width; iv) sternum 6 lateroapically with distinct and well-stepped albeit weak callosity, which slopes rather abruptly towards tergal centre (Fig. 35); shortest distance between inner margins of callosities shorter than to maximally as long as maximum width of callosity (Fig. 35); and v) sternum 8 apically triangularly incised.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573C4474D6A6FF4D3F598A63.taxon	description	Description. FEMALE (Fig. 31): Body length 5 – 6 mm. Head: Head 1.11 – 1.13 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.5 – 1.7 × as long as ocellar diameter. Maximum width of genal area 0.7 – 0.8 × as long as maximum width of compound eye. Second segment of labial palpus 2.1 – 2.3 × as long as first segment and 0.75 – 0.85 × as long as compound eye. Mandible three-toothed and blackish except for preapical reddish zone. Clypeus medially moderately bulged with foremost point in a distance of about one third of clypeal length from lower margin of supraclypeal area. In lateral view, transitional zone between bulging part and marginal zone of clypeus straight. Punctation of clypeus dense with interspaces reaching diameter of half a puncture to rarely one puncture except for narrow apical zone, which is unpunctured and polished. Clypeal punctures oblong rather than circular. Apical margin of clypeus medially sharp and almost straight. Pilosity of paraocular area white, rather dense and predominantly appressed. Clypeus densely covered with rather long and apically wavy to corkscrew-like pollen-collecting hairs. Punctation of supraclypeal area very dense with only linear interspaces except for medioapical zone, where punctation may be more scattered; punctures smaller than those on clypeus. Punctation of frons dense with interspaces rarely exceeding diameter of half a puncture except medially, where interspaces may reach up to one puncture; punctures slightly smaller than those on clypeus. Pilosity of frons similarly dense but slightly shorter than that of clypeus and without or at most single apically wavy to corkscrew-like hairs (Fig. 34). Antenna blackish except for anterior side of flagellum, which is more or less brownish. Antennal segment 3 – 9 distinctly wider than long, segments 10 – 11 almost as long as wide and segment 12 about 1.75 × as long as wide. Mesosoma: Parapsidal line long-linear. Punctation of scutum and scutellum dense with interspaces not exceeding diameter of one puncture, punctures on scutum smaller than those on frons. Punctation of mesepisternum more scattered with interspaces reaching diameter of up to one to one and a half punctures. Basal area of propodeum polished to superficially shagreened except for narrow transverse zone at its base, which is more strongly shagreened. Posterior surface of propodeum adjacent to basal area predominantly polished, its punctation varying in density from scattered beside propodeal pit to very dense on lateral parts. Tegula yellowish-brown on posterior half and more or less blackish on anterior half. Tibial spur of fore leg yellowish to light brown, its apex extended into well stepped tip, which is as long as to slightly longer than basally wide. Toothlike projection at apex of tibia of fore leg dark brown to black. Tibial spurs of middle and hind leg (light) brown to dark brown, long, slender and apically weakly curved. Pilosity on inner side of basitarsus of hind leg yellowish-white. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs dense with interspaces varying in size between diameter of less than half a puncture on lateral tergal sides and diameter of up to one puncture on median tergal parts, punctures of about same size as those on scutum. Marginal zones of terga 1 – 5 dark reddish. Marginal zones of terga 1 – 4 with short white hair bands, which are medially interrupted on terga 1 – 2 and decrease in length and density from tergum 1 to tergum 4 (Fig. 31). Scopa white (Fig. 31). MALE (Fig. 32): Body length 4.5 – 6 mm. Head: Head as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.3 – 1.4 × as long as ocellar diameter. Maximum width of genal area 0.5 – 0.6 × as long as maximum width of compound eye. Second segment of labial palpus 2.2 – 2.3 × as long as first segment and about 0.8 × as long as compound eye. Mandible two-toothed and black except for reddish preapical zone. Clypeus very finely and densely punctured with only linear interspaces except for narrow marginal zone, which is polished and unpunctured. Apical margin of clypeus straight to very shallowly emarginate and with three to five minute teeth. Lower half of frons, paraocular area, supraclypeal area and clypeus covered with dense yellowish-white pilosity, which is erect on frons and supraclypeal area, appressed on paraocular area and semierect on clypeus. Hypostomal area with moderately dense and rather long whitish pilosity, which increases in length towards foramen magnum. Punctation of frons very dense with interspaces varying in size between diameter of less than half a puncture on lateral parts to diameter of up to one puncture in front of anterior ocellus. Antenna dark brown to black except for ventral and posterior side of segments (8) 9 – 13, which are more or less yellowish-brown. Antennal segment 3 as long as wide and slightly longer than segment 4, which is slightly shorter than wide, segments 5 – 11 roughly as long as wide, segment 12 slightly longer than wide and segment 13 about 2 × as long as wide. Antennal segment 13 cylindrical and apically not curved downwards. Mesosoma: Parapsidal line long-linear. Punctation of scutum and scutellum dense with interspaces hardly exceeding diameter of one puncture, punctures on scutum slightly smaller than those on frons. Punctation of mesepisternum more scattered with interspaces reaching diameter of up to one to one and a half punctures. Basal area of propodeum polished to superficially shagreened except for narrow transverse zone at its base, which is more strongly shagreened. Posterior surface of propodeum adjacent to basal area predominantly polished, its punctation varying in density from very scattered beside propodeal pit to very dense on lateral parts. Tegula predominantly yellowish-brown. Tibial spur of fore leg yellowish, its apex extended into short tip, which is about as long as basally wide. Toothlike projection at apex of tibia of fore leg dark brown to black. Tibial spurs of middle and hind leg (light) brown to dark brown, long, slender and apically weakly curved. Veins of fore and hind wing (light) brown to dark brown. Stigma of fore wing (light) brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs 1 – 5 dense with interspaces varying in size between diameter of less than half a puncture on lateral tergal sides and diameter of up to one puncture on median tergal parts, punctures of about same size as those on scutum. Marginal zones of terga 1 – 6 reddish. Marginal zones of terga 1 – 4 (5) with short white hair bands, which are medially interrupted on terga 1 – 2 (Fig. 32). Tergum 6 with very short lateral tooth, its marginal zone slightly raised. Tergum 7 with roundish median pit at its base and medioapically elongated into narrowly triangular tip. Sternum 1 slightly convex. Apical margin of sternum 2 evenly rounded. Apical margin of sternum 3 medially emarginate and with dense patch of long whitish to yellowish hairs distinctly surpassing tergal margin and covering about two fifths of horizontal sternal width. Sternum 4 medioapically more deeply and more widely emarginate than sternum 3, emargination lined with yellowish-white hairs, which are similarly long as hair patch of sternum 3. Apical margin of sternum 5 medially very shallowly emarginate and inconspicuously ciliated with short whitish hairs. Sternum 6 basally with pair of translucent membraneous flaps and lateroapically with distinct and well-stepped albeit weak callosity, which slopes rather abruptly towards tergal centre (Fig. 35); shortest distance between inner margins of callosities shorter than to maximally as long as maximum callosity width (Fig. 35). Sternum 8 apically triangularly incised. Gonoforceps apically rounded and evenly tapering, without stepped tip (Fig. 36).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573C4474D6A6FF4D3F598A63.taxon	distribution	Distribution. Central and southern Spain.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573C4474D6A6FF4D3F598A63.taxon	etymology	Etymology. The species epithet refers to the diagnostic character of both sexes, i. e. the brown to dark brown tibial spurs of middle and hind legs (lat. “ fuscus ” = brown, lat. “ spina ” = spine).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573F446AD6A6F88C3E238FCF.taxon	materials_examined	Holotype. SPAIN: Almería: Nijar, 36 ° 58 ' 36 " N / 2 ° 12 ' 70 " W, 19.5.2019, ♂ (leg. J. Smit). Holotype deposited in the Entomological Collection of ETH Zurich. Paratypes. SPAIN: Almería: Nijar, 36 ° 59 ' 08 " N / 2 ° 14 ' 06 " W, 22.5.2018, 1 ♂ (leg. J. Smit); Granada: Sierra Nevada, Capileira to La Cebadilla, 36.9779 ° N / 3.3548 ° W, 1500 m, 8.6.2021, 1 ♂ (leg. T. Wood); Teruel: Noguera de Albarracin, Barranco de la Olmeda, 40.4616 ° N / 1.6139 ° W, 25.6.2021, 1 ♂ (leg. T. Wood). Paratypes deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573F446AD6A6F88C3E238FCF.taxon	diagnosis	Diagnosis. Hoplitis hispaniae is morphologically very close to H. andalusiae sp. nov. in the female sex and no morphological characters are currently known to separate the females of the two species. Among the European and North African Hoplitis species of the subgenus Micreriades, the females of H. hispaniae and H. andalusiae can be recognised by the following combination of characters: i) head 1.05 – 1.15 × as long as wide; ii) inner margins of compound eyes slightly converging towards clypeus; iii) clypeus basally moderately bulged: in lateral view, transitional zone between bulging part and marginal zone of clypeus straight to very slightly convex; iv) frons between anterior ocellus and antennal sockets with numerous apically wavy to corkscrew-like pollen-collecting hairs, which may be worn off in old individuals; v) punctation of scutum between parapsidal line and median longitudinal furrow rather scattered with interspaces often reaching the diameter of two, rarely more punctures; vi) toothlike projection at apex of tibia of fore leg yellowish-brown; and vii) tibial spurs of middle and hind leg yellowish. The males of H. hispaniae (Fig. 37) are characterised by the following combination of characters: i) head about as long as wide; ii) antennal segment 3 as long as maximally wide; iii) antennal segment 13 cylindrical and apically not distinctly curved downwards; iv) punctation of scutum between parapsidal line and median longitudinal furrow moderately dense with interspaces reaching the diameter of up to one and a half to two punctures; v) apical margin of sternum 3 with dense median patch of long whitish to yellowish hairs distinctly surpassing tergal margin and covering about half of entire sternal width; vi) sternum 6 lateroapically completely plane and without callosities (Fig. 38); vii) sternum 6 medioapically with short, weak and keel-like longitudinal elevation, its apical margin laterally roughly straight and medially emarginate (Fig. 38); and viii) gonoforceps apically rounded rather than truncate and regularly tapering rather than with stepped tip (Fig. 39).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573F446AD6A6F88C3E238FCF.taxon	description	Description. FEMALE: See female description under Hoplitis andalusiae above. MALE (Fig. 37): Body length 6 – 7 mm. Head: Head 0.97 – 1.05 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.4 – 1.5 × as long as ocellar diameter. Maximum width of genal area 0.50 – 0.55 × as long as maximum width of compound eye. Second segment of labial palpus 2.1 – 2.2 × as long as first segment and about 0.75 × as long as compound eye. Mandible two-toothed and completely black except sometimes for dark reddish preapical zone. Clypeus very finely and densely punctured with only linear interspaces except for narrow marginal zone, which is polished and unpunctured. Apical margin of clypeus medially straight to very shallowly emarginate and inconspicuously crenulate. Lower half of frons, paraocular area, supraclypeal area and clypeus covered with dense yellowish-white pilosity, which is (semi) erect on frons and supraclypeal area and appressed on paraocular area and clypeus. Hypostomal area with rather long and dense whitish pilosity, which increases in length towards foramen magnum. Punctation of frons very dense with interspaces rarely reaching diameter of one puncture. Antenna (dark) brown to black. Antennal segment 3 as long as wide and slightly longer than segment 4, which is slightly shorter than wide, segments 5 – 11 roughly as long as wide to slightly shorter, segment 12 slightly longer than wide and segment 13 1.75 – 2.0 × as long as wide. Antennal segment 13 cylindrical and apically not distinctly curved downwards. Posterior side of antennal segments 9 – 13 almost straight in top view. Mesosoma: Parapsidal line long-linear. Punctation of scutum between parapsidal line and median longitudinal furrow moderately dense with interspaces reaching diameter of up to one and a half to two punctures, punctures smaller than those on frons. Punctation of scutellum dense with interspaces rarely exceeding diameter of half a puncture. Punctation of upper part of mesepisternum moderately dense with interspaces reaching diameter of up to one and a half to two punctures, of lower part more scattered. Basal area of propodeum polished except for narrow transverse zone at its base, which is weakly carinate and shagreened. Posterior surface of propodeum adjacent to basal area polished with scattered and fine punctation. Tegula predominantly yellowish-brown. Tibial spur of fore leg yellowish, its apex extended into short tip, which is about as long as basally wide. Toothlike projection at apex of tibia of fore leg yellowish-brown. Tibial spurs of middle and hind leg yellowish, long, slender and almost straight. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs 1 – 5 dense with interspaces varying in size between diameter of less than half a puncture on lateral tergal sides and diameter of one and a half, rarely more punctures on median tergal parts, punctures of about same size as those on scutum. Marginal zones of terga 1 – 6 reddish. Marginal zones of terga 1 – 4 with short white hair bands, which are medially interrupted on terga 1 – 2 and decrease in length and density from tergum 1 to tergum 4 (Fig. 37). Tergum 6 with short lateral tooth, its marginal zone slightly raised. Tergum 7 with roundish median pit at its base and medioapically elongated into narrowly triangular tip. Sternum 1 strongly convex. Sternum 2 mediobasally flat to slightly concave and with distinctly shagreened interspaces between punctures, its apical margin evenly rounded and surpassed by moderately long and sparse white pilosity, which is about as long as hair bands of terga 3 – 4. Apical margin of sternum 3 medially very shallowly emarginate and with dense patch of whitish to yellowish hairs distinctly surpassing tergal margin and covering almost half of horizontal sternal width. Sternum 4 medioapically distinctly more deeply and slightly more widely emarginate than sternum 3, emargination lined with yellowish-white hairs (Fig. 38), which are similarly long as hair patch of sternum 3. Sternum 5 on marginal zone with rather dense but short and (semi) erect whitish pilosity, its apical margin medially slightly emarginate (Fig. 38). Sternum 6 basally with pair of translucent membraneous flaps, medioapically with short, weak and keel-like longitudinal elevation and lateroapically completely plane without callosities (Fig. 38). Apical margin of sternum 6 laterally straight and medially emarginate (Fig. 38). Gonoforceps apically rounded and evenly tapering, without stepped tip (Fig. 39).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573F446AD6A6F88C3E238FCF.taxon	distribution	Distribution. Central and southern Spain.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E573F446AD6A6F88C3E238FCF.taxon	etymology	Etymology. The species epithet refers to the occurrence of the species in Spain (lat. “ Hispania ” = Spain).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57204468D6A6FF4D38528D9F.taxon	materials_examined	Holotype. GREECE: Thessaly: Platania / Volos, 9.4.2005, ♂ (leg. K. Standfuss). Holotype deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57204468D6A6FF4D38528D9F.taxon	diagnosis	Diagnosis. Osmia inflatipes belongs morphologically to the group around Osmia (Helicosmia) leaiana (Kirby, 1802), as shown by the following combination of characters in the male sex (Fig. 40): i) lower half of basal area of propodeum polished and almost without shagreenation; ii) marginal zones of terga 1 – 2 with narrow unpunctured zone along tergal margins; iii) tergum 6 laterally angular but without tooth or projecting edge, resulting in a more or less straight connection between lateral corner and median part of tergal margin; iv) sternum 4 medioapically rounded, its preapical zone with rather dense and half erect to almost appressed yellowish pilosity; and v) gonoforceps apically weakly curved inwards, its inner side only slightly concave (Fig. 43). The males of Osmia inflatipes (Fig. 40) differ from the group around O. leaiana and all other European and North African Osmia (Helicosmia) species by the unusual shape and proportions of the leg segments, which is illustrated by the following characters: i) femur and tibia of hind leg strongly inflated with the femur being only about 1.75 × and the tibia about 2 × as long as maximally wide (Fig. 41); ii) basitarsus of fore leg only about 2 ×, of middle leg about 2.25 × and of hind leg about 2.5 × as long as maximally wide (Fig. 42); and iii) distance between base of tibia and apex of tarsal segment 5 (including claws) of hind leg shorter than metasoma. The female of O. inflatipes is unknown.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57204468D6A6FF4D38528D9F.taxon	description	Description. MALE (Fig. 40): Body length 9 mm. Head: Head 0.93 × as long as wide. Distance between posterior margin of lateral ocellus and preoccipital margin 1.8 × as long as ocellar diameter. Maximum width of genal area 0.9 × as long as maximum width of compound eye. Second segment of labial palpus 2.1 × as long as first segment and 0.75 × as long as compound eye. Mandible two-toothed and black. Apical margin of clypeus weakly rounded except for narrow median part, which is very shallowly emarginate and medially provided with almost imperceivable minute tooth. Frons, paraocular area, supraclypeal area, clypeus and hypostomal area covered with dense and predominantly erect to semierect pilosity, which is white on lower and yellowish on upper half of face. Punctation of frons very dense with only linear interspaces. Antenna dark brown to black. Antennal segment 3 0.85 × as long as wide and 0.75 × as long as segment 4; segments 4 – 12 slightly longer than wide and segment 13 1.6 × as long as wide. Ventral side of antennal segments 6 – 12 slightly convex. Mesosoma: Parapsidal punctiform. Punctation of scutum, scutellum and mesepisternum very dense with only linear interspaces, punctures slightly larger than those on frons. Basal area of propodeum in upper half with narrow zone that is longitudinally carinate and not separated by distinct transverse carina from adjacent lower half, which is polished and unpunctured. Posterior surface of propodeum adjacent to basal area densely punctured with predominantly polished interspaces. Tegula dark brown. Scutum, scutellum and mesepisternum covered with long and erect pilosity, which is yellowish except for hairs on lower half of mesepisternum, which are white. Tibial spur of fore leg yellowish, its apex extended into long and well stepped tip, which is more than 2 × as long as basally wide. Tibial spurs of middle and hind leg brown, long, slender and almost straight. Femur and tibia of hind leg strongly inflated with femur being about 1.75 × and tibia about 2 × as long as maximally wide (Fig. 41). Basitarsus of fore leg about 2 ×, of middle leg about 2.25 × and of hind leg about 2.5 × as long as maximally wide (Fig. 42). Distance between base of tibia and apex of tarsal segment 5 (including claws) of hind leg shorter than metasoma. Pilosity on inner side of basitarsus of hind leg yellowish-brown. Veins of fore and hind wing brown to dark brown. Stigma of fore wing brown in centre and dark brown along margins. Metasoma: Punctation of tergal discs moderately dense with interspaces varying in size between diameter of half a puncture or less on lateral tergal sides and diameter of up to two or three punctures on median parts of terga 2 – 4, punctures slightly smaller than those on scutum. Marginal zones of terga 1 – 5 polished and largely unpunctured. Terga 3 – 5 with loose yellowish hair bands, which do not hide cuticular surface and reach apical margin on tergum 3 or slightly surpass apical margin on terga 4 – 5. Tergum 6 mediopreapically weakly impressed and laterally angular but not protruding, its apical margin medially with small half-elliptical incision. Apical margin of tergum 7 evenly rounded and medially with semicircular incision, whose inner margin is rather densely beset with yellowish hairs (Fig. 43). Sternum 1 weakly convex, its apical margin straight and ciliated with short whitish hairs. Sternum 2 mediopreapically weakly swollen, its apical margin evenly rounded and medially very slightly impressed. Sternum 3 with deep triangular incision, which is lined with yellowish hairs that are directed inwards and do not reach apical tergal margin; maximal width of incision about half as wide as horizontal sternal width. Sternum 4 medioapically rounded, its preapical zone with rather dense and semierect to almost appressed yellowish pilosity, which covers almost half of horizontal sternal width. Apical margin of sternum 5 almost straight and ciliated with short whitish hairs. Genitalia as in Fig. 43.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57204468D6A6FF4D38528D9F.taxon	distribution	Distribution. Eastern Thessaly in Greece.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57204468D6A6FF4D38528D9F.taxon	etymology	Etymology. The species epithet refers to the diagnostic character of the male, i. e. the inflated femur and tibia of the hind legs (lat. “ inflatus ” = inflated, lat. “ pes ” = foot).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FB2939D48BFF.taxon	discussion	Ducke (1899) described Hoplitis papaveris convolvuli as a variety of Hoplitis papaveris (Latreille, 1799) due to slight morphological differences and its pollen specialization to flowers of Convolvulus (Convolvulaceae). Ducke’s taxon was later synonymized by Tkalců (1969) with Hoplitis perezi (Ferton, 1895), which has a wide distribution ranging from the southwestern Mediterranean region over southern Europe and Turkey to Central Asia. Müller (2022) made a nomenclatural mistake and produced a nomen praeoccupatum, when he named a Moroccan Hoplitis species as Hoplitis convolvuli Müller, 2022, a species that is most probably endemic to Morocco and not conspecific with H. perezi. This mistake is corrected here by proposing the following replacement name for the Moroccan endemic: Hoplitis (Anthocopa) trabutiana nom. nov .. The species epithet refers to Convolvulus trabutianus, which is used by the females of this species as an important source for both pollen and brood cell building material and by the males as rendezvous place (Müller 2022).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FE913A5289C7.taxon	materials_examined	Holotype. ITALY: Sardinia: Cala Fuili, 40.256 ° N / 9.624 ° E, 30 m, 28.4.2018, ♀ (leg. D. Bénon). Holotype deposited in the Entomological Collection of ETH Zurich.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FE913A5289C7.taxon	diagnosis	Diagnosis. The females of Osmia sardoa (Fig. 44), which have a body length of 9 – 10 mm, are morphologically very similar to those of O. leaiana, both differ by the following characters from closely related O. (Helicosmia) species: i) apical margin of clypeus medially with two or rarely three teeth, which usually enclose two narrow tufts of yellowish hairs (Fig. 46); ii) base of mandible supramedially with distinct polished bump; and iii) interspaces between punctures of posterior surface of propodeum distinctly shagreened. The females of O. sardoa (Fig. 44) can easily be separated from those of O. leaiana by the following characters: i) pilosity of head, mesosoma and terga foxy red and of same colour as metasomal scopa (Fig. 44, 45; pilosity whitish to yellowish and of distinctly lighter colour than orange metasomal scopa in O. leaiana); ii) marginal zones of terga 1 – 5 with dense foxy-red pilosity forming distinct hair bands (Fig. 45; marginal zones with very sparse whitish to yellowish pilosity not forming distinct hair bands in O. leaiana); iii) pilosity of tergal discs 2 – 4 longer with longest hairs distinctly longer than tarsal segment 3 of hind leg (pilosity shorter with longest hairs about as long as tarsal segment 3 of hind leg in O. leaiana); and iv) metallic sheen of body more extended: central face, mesepisternum and scutellum slightly green metallic and clypeus, gena vertex, scutum, propodeum and terga partly weakly purple metallic (metallic sheen usually limited to terga, which are partly weakly green metallic in O. leaiana). The male of O. sardoa is unknown.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FE913A5289C7.taxon	description	Description. Except for the characters given in the diagnosis, the females of O. sardoa are morphologically identical to those of the widespread and well-known O. leaiana. Therefore, no detailed description of the new species is given here.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FE913A5289C7.taxon	distribution	Distribution. Sardinia.	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
B81AB61E57234468D6A6FE913A5289C7.taxon	etymology	Etymology. The species epithet refers to the occurrence of the species on Sardinia (lat. “ sardous ” = Sardinian).	en	Müller, Andreas, Wood, Thomas J. (2025): Still undescribed bees in Europe-description of nine new European osmiine bee species (Hymenoptera, Megachilidae). Zootaxa 5631 (1): 83-102, DOI: 10.11646/zootaxa.5631.1.3, URL: https://doi.org/10.11646/zootaxa.5631.1.3
