identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9123F87D00935C2BA66BD65439865886.text	9123F87D00935C2BA66BD65439865886.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillo officinalis Dumeril 1816	<div><p>Armadillo officinalis Duméril, 1816</p><p>Fig. 6 L</p><p>Armadillo officinalis Duméril, 1816: 117; Budde-Lund 1885: 16; Preudhomme de Borre 1886: 112; Dollfus 1896 a: 357; Dollfus 1896 b: 527; Arcangeli 1935 a: 2; Vandel 1946: 371.</p><p>Cubaris globator; Eaton 1882: 361.</p><p>Cubaris officinalis; Jackson 1926: 184.</p><p>Material examined.</p><p>Portugal – Arrábida • 28 ♂♂, 63 ♀♀; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001139, 001153, 001156, 001168 • over 1000 individuals (♂ and ♀); 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001232, 001239, 001241, 001244 • 84 ♂♂, 140 ♀♀, 2 juvs; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001305, 001309, 001312, 001318, 001321, 001331, 001334 • 1 ♀; 22.III.2023; same data as for preceding; MNHNC: MB 11: 001362 .</p><p>Distribution.</p><p>Widely distributed in the Mediterranean and Black Sea area.</p><p>Ecological notes.</p><p>This epigean species can also be found in MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was only found in Arrábida, during all seasons, and abundance increasing from spring to summer where it reaches its peak, then decreasing in fall (Fig. 7 L).</p></div>	https://treatment.plazi.org/id/9123F87D00935C2BA66BD65439865886	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
31F00CBFA90B5D8B908AB7D76710C753.text	31F00CBFA90B5D8B908AB7D76710C753.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenoscia minima (Dollfus 1892)	<div><p>Ctenoscia minima (Dollfus, 1892)</p><p>Fig. 6 B</p><p>Philoscia minima Dollfus, 1892: 187, fig. 3.</p><p>Philoscia (Benthana) minima Jackson, 1926: 196, pl. VIII figs 152–165.</p><p>Ctenoscia minima Vandel, 1946: 196, figs 42–49; Schmölzer 1965: 141, fig. 605; Schmölzer 1971: 20; Schmalfuss 2003: 87; Reboleira et al. 2015: 40; Cifuentes and Da Silva 2023: 36.</p><p>Ctenoscia dorsalis; Gregory et al. 2012: 17.</p><p>Material examined.</p><p>Portugal – Sicó • 19 ♀♀; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001130, 001133, 001144, 001149, 001165 • 1 ♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001256 . – Alcobertas • 1 ♂, 3 ♀♀; 39°25'46.5"N, 8°54'59.0"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001185, 001187, 001202 • 6 ♀♀; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001205, 001212, 001218 • 3 ♀♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001294, 001298 • 1 ♂, 2 ♀♀; 01.IV.2023; same data as for preceding; MNHNC: MB 11: 001357, 001360 . – Montejunto • 2 ♂♂, 13 ♀♀; 39°10'11.4"N, 9°04'14.8"W; 22.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001181, 001192 • 2 ♂♂, 3 ♀♀; 21.IX.2022; same data as for preceding; MNHNC: MB 11: 001223, 001231 • 2 ♂♂; 21.XII.2022; same data as for preceding; MNHNC: MB 11: 001340 • 2 ♀♀; 21.III.2023; same data as for preceding; MNHNC: MB 11: 001385 . – Arrábida • 17 ♀♀, 1 juv.; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001137, 001151, 001154, 001172, 001176 • 4 ♂♂, 10 ♀♀, 2 juvs; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001233, 001237, 001245 • 2 ♂♂, 11 ♀♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001306, 001314, 001324 .</p><p>Distribution.</p><p>Portugal, Spain, Italy, France (Corsica), Malta, Tunisia, and Morocco.</p><p>Ecological notes.</p><p>This epigean species can also be found in caves and MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in four locations, Sicó, Alcobertas, Montejunto and Arrábida. In Sicó it was captured during spring and summer, with a higher abundance in spring (Fig. 7 B). In Alcobertas it was captured during all seasons, with abundance increasing towards summer and then decreasing till winter (Fig. 7 B). In Montejunto this species was also captured during all seasons, with abundance decreasing steadily towards fall and then staying the same in winter (Fig. 7 B). In Arrábida it was captured during spring, summer, and fall, with a steady decrease in abundance from spring to fall (Fig. 7 B).</p></div>	https://treatment.plazi.org/id/31F00CBFA90B5D8B908AB7D76710C753	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
6561F11ACA8A5B5C9CE9923897D561CC.text	6561F11ACA8A5B5C9CE9923897D561CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eluma caelata (Miers 1877)	<div><p>Eluma caelata (Miers, 1877)</p><p>Fig. 6 H</p><p>Armadillidium caelatum Miers, 1877: 665, pl. 7 fig. 3.</p><p>Rhacodes inscriptus Eaton, 1882: 360 (not Rhacodes inscriptus C. L. Koch in Rosenhauer 1856: 422).</p><p>Eluma purpurascens Budde-Lund, 1885: 48, 294; Preudhomme de Borre 1886: 112; Dollfus 1889 a: 126; Dollfus 1889 b: 306; Dollfus 1896 a: 357; Norman 1899: 69; Verhoeff 1908: 370; Vandel 1946: 357, fig. 158.</p><p>Eluma caelatum Arcangeli, 1935 a: 3; Arcangeli 1948: 242; Schmölzer 1965: 303, fig. 1213; Schmölzer 1971: 58; Schmalfuss 2003: 107.</p><p>Eluma caelata Gregory et al., 2012: 21, tab. 2, 3; Reboleira et al. 2015: 47; Recuero et al. 2022: 1315, tab. 1; Cifuentes and Da Silva 2023: 29, figs 1 A, 2 A-C, 4 G.</p><p>Material examined.</p><p>Portugal – Serra de Estrela • 1 ♀; 40°17'22.9"N, 7°35'19.7"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001121 • 1 ♂, 1 ♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001250 • 5 ♂♂, 7 ♀♀; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001268, 001270, 001271, 001272 • 4 ♂♂, 5 ♀♀; 19.XII.2022; same data as for preceding; MZUF 10034 • 1 ♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001347 . – Sicó • 3 ♀♀; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001126, 001148, 001163 • 2 ♂♂, 3 ♀♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001251, 001254, 001265, 001267 • 1 ♀; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001280 • 3 ♀♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001369, 001380 . – Alcobertas • 34 ♂♂, 27 ♀♀; 39°25'46.5"N, 8°54'59.0"W; 1.IV.2023; leg. R. P. EusébioEusébio, R. P. leg; MSS pitfall trap; MNHNC: MB 11: 001349, 001359 . – Montejunto • 14 ♂♂, 9 ♀♀; 39°10'11.4"N, 9°04'14.8"W; 23.VII.2022;; MSS pitfall trap; MNHNC: MB 11: 001159, 001182, 001183, 001193 • 1 ♂, 11 ♀♀; 21.IX.2022; same data as for preceding; MNHNC: MB 11: 001221, 001224, 001228 • 11 ♂♂, 4 ♀♀; 21.XII.2022; same data as for preceding; MNHNC: MB 11: 001327, 001336, 001342, 001345 • 38 ♂♂, 26 ♀♀, 1 juv.; 21.III.2023; same data as for preceding; MNHNC: MB 11: 001388, 001391, 001395, 001398, 001401 . – Arrábida • 8 ♂♂, 9 ♀♀; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001157 • 1 ♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001311 • 1 ♂, 2 ♀♀; 22.III.2023; same data as for preceding; MNHNC: MB 11: 001367 .</p><p>Distribution.</p><p>United Kingdom, France, Iberian Peninsula, The Netherlands, Belgium, Morocco, and Western Sahara. It was introduced to Australia (Tasmania) and French Guiana.</p><p>Ecological notes.</p><p>This epigean species can also be found in caves and MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in all locations. In Serra da Estrela, Sicó and Montejunto it was captured during all seasons. In Serra da Estrela abundance increases towards fall then decreases in winter (Fig. 7 H). In Sicó abundance increases toward summer, decreases in fall and increases again in winter (Fig. 7 H). In Montejunto abundance decreases in summer then increases steadily towards winter (Fig. 7 H). In Alcobertas it was captured only in winter (Fig. 7 H). In Arrábida it was captured during spring, fall and winter, with abundance decreasing from its peak in spring towards fall, then increasing slightly in winter (Fig. 7 H).</p></div>	https://treatment.plazi.org/id/6561F11ACA8A5B5C9CE9923897D561CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
06ADCA74A36F5F7198D3AC53E41699DC.text	06ADCA74A36F5F7198D3AC53E41699DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eluma cristata Eusébio & Taiti & Reboleira 2025	<div><p>Eluma cristata sp. nov.</p><p>Figs 2, 3, 4, 5, 6 K</p><p>Eluma tuberculata Reboleira et al., 2015: 49 (partim: specimens from MSS near Gruta de Alcobertas, Estremenho Karst Massif).</p><p>Type material.</p><p>Holotype. Portugal • ♂; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.9163885&amp;materialsCitation.latitude=39.429585" title="Search Plazi for locations around (long -8.9163885/lat 39.429585)">Alcobertas</a>; 39°25'46.5"N, 8°54'59.0"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001198 . Paratypes: Portugal • 137 ♂♂, 138 ♀♀; same data as holotype; MNHNC: MB 11: 001186, 001189, 001195, 001198, 001200 • 115 ♂♂, 121 ♀♀; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001203, 001207, 001209, 001213, 001216 • 5 ♂♂, 10 ♀♀; same data as for preceding; MZUF 10038 • 280 ♂♂, 240 ♀♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001291, 001295, 001296, 001301, 001302 • 10 ♂♂, 10 ♀♀; same data as for preceding; MZUF 10039 • 68 ♂♂, 103 ♀♀; 1.IV.2023; same data as for preceding; MNHNC: MB 11: 001348, 001351, 001354, 001356 .</p><p>Description.</p><p>Maximum length: ♂ 7.5 mm; ♀ 9.5 mm. Body strongly convex, able to roll up into perfect ball, euspheric type. Colour light brown, as in E. matae and E. tuberculata . Dorsal surface with ornamentation consisting of protruding crests on posterior margins of cephalon, pereonites 1–7 tergites and median part of pleonites 1–5, each crest ending in lobes of irregular shape and number; telson with two small paramedian tubercles (Figs 2 A – E, 3 A, G, H, 6 K). Dorsum with scattered pointed scale-setae (Fig. 3 B) and one line of noduli laterales per side located near top of crests on pereonites 1–7 far from lateral margins (Fig. 3 G). Cephalon (Figs 2 A – D, 3 A, 6 K) with triangular scutellum distinctly separated from and not bent over vertex; distinct frontal line and no postscutellar line; oblique antennary lobes, quadrangular and directed frontwards. Eye consisting of single large ocellus as in Fig. 2 A, C. Pereonite 1 (Figs 2 A, C, 3 C, D, 6 K) with deep dorsal groove on lateral margin, schisma on posterior corners with rounded inner lobe distinctly shorter than outer lobe; posterior margin slightly concave at sides. Pereonites 2 and 3 (Fig. 3 E, F) epimera quadrangular with small and rounded lobe on ventral side. Pereonites 4–7 with quadrangular epimera and straight posterior margins. Pleonites 3–5 (Fig. 3 G) with rectangular epimera, slightly divergent. Telson (Fig. 3 G, H) triangular, wider than long, with slightly concave sides and rounded apex. Antennula (Fig. 3 I) of three articles, second article much shorter than first and third, third article with tuft of superimposed aesthetascs subapically. Antenna (Fig. 4 A) with flagellum slightly shorter than fifth article of peduncle, second flagellar article more than twice as long as first, bearing two aesthetascs on middle part. Mandibles (Fig. 4 B, C) with dichotomized molar penicil and three or four free penicils; right mandible with one penicil and left mandible with two penicils on hairy lobe. Maxillula (Fig. 4 D) outer lobe with 4 + 6 (5 cleft) teeth; inner lobe with two thick subapical penicils and small triangular distal point. Maxilla (Fig. 4 E) apically setose with outer lobe rounded, much wider than inner lobe. Maxilliped (Fig. 4 F) with quadrangular endite bearing two triangular spines on distal margin and large subapical seta; basal article of palp with two large setae. Uropod (Fig. 4 G) flattened; exopod about twice as wide as long with slightly concave distal margin; endopod slightly protruding backwards compared with exopod.</p><p>Male. Pereopod 1–3 (Fig. 5 A) with line of pointed setae on sternal margins of carpus and, to lesser extent, merus. Pereopod 7 (Fig. 5 B) ischium with sternal margin distinctly concave, distal margin with rounded lobe and four stout setae; merus with sternal margin slightly convex; merus and carpus with line of long setae on sternal margin. Genital papilla (Fig. 5 C) fusiform. Pleopod 1 (Fig. 5 C) exopod with large distal lobe bent outwards and apically rounded; endopod with distal part quadrangular and short apical lobe directed outwards. Pleopod 2 (Fig. 5 D) exopod triangular with a line of about five short setae on outer margin; endopod slightly longer than exopod. Pleopods 3–5 exopods as in Fig. 5 E – G.</p><p>Etymology.</p><p>From Latin cristatus = crested. The name refers to the high crests on the posterior margins of the body segments.</p><p>Remarks.</p><p>At present, the genus Eluma comprises only five species, i. e., the four species included here and E. praticola Taiti &amp; Rossano, 2015 from northern Morocco (see Schmalfuss 2003; Taiti and Rossano 2015; Cifuentes and Da Silva 2023). Eluma cristata sp. nov. is readily distinct from all the other species by the protruding dorsal crests on the posterior margins of the cephalon, pereonites and, to a lesser extent, pleonites. Moreover, it differs from E. caelata in lacking the basal triangular lobe on the male pereopod 7 merus (see fig. 32 F, G in Taiti and Rossano 2015); from this species and E. praticola also in the male pleopod 1 endopod with thickset distal part, not bent outwards; from E. matae also in the much shorter dorsal scale-setae of different shape (see fig. 4 A – F in Cifuentes and Da Silva 2023).</p><p>Ecological notes.</p><p>This species was only found in this MSS habitat.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was only found in Alcobertas, during all seasons, and with a decrease in abundance from spring to summer, highest abundance during fall and a new decrease in winter (Fig. 7 K).</p></div>	https://treatment.plazi.org/id/06ADCA74A36F5F7198D3AC53E41699DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
5857BBC77EF45E4BA05D019B43C83A8C.text	5857BBC77EF45E4BA05D019B43C83A8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eluma matae Cifuentes and Da Silva 2023	<div><p>Eluma matae Cifuentes and Da Silva, 2023</p><p>Fig. 6 J</p><p>Eluma matae Cifuentes &amp; Da Silva, 2023: 33, figs 1 B, 3, 4 A-F, I, K-Q.</p><p>Material examined.</p><p>Portugal – Montejunto • 6 ♂♂, 10 ♀♀; 39°10'11.4"N, 9°04'14.8"W; 22.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001177, 001184, 001194) • 10 ♀♀, 6 juvs; 21.IX.2022; same data as for preceding; MNHNC: MB 11: 001225, 001227, 001229 • 9 ♂♂, 13 ♀♀; 21.XII.2022; same data as for preceding; MNHNC: MB 11: 001328, 001335, 001337, 001344 • 5 ♂♂, 5 ♀♀; 21.XII.2022; same data as for preceding; MZUF 10037 • 30 ♂♂, 44 ♀♀, 1 juv.; 21.III.2023; same data as for preceding; MNHNC: MB 11: 001390, 001392, 001396, 001399, 001402 .</p><p>Distribution.</p><p>Endemic to Portugal.</p><p>Ecological notes.</p><p>This epigean species can also be found in MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was only found in Montejunto, during all seasons, and with an increase in abundance from spring to winter (Fig. 7 J).</p></div>	https://treatment.plazi.org/id/5857BBC77EF45E4BA05D019B43C83A8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
6DE9514DFFA75A32BF914D3F2AF3D8D8.text	6DE9514DFFA75A32BF914D3F2AF3D8D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eluma tuberculata Cruz 1991	<div><p>Eluma tuberculata Cruz, 1991</p><p>Fig. 6 I</p><p>Eluma tuberculata Cruz, 1991: 73, fig. 1 a – f; Schmalfuss 2003: 107; Reboleira et al. 2015: 49, figs 30, 31 (nec specimens from MSS near Gruta de Alcobertas, Estremenho Massif); Cifuentes and Da Silva 2023: 33, figs 1 C, 2 D-F, 4 H.</p><p>Material examined.</p><p>Portugal – Sicó • 68 ♂♂, 114 ♀♀; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001124, 001125, 001132, 001146, 001162 • 5 ♂♂, 5 ♀♀; 20.VII.2022; same data as for preceding; MZUF 10035 • 39 ♂♂, 93 ♀♀, 13 juvs; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001252, 001253, 001260, 001261, 001266 • 10 ♂♂, 10 ♀♀; 19.IX.2022; same data as for preceding; MZUF 10036 • 27 ♂♂, 78 ♀♀, 14 juvs; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001276, 001279, 001281, 001286, 001289 • 23 ♂♂, 35 ♀♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001370, 001375, 001377, 001378, 001384 .</p><p>Distribution.</p><p>Endemic to Portugal.</p><p>Ecological notes.</p><p>This species can be found in caves and MSS habitats.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was only found in Sicó, during all seasons, and with a decrease in abundance from spring to winter (Fig. 7 I).</p></div>	https://treatment.plazi.org/id/6DE9514DFFA75A32BF914D3F2AF3D8D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
A246634896EE51ABAFE9FF6FA6EF0D6E.text	A246634896EE51ABAFE9FF6FA6EF0D6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porcellio dilatatus Brandt 1831	<div><p>Porcellio dilatatus Brandt, 1831</p><p>Fig. 6 E</p><p>Porcellio dilatatus Brandt in Brandt and Ratzeburg 1831: 78, Pl. 12, fig. 6 c, d; Vandel 1946: 316, fig. 135; Schmölzer 1971: 44.</p><p>Porcellio (Porcellio) dilatatus Arcangeli, 1935 a: 12 .</p><p>Porcellio dilatatus dilatatus; Reboleira et al. 2015: 46.</p><p>Material examined.</p><p>Portugal – Sicó • 1 ♀; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001122 • 1 ♂; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001274 • 1 ♂, 3 ♀♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001371, 001382 . – Montejunto • 1 ♂; 39°10'11.4"N, 9°04'14.8"W; 21.III.2023; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001389 . – Arrábida • 22 ♂♂, 41 ♀♀, 182 juvs; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001135, 001138, 001140, 001150, 001152, 001158, 001169, 001170, 001173, 001174 • 2 ♂♂, 12 ♀♀, 20 juvs; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001235, 001238, 001243, 001247 • 28 ♂♂, 23 ♀♀, 32 juvs; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001304, 001308, 001310, 001316, 001317, 001319, 001322, 001326, 001330, 001333 • 16 ♂♂, 13 ♀♀; 22.III.2023; same data as for preceding; MNHNC: MB 11: 001361, 001364, 001365, 001366 .</p><p>Distribution.</p><p>Common across Europe and introduced into many parts of the world.</p><p>Ecological notes.</p><p>This species is mainly found in caves and MSS habitats but does not show any adaptation to subterranean habitats.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in three locations, Sicó, Montejunto and Arrábida. In Sicó and Montejunto it was found in very low numbers (Fig. 7 E). In Arrábida it was captured during all seasons, with abundance decreasing from spring to summer, then increasing slightly in fall and decreasing again in winter (Fig. 7 E).</p></div>	https://treatment.plazi.org/id/A246634896EE51ABAFE9FF6FA6EF0D6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
79A2CEE4792E501BB0BEA217F19148DF.text	79A2CEE4792E501BB0BEA217F19148DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porcellio dispar Verhoeff 1901	<div><p>Porcellio dispar Verhoeff, 1901</p><p>Fig. 6 F</p><p>Porcellio dispar Verhoeff, 1901: 407; Vandel 1946: 289, figs 110–111; Schmölzer 1965: 228; Schmölzer 1971: 36; Schmalfuss, 2003: 222; Cifuentes and Da Silva 2023: 36.</p><p>Porcellio (Euporcellio) dispar Verhoeff, 1907: 269.</p><p>Porcellionides dimorphus Jackson, 1926: 186, pl. I figs 1–22.</p><p>Porcellio (Porcellio) dispar Arcangeli, 1935 a: 10, figs 3–7.</p><p>Material examined.</p><p>Portugal – Sicó • 6 ♀♀; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001131, 001145, 001164 • 1 ♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001264 • 2 ♀♀; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001277 • 3 ♀♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001373 . – Montejunto • 2 ♂♂, 4 ♀♀; 39°10'11.4"N, 9°04'14.8"W; 23.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001160, 001179 • 1 ♀; 21.XII.2022; same data as for preceding; MNHNC: MB 11: 001343 • 3 ♂♂, 23 ♀♀; 21.III.2023; same data as for preceding; MNHNC: MB 11: 001386, 001394 .</p><p>Distribution.</p><p>Portugal, NW Spain, and France.</p><p>Ecological notes.</p><p>This epigean species can also be found in caves and MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in two locations, Sicó and Montejunto. In Sicó it was captured in all seasons, with abundance decreasing from spring to summer, then increasing slightly until winter (Fig. 7 F). In Montejunto it was captured during spring, fall and winter, with abundance decreasing from spring to fall and increasing to its highest in winter (Fig. 7 F).</p></div>	https://treatment.plazi.org/id/79A2CEE4792E501BB0BEA217F19148DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
F1535111AE995F6491CAC6FBC4D0B7F6.text	F1535111AE995F6491CAC6FBC4D0B7F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porcellio novus Arcangeli 1935	<div><p>Porcellio novus Arcangeli, 1935</p><p>Fig. 6 G</p><p>Porcellio (Porcellio) novus Arcangeli, 1935 b: 40, figs 1–6.</p><p>Porcellio novus Vandel, 1946: 322; Schmölzer 1955: 314; Schmölzer 1965: 230, fig. 835; Schmölzer 1971: 38; Schmalfuss, 2003: 231.</p><p>Material examined.</p><p>Portugal – Alcobertas • 2 ♂♂, 3 ♀♀; 39°25'46.5"N, 8°54'59.0"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001196 • 6 ♂♂, 10 ♀♀, 1 juv.; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001208, 001210, 001214, 001219 • 1 ♂, 3 ♀♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001290, 001292 • 2 ♀♀; 1.IV.2023; same data as for preceding; MNHNC: MB 11: 001355 .</p><p>Distribution.</p><p>Portugal and western Spain.</p><p>Ecological notes.</p><p>This epigean species can also be found in MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in one location, Alcobertas, during all seasons, with abundance increasing from spring to summer, then decreasing steadily towards winter (Fig. 7 G).</p></div>	https://treatment.plazi.org/id/F1535111AE995F6491CAC6FBC4D0B7F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
43BCC4584F9857AD9854CB3426332654.text	43BCC4584F9857AD9854CB3426332654.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porcellio scaber Latreille 1804	<div><p>Porcellio scaber Latreille, 1804</p><p>Fig. 6 D</p><p>Porcellio scaber Latreille, 1804: 45; Dollfus 1897: 206; Vandel 1946: 297; Gregory et al. 2012: 20, fig. 6.</p><p>Porcellio (Porcellio) scaber Arcangeli, 1935 a: 9 .</p><p>Porcellio (Euporcellio) scaber var. scabrior Verhoeff, 1907: 266.</p><p>Porcellio scaber scaber Schmölzer, 1971: 39.</p><p>Material examined.</p><p>Portugal – Serra de Estrela • 1 ♂, 1 ♀; 40°17'22.9"N, 7°35'19.7"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001167 • 1 ♂, 9 ♀♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001240, 001248, 001249 • 1 ♂, 1 ♀; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001269, 001273 .</p><p>Distribution.</p><p>Widespread in Europe and introduced into many parts of the world.</p><p>Ecological notes.</p><p>This epigean species can also be found in MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in one location, Serra da Estrela, during spring, summer, and fall, and with abundance peaking in summer (Fig. 7 D).</p></div>	https://treatment.plazi.org/id/43BCC4584F9857AD9854CB3426332654	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
3BFD377A885757818359D7ED4BBDEACA.text	3BFD377A885757818359D7ED4BBDEACA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porcellionides cingendus (Kinahan 1857)	<div><p>Porcellionides cingendus (Kinahan, 1857)</p><p>Fig. 6 C</p><p>Porcellio cingendus Kinahan, 1857: 279, figs. 1, 2, 4, 6, 8, 9.</p><p>Porcellio (Porcellionides) lusitanorum Arcangeli, 1935 a: 21, figs 13–18.</p><p>Metoponorthus (Lusitoniscus) cingendus Vandel, 1946: 274, figs 91–97; Schmölzer 1965: 195, figs 698–699; Schmölzer 1971: 33.</p><p>Porcellionides cingendus Schmalfuss, 2003: 240; Gregory et al. 2012: 18; Reboleira et al. 2015; Cifuentes and Da Silva 2024: 4, figs 1, 2 A, 3 A, 4 A – D, 5, tab. 1.</p><p>Material examined.</p><p>Portugal – Sicó • 54 ♂♂, 58 ♀♀, 2 juvs; 39°58'51.8"N, 8°33'13.1"W; 20.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001123, 001127, 001128, 001129, 001134, 001143, 001147, 001166 • 18 ♂♂, 38 ♀♀; 19.IX.2022; same data as for preceding; MNHNC: MB 11: 001255, 001257, 001258, 001259, 001262, 001263 • 32 ♂♂, 31 ♀♀; 19.XII.2022; same data as for preceding; MNHNC: MB 11: 001275, 001278, 001282, 001284, 001288 • 30 ♂♂, 6 ♀♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001372, 001374, 001376, 001381, 001383 . – Alcobertas • 6 ♂♂, 21 ♀♀; 39°25'46.5"N, 8°54'59.0"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001188, 001197, 001199, 001201 • 13 ♂♂, 58 ♀♀; 20.IX.2022; same data as for precedings; MNHNC: MB 11: 001204, 001211, 001215, 001217 • 34 ♂♂, 46 ♀♀; 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001293, 001297, 001299, 001303 • 56 ♂♂, 23 ♀♀; 01.IV.2023; same data as for preceding; MNHNC: MB 11: 001350, 001352, 001353, 001358 . – Montejunto • 32 ♂♂, 44 ♀♀; 39°10'11.4"N, 9°04'14.8"W; 23.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001161, 001178, 001180, 001190 • 3 ♂♂, 33 ♀♀; 21.IX.2022; same data as for preceding; MNHNC: MB 11: 001220, 001222, 001226, 001230 • 55 ♂♂, 57 ♀♀, 2 juvs; 21.XII.2022; same data as for preceding; MNHNC: MB 11: 001329, 001338, 001339, 001346 • over 500 specimens (♂ and ♀); 21.III.2023; same data as for preceding; MNHNC: MB 11: 001387, 001393, 001397, 001400, 001403 . – Arrábida • 11 ♂♂, 17 ♀♀, 3 juvs; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001136, 001142, 001155, 001171, 001175 • 18 ♂♂, 64 ♀♀, 46 juvs; 20.IX.2022; same data as for preceding; MNHNC: MB 11: 001234, 001236, 001242, 001246 • over 200 specimens (♂ and ♀); 20.XII.2022; same data as for preceding; MNHNC: MB 11: 001307, 001315, 001320, 001325, 001332 • 2 ♂♂, 2 ♀♀, 1 juv.; 22.III.2023; same data as for preceding; MNHNC: MB 11: 001363, 001368 .</p><p>Distribution.</p><p>United Kingdom, France and Iberian Peninsula.</p><p>Ecological notes.</p><p>This epigean species can also occur in caves and MSS.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in four locations, Sicó, Alcobertas, Montejunto and Arrábida. Specimens were found during all seasons in the four locations. In Sicó there was a steady decrease in abundance from spring to winter, in Alcobertas there was an increase from spring to winter, in Montejunto abundance decreased from spring to summer and then increased until winter (Fig. 7 C). In Arrábida abundance increased steadily until fall and then decreased to exceptionally low numbers in winter (Fig. 7 C).</p></div>	https://treatment.plazi.org/id/3BFD377A885757818359D7ED4BBDEACA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
2723060D34EE5863A4E05439DC7CF412.text	2723060D34EE5863A4E05439DC7CF412.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichoniscoides machadoi Vandel 1946	<div><p>Trichoniscoides machadoi Vandel, 1946</p><p>Fig. 6 A</p><p>Trichoniscoides machadoi Vandel, 1946: 168, figs 8–12; Vandel 1947: 270; Vandel 1948: 129, fig. 14; Vandel 1952: 262, figs 2, 9; Schmölzer 1965: 37, figs 79–82; Schmölzer 1971: 4, 75, 139, map 4; Schmalfuss 2003: 303; Reboleira et al. 2015: 6, fig. 2; Cifuentes 2019: 48; Cifuentes and Da Silva 2023: 7.</p><p>Material examined.</p><p>Portugal – Sicó • 2 ♂♂, 3 ♀♀; 39°58'51.8"N, 8°33'13.1"W; 19.XII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001283, 001285, 001287 • 1 ♀; 20.III.2023; same data as for preceding; MNHNC: MB 11: 001379 . – Arrábida • 1 ♂, 2 ♀♀; 38°28'59.2"N, 9°00'09.9"W; 21.VII.2022; leg. R. P. Eusébio; MSS pitfall trap; MNHNC: MB 11: 001141 • 3 ♂♂; 22.III.2023; same data as for preceding; MNHNC: MB 11: 001313, 001323 .</p><p>Distribution.</p><p>Endemic to Portugal. This species is widely distributed throughout the country.</p><p>Ecological notes.</p><p>This species can be found in caves, MSS, and endogean habitats.</p><p>Seasonal patterns in Portuguese MSS habitats.</p><p>This species was found in two locations, Sicó and Arrábida. In Sicó it was captured during fall and winter, with a higher abundance in fall (Fig. 7 A). In Arrábida it was captured during spring and fall in equal abundances (Fig. 7 A).</p></div>	https://treatment.plazi.org/id/2723060D34EE5863A4E05439DC7CF412	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Eusébio, Rita P.;Taiti, Stefano;Reboleira, Ana Sofia P. S.	Eusébio, Rita P., Taiti, Stefano, Reboleira, Ana Sofia P. S. (2025): Terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Mesovoid Shallow Substratum habitats of Portugal. Subterranean Biology 53: 129-153, DOI: 10.3897/subtbiol.53.163632
