taxonID	type	description	language	source
62093EBB9A62525C81D2B2CB14830E6D.taxon	description	Fig. 1	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
62093EBB9A62525C81D2B2CB14830E6D.taxon	diagnosis	Diagnosis. The species can be recognised by having sternite 5 with a rounded, highly elevated, setiferous process between coxae 4, and gonopod conformation (process a leaf – shaped, shorter than slender, spiniform process b).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
62093EBB9A62525C81D2B2CB14830E6D.taxon	distribution	Distribution. Ha Giang Province (Ha Giang) (Attems 1937, 1938), Lao Cai (this study).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
77148E6FFFCD55A18C40426FE391D158.taxon	type_taxon	Type species. Kronopolites svenhedini (Verhoeff, 1934) corrected by Xiong et al. (2025).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
A00CB8421B0F578394789767295777C3.taxon	description	Fig. 4	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
A00CB8421B0F578394789767295777C3.taxon	diagnosis	Diagnosis. The species is similar to K. acuminatus (Attems, 1937), but distinguished by the form of gonopod processes a and b (process a shorter than process b in K. acuminatus, vice versa in K. biagrilectus).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
A00CB8421B0F578394789767295777C3.taxon	distribution	Distribution. Widely distributed in southern China (Golovatch 2020) and Vietnam (this study).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
8889CB4C39B056689152F2909B07B2B6.taxon	description	Figs 5, 6, 7, 8	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
8889CB4C39B056689152F2909B07B2B6.taxon	diagnosis	Diagnosis. The species differs from its congeners in colouration pattern (tergites mostly dark while other parts of body whitish yellow), gonopod conformation (lamina l present laterally, subrectangular; processes a and b both leaf-shaped, but pointed; a shorter than b in length; process a subhelicoid. Solenophore clearly curved, long, expanded distomesally, bipartite. Solenomere longer than solenophore, ribbon-shaped, coiled).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
8889CB4C39B056689152F2909B07B2B6.taxon	description	Description. Size: Length 48.38 mm, width of midbody pro- and metazona 3.41 and 4.87 mm, respectively. Colouration: tergites mostly dark; head blackish brown; prozonae, pleurites, sternites, antennae, telson, and legs whitish yellow (Figs 5, 6). Head: Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae moderately long (Fig. 5 A, B), extending behind body segment 3 when stretched laterally; antennomere 2 = 3 = 4> 6> 5> 7 = 1, antennomere 7 with four conical sensories. Collum: traces of setae hardly seen; lateral incisions absent; caudal corner of paraterga very broadly rounded, declined ventrad, produced behind rear tergal margin (Fig. 5 A, B). Body rings: In width, segment 4 <3 <head <5 <collum <body ring 2 <6 – 17, thereafter body gently and gradually tapering. Tegument smooth and shining, prozonae finely shagreened, metaterga finely rugulose (Figs 5 A – D, 6 A, B); surface below paraterga finely microgranulate (Fig. 5 C, F). Tergal setae all broken, traces hardly visible. Axial line distinct on anterior halves of metazonae (Figs 5 D, 6 A). Transverse sulcus usually distinct (Fig. 5 D), slightly incomplete on body rings 4 and 19, complete on metaterga 5 – 18, narrow, line-shaped, shallow, reaching bases of paraterga. Stricture between pro – and metazonae evident, broad and deep, ribbed at bottom down to base of paraterga (Figs 5 D, 6 A). Pleurosternal carinae complete crests with a sharp caudal tooth on body rings 2 – 7 (Fig. 5 C, F), thereafter increasingly strongly reduced until body ring 17. Paraterga strongly developed, lying rather high (at upper 1 / 3 of body), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, starting from segment 15 extending increasingly well beyond rear tergal margin (Figs 5 C, D, F, 6 A, B); lateral edge without incisions; posterior edge nearly straight. Calluses on paraterga narrow, delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, lying in an ovoid groove at ~ 1 / 4 in front of posterior edge of metaterga (Figs 5 C, D, 6 A). Telson: Epiproct (Figs 5 F, 6 A – C) conical, flattened dorsoventrally, with two small apical papillae; tip subtruncate; pre – apical papillae small, lying close to tip. Hypoproct sub – semicircular, setiferous knobs at caudal edge small and well – separated (Fig. 6 C). Sterna: densely setose, without modifications except sternum 5 th with a bifid tongued – shaped, setose cone between male coxae 4 (Fig. 6 D). Legs (Fig. 6 C, F): rather long and slender, midbody ones ~ 1.2 – 1.3 times as long as body height; prefemora without modifications, tarsal brushes present on pregonopodal legs. Gonopods (Figs 7, 8) typical Kronopolites species; coxite (co) long, subcylindrical, a little curved caudad, sparsely setose distoventrally. Prefemur (pref) densely setose, ~ 1 / 3 as long as femorite + postfemoral part. Femorite (fe) long, cylindrical, slightly constricted medially, simple without any modifications; mesal side strongly grooved; lateral side with a distinct transverse sulcus demarcating the postfemoral region. Postfemoral part well developed; lamina l present laterally, subrectangular; processes a and b both leaf-shaped, but pointed; a shorter than b in length; process a subhelicoid. Solenophore clearly curved, long, expanded distomesally, bipartite. Solenomere longer than solenophore, ribbon-shaped, coiled.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
8889CB4C39B056689152F2909B07B2B6.taxon	etymology	Etymology. Contrastus, an adjective epithet is used to emphasise the contrast colouration of body.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
F7CEF392741C5373A2B275947A95601E.taxon	description	Fig. 2	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
F7CEF392741C5373A2B275947A95601E.taxon	diagnosis	Diagnosis. The species can be recognised by black-brown colouration, the absence of cones or lamina between coxae 4 on sternite 5, and gonopod conformation (process a shorter and coiled, while process b longer, straight, digitiform and subhelicoid). The species differs from its close congener, K. ramosus, in shape and length of process a and b (a short, wider coiled process a and a longer, straight, digitiform, subhelicoid process b in K. montanus vs a short and thin, curved, process a with an apical hook and a longer, straight, acuminate process b in K. ramosus).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
F7CEF392741C5373A2B275947A95601E.taxon	distribution	Distribution. Lao Cai Province (Sapa) (Golovatch 2009); Son La Province (Xuan Nha NR; Ta Sua NR); Phu Tho Province (Xuan Son NP) (this study).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
229EDB61ACDB56A9A58065B36AD7244F.taxon	description	Fig. 3	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
229EDB61ACDB56A9A58065B36AD7244F.taxon	diagnosis	Diagnosis. The species can be distinguished by having a uniformly dark colouration and yellow legs, sternite 5 without any processes between male coxae 4, gonopod process a unciform while process b straight, both a and b slender and acuminate, sharing a broad lobe-shaped base.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
229EDB61ACDB56A9A58065B36AD7244F.taxon	distribution	Distribution. Nghe An Province (Pu Mat NP) (Golovatch and Semenyuk 2021).	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
02E7C2BCC3ED5697A8FC4ED8B07F8982.taxon	description	Figs 9, 10, 11	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
02E7C2BCC3ED5697A8FC4ED8B07F8982.taxon	diagnosis	Diagnosis. The new species is similar to K. montanus Golovatch, 2009, but differs in the following characters: gonopod process a curved, serrated, acuminate leaf-shaped, as long as distally serrated, ribbon-shaped process b.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
02E7C2BCC3ED5697A8FC4ED8B07F8982.taxon	description	Description. Size: length 32.01 mm, width of midbody pro- and metazona 2.67 mm and 3.67 mm, respectively. Colouration (Fig. 9): almost uniformly brownish yellow after long preservation in ethanol. Head (Fig. 9 A – C): Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae claviform, moderately long (Fig. 9 A – C), extending behind body segment 3 when stretched laterally; antennomere 2 = 3 = 4> 5> 6> 7 = 1, antennomere 7 with four conical sensories. Collum: traces of setae hardly seen; lateral incisions absent; caudal corner of paraterga very broadly rounded, declined ventrad, produced behind rear tergal margin (Fig. 9 A, B). Body rings: In width, segment 4 <3 <head <5 <segment 2 <collum <6 – 17, thereafter body gently and gradually tapering. Tegument (Fig. 9 A – E) smooth and shining, prozonae finely shagreened; surface below paraterga finely rugulose. Tergal setae all broken, traces hardly seen. Axial line visible, especially on midbody tergites. Transverse sulcus usually distinct (Fig. 9 A), slightly incomplete on body rings 4 and 19, complete on metaterga 5 – 18, narrow, line – shaped, shallow, reaching bases of paraterga. Stricture between pro – and metazonae evident, broad and deep, striolated at bottom down to base of paraterga (Fig. 9 D). Pleurosternal carinae complete crests on body rings 2 – 7; thereafter, increasingly strongly reduced until body ring 17. Paraterga strongly developed (Fig. 9 A, D), lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded; lateral edge without incisions; posterior edge nearly straight. Calluses on paraterga narrow, delimited by a sulcus both dorsally and ventrally. Ozopores (Fig. 9 D, E) evident, lateral, lying in an ovoid groove at ~ 1 / 4 in front of posterior edge of metaterga. Telson: damaged. Sterna: densely setose, without modifications (Fig. 9 C, F). Legs: rather long and slender, midbody ones ~ 1.2 – 1.3 times as long as body height; prefemora without modifications, tarsal brushes present only on pregonopodal legs Gonopods (Figs 10, 11) a typical Kronopolites species; coxite (co) stout, subcylindrical, sparsely setose distoventrally. Prefemur (pref) densely setose, ~ 1 / 3 as long as femorite + postfemoral part. Femorite somewhat stout, evidently grooved mesally, demarcated from postfemoral region by a distinct oblique sulcus laterally. Lamina l present, producing into a small spine z distally; process a leaf-shaped, mesal margin serrated, clearly shorter than distally-serrated, ribbon-shaped process b; solenophore well developed, bipartite, longer than a short flagelliform solenomere which parly sheathed by solenophore.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
02E7C2BCC3ED5697A8FC4ED8B07F8982.taxon	etymology	Etymology. Serratus, an adjective epithet, is used to emphasise the serrated process a and b of the gonopod.	en	Nguyen, Anh D., Vu, Tam T. T., Phung, Hong Luong T., Tran, Duc-Luong, Le, Hung-Anh (2025): Mountainous millipedes in Vietnam. V. The millipede genus Kronopolites Attems, 1914 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species. ZooKeys 1249: 223-245, DOI: 10.3897/zookeys.1249.155280
