identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
62093EBB9A62525C81D2B2CB14830E6D.text	62093EBB9A62525C81D2B2CB14830E6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites acuminatus Attems 1937	<div><p>Kronopolites acuminatus Attems, 1937</p><p>Fig. 1</p><p>Kronopolites acuminatus Attems, 1937: 52; Attems 1938: 227, fig. 53; Jeekel 1968: 59; Golovatch 2009: 121, in key;</p><p>Kronopolites acuminatus acuminatus: – Hoffman 1963: 584, established new subspecific status; Golovatch 1983: 181; Enghoff et al. 2004: 38.</p><p>Type specimens.</p><p>NHMW.</p><p>Material examined.</p><p>Vietnam • Lao Cai Province • 1 male, 2 females; Hoang Lien National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.7803&amp;materialsCitation.latitude=22.35087" title="Search Plazi for locations around (long 103.7803/lat 22.35087)">on the way to Fanxipan Mt.</a>; 22.35087°N, 103.78030°E; primary forest; 2,000 m a. s. l.; 28 September 2005; Anh D. Nguyen leg.; IEBR-Myr 115 • 1 male, 1 female; Hoang Lien National Park, Tram Ton, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.77541&amp;materialsCitation.latitude=22.35347" title="Search Plazi for locations around (long 103.77541/lat 22.35347)">Tram Ton station, observation tower</a>; 22.35347°N, 103.77541°E; secondary forest; 1,900 m a. s. l.; 23 March 2007; Anh D. Nguyen leg.; IEBR-Myr 187 • 1 male, 2 females; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.64392&amp;materialsCitation.latitude=22.61148" title="Search Plazi for locations around (long 103.64392/lat 22.61148)">Bat Xat Natural Reserve</a>; 22.61148°N, 103.64392°E; natural forest; 28 December 2017; Hung D. Nguyen leg.; IEBR-Myr 760 • 1 male, 1 female; same data as IEBR-Myr 760; IEBR-Myr 761 .</p><p>Diagnosis.</p><p>The species can be recognised by having sternite 5 with a rounded, highly elevated, setiferous process between coxae 4, and gonopod conformation (process a leaf – shaped, shorter than slender, spiniform process b).</p><p>Distribution.</p><p>Ha Giang Province (Ha Giang) (Attems 1937, 1938), Lao Cai (this study).</p><p>Remarks.</p><p>The species has been known only from northern Vietnam.</p></div>	https://treatment.plazi.org/id/62093EBB9A62525C81D2B2CB14830E6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
77148E6FFFCD55A18C40426FE391D158.text	77148E6FFFCD55A18C40426FE391D158.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites Attems 1914	<div><p>Genus Kronopolites Attems, 1914</p><p>Type species.</p><p>Kronopolites svenhedini (Verhoeff, 1934) corrected by Xiong et al. (2025).</p><p>Remarks.</p><p>The genus was recently revised by Likhitrakarn et al. (2015). The genus can be recognised by the gonopod solenophore typically carrying a fork consisting of two basal processes: curved or suberect, anteriad-directed process a and posteriad-directed process b.</p><p>Most species of Kronopolites live in montane forest habitats usually found at elevations between ca 400 m and 2,700 m a. s. l.</p></div>	https://treatment.plazi.org/id/77148E6FFFCD55A18C40426FE391D158	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
A00CB8421B0F578394789767295777C3.text	A00CB8421B0F578394789767295777C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites biagrilectus Hoffman 1963	<div><p>Kronopolites biagrilectus Hoffman, 1963</p><p>Fig. 4</p><p>Material examined.</p><p>Vietnam • 1 male; Dien Bien Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.38603&amp;materialsCitation.latitude=22.29225" title="Search Plazi for locations around (long 102.38603/lat 22.29225)">Muong Nhe Natural Reserve</a>; 22.29225°N, 102.38603°E; 784 m a. s. l.; natural forest; 7 May 2018; Hung D. Nguyen leg.; IEBR-Myr 756 .</p><p>Diagnosis.</p><p>The species is similar to K. acuminatus (Attems, 1937), but distinguished by the form of gonopod processes a and b (process a shorter than process b in K. acuminatus, vice versa in K. biagrilectus).</p><p>Distribution.</p><p>Widely distributed in southern China (Golovatch 2020) and Vietnam (this study).</p><p>Remarks.</p><p>The species was proposed as a subspecies, K. acuminatus biagrilectus Hoffman, 1963, and considered as a northern form of K. acuminatus acuminatus . The only difference between the two species is the length ratio of processes a and b. The species is recorded herewith from Vietnam for the first time. Our specimen fits well the description of Hoffman (1963).</p></div>	https://treatment.plazi.org/id/A00CB8421B0F578394789767295777C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
8889CB4C39B056689152F2909B07B2B6.text	8889CB4C39B056689152F2909B07B2B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites contrastus Nguyen & Vu & Phung & Tran & Le 2025	<div><p>Kronopolites contrastus sp. nov.</p><p>Figs 5, 6, 7, 8</p><p>Material examined.</p><p>Holotype: Vietnam • 1 male; Tuyen Quang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.11154&amp;materialsCitation.latitude=22.20248" title="Search Plazi for locations around (long 105.11154/lat 22.20248)">Cham Chu Nature Reserve</a>; 22.20248°N, 105.11154°E; limestone forest; July 2018; Dai D. Nguyen leg.; IEBR-Myr 718.</p><p>Diagnosis.</p><p>The species differs from its congeners in colouration pattern (tergites mostly dark while other parts of body whitish yellow), gonopod conformation (lamina l present laterally, subrectangular; processes a and b both leaf-shaped, but pointed; a shorter than b in length; process a subhelicoid. Solenophore clearly curved, long, expanded distomesally, bipartite. Solenomere longer than solenophore, ribbon-shaped, coiled).</p><p>Description.</p><p>Size: Length 48.38 mm, width of midbody pro- and metazona 3.41 and 4.87 mm, respectively.</p><p>Colouration: tergites mostly dark; head blackish brown; prozonae, pleurites, sternites, antennae, telson, and legs whitish yellow (Figs 5, 6).</p><p>Head: Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae moderately long (Fig. 5 A, B), extending behind body segment 3 when stretched laterally; antennomere 2 = 3 = 4&gt; 6&gt; 5&gt; 7 = 1, antennomere 7 with four conical sensories.</p><p>Collum: traces of setae hardly seen; lateral incisions absent; caudal corner of paraterga very broadly rounded, declined ventrad, produced behind rear tergal margin (Fig. 5 A, B).</p><p>Body rings: In width, segment 4 &lt;3 &lt;head &lt;5 &lt;collum &lt;body ring 2 &lt;6–17, thereafter body gently and gradually tapering. Tegument smooth and shining, prozonae finely shagreened, metaterga finely rugulose (Figs 5 A – D, 6 A, B); surface below paraterga finely microgranulate (Fig. 5 C, F). Tergal setae all broken, traces hardly visible. Axial line distinct on anterior halves of metazonae (Figs 5 D, 6 A). Transverse sulcus usually distinct (Fig. 5 D), slightly incomplete on body rings 4 and 19, complete on metaterga 5–18, narrow, line-shaped, shallow, reaching bases of paraterga. Stricture between pro – and metazonae evident, broad and deep, ribbed at bottom down to base of paraterga (Figs 5 D, 6 A). Pleurosternal carinae complete crests with a sharp caudal tooth on body rings 2–7 (Fig. 5 C, F), thereafter increasingly strongly reduced until body ring 17.</p><p>Paraterga strongly developed, lying rather high (at upper 1 / 3 of body), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, starting from segment 15 extending increasingly well beyond rear tergal margin (Figs 5 C, D, F, 6 A, B); lateral edge without incisions; posterior edge nearly straight. Calluses on paraterga narrow, delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, lying in an ovoid groove at ~ 1 / 4 in front of posterior edge of metaterga (Figs 5 C, D, 6 A).</p><p>Telson: Epiproct (Figs 5 F, 6 A – C) conical, flattened dorsoventrally, with two small apical papillae; tip subtruncate; pre – apical papillae small, lying close to tip. Hypoproct sub – semicircular, setiferous knobs at caudal edge small and well – separated (Fig. 6 C).</p><p>Sterna: densely setose, without modifications except sternum 5 th with a bifid tongued – shaped, setose cone between male coxae 4 (Fig. 6 D).</p><p>Legs (Fig. 6 C, F): rather long and slender, midbody ones ~ 1.2–1.3 times as long as body height; prefemora without modifications, tarsal brushes present on pregonopodal legs.</p><p>Gonopods (Figs 7, 8) typical Kronopolites species; coxite (co) long, subcylindrical, a little curved caudad, sparsely setose distoventrally. Prefemur (pref) densely setose, ~ 1 / 3 as long as femorite + postfemoral part. Femorite (fe) long, cylindrical, slightly constricted medially, simple without any modifications; mesal side strongly grooved; lateral side with a distinct transverse sulcus demarcating the postfemoral region. Postfemoral part well developed; lamina l present laterally, subrectangular; processes a and b both leaf-shaped, but pointed; a shorter than b in length; process a subhelicoid. Solenophore clearly curved, long, expanded distomesally, bipartite. Solenomere longer than solenophore, ribbon-shaped, coiled.</p><p>Etymology.</p><p>Contrastus, an adjective epithet is used to emphasise the contrast colouration of body.</p><p>Remarks.</p><p>The new species is similar to K. fuscocingulatus from northern Thailand by having a sternal lobe between male coxae 4, and processes a and b of gonopod being nearly independent, subequal in length. However, two species are distinguished by shape of sternal lobe (bifid tongue-shaped vs roundly subquadrate), and shape of process a and b (spiniform, blunt, short, and stout vs ribbon-shaped, blunt, slender and long).</p></div>	https://treatment.plazi.org/id/8889CB4C39B056689152F2909B07B2B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
F7CEF392741C5373A2B275947A95601E.text	F7CEF392741C5373A2B275947A95601E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites montanus Golovatch 2009	<div><p>Kronopolites montanus Golovatch, 2009</p><p>Fig. 2</p><p>Kronopolites montanus Golovatch, 2009: 121, figs 9–16; Nguyen and Sierwald 2013: 1288; Likhitrakarn et al. 2015: 31.</p><p>Type specimens.</p><p>ZMUM.</p><p>Material examined.</p><p>Vietnam • Son La Province • 1 male; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.67119&amp;materialsCitation.latitude=20.721195" title="Search Plazi for locations around (long 104.67119/lat 20.721195)">Xuan Nha Natural Reserve</a>; 20.7211944°N, 104.67119°E; 566 m a. s. l.; residential area; 19 May 2018; Hung D. Nguyen leg.; IEBR-Myr 266 • 2 males; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.49196&amp;materialsCitation.latitude=21.33535" title="Search Plazi for locations around (long 104.49196/lat 21.33535)">Ta Xua Natural Reserve</a>; 21.33535°N, 104.49196°E; 490 m a. s. l.; bamboo forest; 9 February 2017; Son X. Le &amp; Ha T. Vu leg.; IEBR-Myr 581 • 4 males, 3 females, 2 juveniles; same locality as IEBR-Myr 581; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.36719&amp;materialsCitation.latitude=21.29298" title="Search Plazi for locations around (long 104.36719/lat 21.29298)">around Chieu village, near irrigation lake</a>; 21.29298°N, 104.36719°E; 450 m a. s. l.; 11 February 2017; IEBR-Myr 582 • 1 male, 1 female; same locality as IEBR-Myr 581; 21.33978°N, 104.68736°E; 400 m a. s. l.; cultivated land; 10 February 2017; Son X. Le &amp; Ha T. Vu leg.; IEBR-Myr 584 • 2 males, 1 female; same locality as IEBR-Myr 581; mixed forest; 30 May 2018; Hung D. Nguyen leg.; IEBR-Myr 752 • 2 males, 1 female; same locality as IEBR-Myr 581; mixed forest; 30 April 2018; Hung D. Nguyen leg.; IEBR-Myr 753 • 1 male; same locality as IEBR-Myr 581; natural forest; 30 May 2018; Hung D. Nguyen leg.; IEBR-Myr 757; Lao Cai Province • 1 male, 5 females, 2 juveniles; Van Ban District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.00361&amp;materialsCitation.latitude=22.03536" title="Search Plazi for locations around (long 104.00361/lat 22.03536)">Nam Xay commune</a>; 22.03536°N, 104.00361°E; 600–700 m a. s. l.; bamboo forest; 28 March 2005; Anh D. Nguyen leg.; IEBR-Myr 110 • 1 male; same locality as IEBR-Myr 110; bamboo forest; 900 m a. s. l.; 12 April 2005; Anh D. Nguyen leg.; IEBR-Myr 112 • 1 male; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.78133&amp;materialsCitation.latitude=22.41475" title="Search Plazi for locations around (long 103.78133/lat 22.41475)">Hoang Lien National Park</a>; 22.41475°N, 103.78133°E; 2,000 m a. s. l.; nature forest; 20–29 March 2007; Anh D. Nguyen leg.; IEBR-Myr 186 • 1 male; same locality as IEBR-Myr 186; 1,900 m a. s. l.; bamboo forest; 20–29 March 2007; Anh D. Nguyen leg.; IEBR-Myr 189 • 1 male, 1 female; Hoang Lien National Park; 1,526 m a. s. l.; mixed forest; 16 March 2018; Hung D. Nguyen leg.; IEBR-Myr 763 ; Phu Tho Province • 1 male; Xuan Son National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.97299&amp;materialsCitation.latitude=21.12521" title="Search Plazi for locations around (long 104.97299/lat 21.12521)">on the way to Lang village</a>; 21.12521°N, 104.97299°E; 16 March 2006; Nguyen Van Quang leg.; IEBR-Myr 26 • 1 male, 1 female; same locality as the IEBR-Myr 26; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.93149&amp;materialsCitation.latitude=21.13776" title="Search Plazi for locations around (long 104.93149/lat 21.13776)">Lap village</a>; 21.13776°N, 104.93149°E; 17 January 2006; Nguyen Van Quang leg.; IEBR-Myr 114 • 1 male, 1 female; Xuan Son National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.93149&amp;materialsCitation.latitude=21.13776" title="Search Plazi for locations around (long 104.93149/lat 21.13776)">Lap village</a>; 21.13776°N, 104.93149°E; pitfall trapping; 6 April 2011; An &amp; Luong leg.; IEBR-Myr 173 .</p><p>Diagnosis.</p><p>The species can be recognised by black-brown colouration, the absence of cones or lamina between coxae 4 on sternite 5, and gonopod conformation (process a shorter and coiled, while process b longer, straight, digitiform and subhelicoid). The species differs from its close congener, K. ramosus, in shape and length of process a and b (a short, wider coiled process a and a longer, straight, digitiform, subhelicoid process b in K. montanus vs a short and thin, curved, process a with an apical hook and a longer, straight, acuminate process b in K. ramosus).</p><p>Distribution.</p><p>Lao Cai Province (Sapa) (Golovatch 2009); Son La Province (Xuan Nha NR; Ta Sua NR); Phu Tho Province (Xuan Son NP) (this study).</p><p>Remarks.</p><p>The species is widely distributed in northwestern Vietnam.</p></div>	https://treatment.plazi.org/id/F7CEF392741C5373A2B275947A95601E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
229EDB61ACDB56A9A58065B36AD7244F.text	229EDB61ACDB56A9A58065B36AD7244F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites ramosus Golovatch & Semenyuk 2021	<div><p>Kronopolites ramosus Golovatch &amp; Semenyuk, 2021</p><p>Fig. 3</p><p>Kronopolites ramosus Golovatch &amp; Smenyuk, 2021: 479, figs 31–46.</p><p>Type specimens.</p><p>ZMUM.</p><p>Material examined.</p><p>Vietnam • Ha Tinh Province • 1 male, 1 female; Huong Son District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15382&amp;materialsCitation.latitude=18.46268" title="Search Plazi for locations around (long 105.15382/lat 18.46268)">Son Kim commune</a>; 18.46268°N, 105.15382°E; 350 m a. s. l.; natural forests; 3 May 2004; Anh D. Nguyen leg.; IEBR-Myr 111 ; Nghe An Province • 3 males, 6 females, 1 juvenile; Pu Mat National Park; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.63714&amp;materialsCitation.latitude=19.07991" title="Search Plazi for locations around (long 104.63714/lat 19.07991)">Khe Thoi</a>; 19.07991°N, 104.63714°E; closed forest, near stream; 4–10 April 2011; Anh D. Nguyen leg; IEBR-Myr 175 • 7 males, 5 females; same as the sample IEBR-Myr 175; IEBR-Myr 176 • 2 females, 2 juveniles; Pu Mat National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.80081&amp;materialsCitation.latitude=18.9715" title="Search Plazi for locations around (long 104.80081/lat 18.9715)">Thac Chem waterfall</a>; 18.97150°N, 104.80081°E; 430 m a. s. l.; evergreen closed forest; 4–10 April 2011; Anh D. Nguyen leg.; IEBR-Myr 199 • 5 males, 3 females; Pu Mat National Park; same as the sample IEBR-Myr 175; IEBR-Myr 553 • 1 male; Pu Mat National Park; same as the sample IEBR-Myr 175; IEBR-Myr 174 .</p><p>Diagnosis.</p><p>The species can be distinguished by having a uniformly dark colouration and yellow legs, sternite 5 without any processes between male coxae 4, gonopod process a unciform while process b straight, both a and b slender and acuminate, sharing a broad lobe-shaped base.</p><p>Distribution.</p><p>Nghe An Province (Pu Mat NP) (Golovatch and Semenyuk 2021).</p><p>Remarks.</p><p>The species has been known only from Vietnam.</p><p>Remarks.</p><p>As mentioned above, two species, K. montanus and K. ramosus, show high similarity in morphology, especially gonopod conformation (see Figs 2, 3). The two species can be differentiated based on their gonopod processes: K. montanus has a short, wider coiled process a and a longer, straight, digitiform, subhelicoid process b, while K. ramosus has a short and thin, curved, process a with an apical hook and a longer, straight, acuminate process b. In addition, two species have high significant genetic divergence (COI distance = 11.6 %). Therefore, two species are clearly distinguished from each other.</p></div>	https://treatment.plazi.org/id/229EDB61ACDB56A9A58065B36AD7244F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
02E7C2BCC3ED5697A8FC4ED8B07F8982.text	02E7C2BCC3ED5697A8FC4ED8B07F8982.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kronopolites serratus Nguyen & Vu & Phung & Tran & Le 2025	<div><p>Kronopolites serratus sp. nov.</p><p>Figs 9, 10, 11</p><p>Material examined.</p><p>Holotype: Vietnam • 1 male; Lao Cai Province, Hoang Lien National Park, Thac Bac waterfall; 22.36350°E, 103.77754°E; 1,950 m a. s. l.; regenerated forest; 28 November 2005; Anh D. Nguyen leg.; IEBR-Myr 109.</p><p>Diagnosis.</p><p>The new species is similar to K. montanus Golovatch, 2009, but differs in the following characters: gonopod process a curved, serrated, acuminate leaf-shaped, as long as distally serrated, ribbon-shaped process b.</p><p>Description.</p><p>Size: length 32.01 mm, width of midbody pro- and metazona 2.67 mm and 3.67 mm, respectively.</p><p>Colouration (Fig. 9): almost uniformly brownish yellow after long preservation in ethanol.</p><p>Head (Fig. 9 A – C): Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae claviform, moderately long (Fig. 9 A – C), extending behind body segment 3 when stretched laterally; antennomere 2 = 3 = 4&gt; 5&gt; 6&gt; 7 = 1, antennomere 7 with four conical sensories.</p><p>Collum: traces of setae hardly seen; lateral incisions absent; caudal corner of paraterga very broadly rounded, declined ventrad, produced behind rear tergal margin (Fig. 9 A, B).</p><p>Body rings: In width, segment 4 &lt;3 &lt;head &lt;5 &lt;segment 2 &lt;collum &lt;6–17, thereafter body gently and gradually tapering. Tegument (Fig. 9 A – E) smooth and shining, prozonae finely shagreened; surface below paraterga finely rugulose. Tergal setae all broken, traces hardly seen. Axial line visible, especially on midbody tergites. Transverse sulcus usually distinct (Fig. 9 A), slightly incomplete on body rings 4 and 19, complete on metaterga 5–18, narrow, line – shaped, shallow, reaching bases of paraterga. Stricture between pro – and metazonae evident, broad and deep, striolated at bottom down to base of paraterga (Fig. 9 D). Pleurosternal carinae complete crests on body rings 2–7; thereafter, increasingly strongly reduced until body ring 17.</p><p>Paraterga strongly developed (Fig. 9 A, D), lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded; lateral edge without incisions; posterior edge nearly straight. Calluses on paraterga narrow, delimited by a sulcus both dorsally and ventrally. Ozopores (Fig. 9 D, E) evident, lateral, lying in an ovoid groove at ~ 1 / 4 in front of posterior edge of metaterga.</p><p>Telson: damaged.</p><p>Sterna: densely setose, without modifications (Fig. 9 C, F).</p><p>Legs: rather long and slender, midbody ones ~ 1.2–1.3 times as long as body height; prefemora without modifications, tarsal brushes present only on pregonopodal legs</p><p>Gonopods (Figs 10, 11) a typical Kronopolites species; coxite (co) stout, subcylindrical, sparsely setose distoventrally. Prefemur (pref) densely setose, ~ 1 / 3 as long as femorite + postfemoral part. Femorite somewhat stout, evidently grooved mesally, demarcated from postfemoral region by a distinct oblique sulcus laterally. Lamina l present, producing into a small spine z distally; process a leaf-shaped, mesal margin serrated, clearly shorter than distally-serrated, ribbon-shaped process b; solenophore well developed, bipartite, longer than a short flagelliform solenomere which parly sheathed by solenophore.</p><p>Remarks.</p><p>Although the new species is described based on only a male, its gonopod distinctly differs from that of K. montanus in gonopod process a being curved, serrated, acuminate leaf-shaped, as long as distally serrated, ribbon-shaped process b. Both species are also found in Hoang Lien National Park at the high elevation of more than 1,900 m a. s. l.</p><p>Etymology.</p><p>Serratus, an adjective epithet, is used to emphasise the serrated process a and b of the gonopod.</p></div>	https://treatment.plazi.org/id/02E7C2BCC3ED5697A8FC4ED8B07F8982	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nguyen, Anh D.;Vu, Tam T. T.;Phung, Hong Luong T.;Tran, Duc-Luong;Le, Hung-Anh	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
