taxonID	type	description	language	source
CE7A878EFFE52910FF0DE564FC0D256A.taxon	description	Figs 1 - 19	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE52910FF0DE564FC0D256A.taxon	materials_examined	Material examined. Holotype: male, Southern Ecuador, 3 ° 58 ' S 79 ° 50 ' W, Estación Científica San Francisco, 2000 m a. s. l., upper organic soil layer in mostly undisturbed rain forest, 01.04.2008 (ZISP). – Paratypes: five males and one female in SZMN, and five males in PC, same locality and date as holotype.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE52910FF0DE564FC0D256A.taxon	diagnosis	Diagnosis. Body size 647 - 680 × 332 - 356. Surface of notogaster and anogenital region smooth or foveolate. Interbothridial region with two pairs of muscle sigilla. Interlamellar setae longer than rostral and lamellar setae; all setiform, ciliate unilaterally. Sensilli setiform, with four to five long branches. Notogastral setae long, ciliate unilaterally. Epimeral region with muscle sigilla. Anogenital setae smooth (except ciliate adanal setae). Lyrifissures iad located in inverse apoanal or paraanal position.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE52910FF0DE564FC0D256A.taxon	description	Description Measurements: Body length 647 (holotype), 647 - 680 (mean 660; 11 paratypes); notogaster width 348 (holotype), 332 - 356 (mean 346; 11 paratypes). Integument (Figs 1 - 3): Body colour brown to yellow-brownish. Dorsal surface of prodorsum smooth; lateral sides with small microgranulate parts between bothridia and acetabula I, II. Interbothridial region with two pairs of distinct muscle sigilla. Surface of notogaster and anogenital region smooth or rarely foveolate (diameter foveolae up to 4 µm). Distance between foveolae considerable longer than diameter of foveolae. Epimeral region with numerous muscle sigilla. Prodorsum (Figs 1, 3 - 8): Rostrum rounded. Lamellar lines thin, not visible absolutely in many specimens. Triangular tubercle located posteriorly to each bothridium and interlamellar seta. Rostral (ro, 45 - 53), lamellar (le, 53 - 61), interlamellar (in, 90 - 102) and exobothridial (ex, 24 - 36) setae setiform, with short cilia unilaterally. Sensilli (ss, 127 - 143) setiform, having four (rarely five) long branches in medio-distal part. Notogaster (Fig. 1): Anterior border convex. A pair of setae c is absent (only alveoli present). Twelve other pairs of notogastral setae long (123 - 151), setiform, with short cilia unilaterally. Setae in anterior half of body slightly shorter than in posterior half. Lyrifissures ia, im, ip, ih and ips distinct, located typically for genus. Opisthonotal gland openings (gla) located posteriorly to setae h 3. Gnathosoma (Figs 9 - 12): Subcapitulum longer than wide (155 - 168 × 98 - 110). Subcapitular setae (a, m, h) setiform, smooth, similar in length (20 - 28). Two pairs of adoral setae (or 1, or 2, 4 - 6) present, thorn-like. Palps (82 - 94) with setation 0 - 2 - 1 - 3 - 9 (+ ω). Setae of femora with long, thickened cilia; setae of genua and tibia with short cilia unilaterally; setae of tarsi smooth. Solenidion long, thickened, blunt-ended, pressed to the palptarsus surface, attached with distal seta u " in distal half. Chelicerae (159 - 168) chelate-dentate. Cheliceral setae setiform, barbed; cha (36 - 45) longer than chb (24 - 28). Trägårdh’ s organ (Tg) distinct. Epimeral and lateral podosomal regions (Figs 2, 3, 13, 14): Epimeral border IV convex posteriorly. Epimeral setae setiform; 1 c, 3 c and 4 c with short cilia unilaterally, others smooth. Setae 3 c, 4 a and 4 c (53 - 57) longer than 1 b, 1 c, 3 b (32 - 41) and 1 a, 2 a, 3 a, 4 b (16 - 20). Pedotecta I (Pd I) convex. Discidia (dis) triangular, blunt-ended. ro Anogenital region (Figs 2, 15, 16): Five pairs of genital (g 1 - g 5, 16 - 20), one pair of aggenital (ag, 16 - 20) and two pairs of anal (an 1, 36 - 49; an 2, 28 - 32) setiform, smooth. Three pairs of adanal setae (ad 1, 98 - 118; ad 2, 77 - 90; ad 3, 49 - 57) setiform, with short cilia unilaterally. Lyrifissures iad located in inverse apoanal (paraanal in two specimens; left lyrifissure inverse apoanal, right lirifissure paraanal in one specimen) position. Legs (Figs 17 - 19): Generally, similar to Gittella ecuadoriensis Ermilov & Kalúz, 2012. Formulae of leg setation and solenidia: I (1 - 5 - 2 - 4 - 20) [1 - 2 - 2], II (1 - 5 - 2 - 4 - 16) [1 - 1 - 2], III (2 - 3 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 - 1 - 0]; homology of setae and solenidia indicated in Table 1. Setae p on tarsi I setiform; on tarsi II-IV thorn-like. Setae tc' on leg II, tc " on leg III, a " on leg IV with thickened cilia bilaterally. Setae a', s, po' on leg II, a ", s, po " on leg III, po " on leg IV with thickened cilia unilaterally. Famulus (e) setiform, straight or bent, blunt-ended. Solenidia ω 1, ω 2 on tarsi I, II, Φ 2 on tibiae I, Φ on tibiae II, σ on tibiae III thickened, blunt-ended. Other solenidia thinner and longer, weakly blunt distally.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE52910FF0DE564FC0D256A.taxon	etymology	Etymology. The specific name oariabilis refers to the variability of some morphological characters (body surface, orientation of lyrifissures iad).	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE52910FF0DE564FC0D256A.taxon	discussion	Remarks. In having the combination of morphological characters (setiform, branched sensilli; long and ciliate notogastral setae; length of adanal setae; presence of two pairs of interbothridial muscle sigilla), Gittella oariabilis spec. nov. is most similar to Gittella maxima (Balogh & Mahunka, 1981) (see Balogh & Mahunka 1981), however, it differs from the latter by the morphology of sensilli (with long four to five branches in the new species versus with four long and four short branches in G. maxima), presence of muscle sigilla in epimeral region (versus absent in G. maxima), morphology of notogastral setae (ciliate unilaterally in the new species versus ciliate not unilaterally in G. maxima).	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	description	Figs 20 - 34	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	materials_examined	Material examined. Holotype: male, Southern Ecuador, 3 ° 58 ' S 79 ° 50 ' W, Estación Científica San Francisco, 2000 m a. s. l., upper organic soil layer in mostly undisturbed rain forest, 01.04.2008 (ZISP). – Paratypes: one male and two females in SZMN, and three males in PC, same locality and date as holotype.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	diagnosis	Diagnosis. Body size 311 - 328 × 147 - 159. Body surface smooth. Lamellar lines distinct. Interbothridial region with two pairs of muscle sigilla. Lamellar setae longer than rostral and interlamellar setae. Sensilli with dilated head, having six long branches. Notogastral setae of medium size, ciliate. Epimeral region with muscle sigilla. Anogenital setae smooth (except ciliate adanal setae). Lyrifissures iad located in inverse apoanal position.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	description	Description Measurements: Body length 311 (holotype), 311 - 328 (mean 317; six paratypes); notogaster width 147 (holotype), 147 - 159 (mean 155; six paratypes). Integument (Figs 20 - 22): Body colour yellow-brownish. Body surface smooth; lateral sides with small microgranulate parts between bothridia and acetabula I, II. Interbothridial region with two pairs of distinct muscle sigilla. Epimeral region with numerous muscle sigilla. Prodorsum (Figs 20, 22 - 27): Rostrum rounded. Lamellar lines well developed. Triangular tubercle located posteriorly to each bothridium and interlamellar seta. Rostral (16 - 20), lamellar (24 - 32), interlamellar (16 - 20) and exobothridial (8 - 12) setae setiform, with very short cilia unilaterally. Sensilli (61 - 69) with dilated head, having six long branches. Notogaster (Figs 20, 28): Anterior border convex. A pair of setae c is absent (only alveoli present). Twelve other pairs of notogastral setae long (36 - 45), setiform, with short cilia unilaterally. Lyrifissures ia, im, ip, ih and ips distinct, located typically for genus. Opisthonotal gland openings located posteriorly to setae h 3. Gnathosoma: Subcapitulum longer than wide (82 - 90 × 49 - 53). Subcapitular setae (a, m, h) setiform, smooth, similar in length (16 - 20). Two pairs of adoral setae present, thorn-like (4). Palps (49 - 53) with setation 0 - 2 - 1 - 3 - 9 (+ ω). Setae of femora with long, thickened cilia; setae of genua and tibia with short cilia unilaterally; setae of tarsi smooth. Solenidion long, thickened, blunt-ended, pressed to the palptarsus surface, attached with distal seta u " in distal half. Chelicerae (82 - 90) chelate-dentate. Cheliceral setae setiform, barbed; cha (24 - 28) longer than chb (16). Trägårdh’ s organ distinct. Epimeral and lateral podosomal regions (Figs 21, 22, 29 - 31): Epimeral border IV convex posteriorly. Epimeral setae setiform; 1 c, 3 c and 4 c with short cilia unilaterally, others smooth. Setae 1 b, 1 c, 3 b, 3 c, 3 c, 4 a and 4 c (16 - 20) longer than others (6 - 12). Pedotecta I convex. Discidia triangular, blunt-ended. Anogenital region (Figs 21, 32 - 34): Five pairs of genital (4 - 6), one pair of aggenital (4 - 6) and two pairs of anal (an 1, 8 - 12; an 2, 4 - 8) setiform, smooth. Three pairs of adanal setae (ad 1, ad 2, 24 - 32; ad 3, 12 - 16) setiform, with short cilia unilaterally. Lyrifissures iad located in inverse apoanal position. Legs: Similar to Gittella oariabilis spec. nov. Homology of setae and solenidia indicated in Table 1.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	etymology	Etymology. The specific name minor refers to the small body size.	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
CE7A878EFFE02915FF00E21FFBDC2527.taxon	discussion	Remarks. In having the combination of morphological characters (smooth body surface; sensilli with dilated head, having branches; ciliate notogastral setae of medium length), Gittella minor spec. nov. is most similar to Gittella insularis Mahunka, 1998 (see Mahunka 1998), however, it differs from the latter by the smaller body size (311 - 328 × 147 - 159 in the new species versus 576 - 642 × 255 - 297 in G. insularis), presence of two pairs of interbothridial muscle sigilla (versus three pairs in G. insularis), ciliate rostral and interlamellar setae (versus smooth in G. insularis).	en	Ermilov, Sergey G., Sandmann, Dorothee, Marian, Franca, Maraun, Mark (2013): Two new oribatid mite species of the genus Gittella from Ecuador. Spixiana 36 (1): 1-8, DOI: 10.5281/zenodo.16851109
