taxonID	type	description	language	source
D70F637FFFCB6D4A1DE2506206AF23D7.taxon	description	(Fig. 12 B, D, G, H)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4A1DE2506206AF23D7.taxon	materials_examined	Material examined. HOLOTYPE OF MASARIS SMIRNOVI: ♂, labeled “ [golden disc] // Dist. Perovsk [Kyzylorda] / 29 V 923. / S. Tarbinskij // Тип [Type] / Masaris / smirnovi Kost. / = M. carli Schult / Синон. устан. / Ю. Костылев [Synonymy established by G. Kostylev] / 23 VI 41 [G. Kostylev’s signature] // Masaris / smirnovi / n. sp. / Typus // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotype ♂ / Masaris smirnovi / Kostylev, 1925 [red label] ” [ZMMU]. HOLOTYPE OF MASARIS ELEGANS: ♂, labeled “ Turkmenistan / Farab 29.4. / 1991 Halada // ♂ // Masaris / elegans nov. spec. ♂ / J. Gusenleitner, det. 2002 / Holotypus // Holotypus [red label] ” [OLML: ZOBODAT 2025 a]. PARATYPE OF MASARIS ELEGANS: KAZAKHSTAN. Almaty Province: Charyn Valley, W Chundzha, 43 ° 37 ʹN, 79 ° 21 ʹE, 650 m, 29 – 31. V. 2001, 1 ♀, leg. M. Hauser [OLML: ZOBODAT 2025 b]. ADDITIONAL MATERIAL: UZBEKISTAN: Bukhara Region, Khargush [Xargoʻsh], 27. V. 1928, 1 ♀, leg. A. Gerasimov [ZISP]. KAZAKHSTAN. Aktobe Province: Turgay Oblast, Aja, 17. VIII. 1899, 1 ♂, leg. P. Sushkin [ZISP]. Almaty Province: 20 km SE Akzhar, 10. VI. 1986, 3 ♂, leg. V. Kazenas [FSCV]; Vicinity of Akzhar, 44 ° 52 ′ 49 ″ N, 75 ° 53 ′ 11 ″ E, 397 m, on Halimodendron halodendron, 26. V. 2015, 1 ♂, leg. K. Fadeev [ZISP]; 18 km SWW Chundzha, 43.5028 ° N, 79.2134 ° E, 11. V. 2024, 1 ♂, leg. M. Proshchalykin [CAFK]; ibid., on Dodartia orientalis, 11. V. 2024, 6 ♀, leg. M. Proshchalykin [CAFK]; ibid., on Dodartia orientalis, 11. V. 2024, 8 ♂, leg. A. Fateryga [6 ♂, CAFK; 2 ♂, CVMM]; ibid., on Dodartia orientalis, 12. V. 2024, 1 ♂, leg. A. Fateryga [CAFK]; ibid., on Dodartia orientalis, 15. V. 2024, 2 ♀, leg. A. Fateryga [CVMM]. Jetisu Province: 60 km NW Ushtobe, Karatal River, 8. VI. 1986, 1 ♀, leg. V. Kazenas [FSCV]. Kyzylorda Province: Baigakum bei Djulek, 1. V. 1907, 1 ♀, leg. L. Wollmann [ZISP]; ibid., 10. V. 1908, 1 ♀, leg. L. Wollmann [ZISP]; ibid., 11. V. 1908, 2 ♀, leg. L. Wollmann [ZISP]; ibid., 12. V. 1908, 1 ♀, 1 ♂, leg. L. Wollmann [ZISP]; Terenj-Uzjak, 30. V. 1925, 1 ♂, leg. A. Zhelochovtsev [ZMMU]; Perovsk [Kyzylorda], 18. V. 1928, 2 ♂, leg. V. Popov [ZISP]; ibid., 21. V. 1928, 1 ♂, leg. V. Popov [ZISP]. Ulytau Province: S Jezkazgan, S part of Samenkum Sands, 29. V. 1962, 1 ♂, leg. V. Tobias [ZIPS].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4A1DE2506206AF23D7.taxon	distribution	Distribution. Turkmenistan, Uzbekistan, Kazakhstan (Aktobe, Almaty, Jetisu, Kyzylorda, and Ulytau provinces).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4A1DE2506206AF23D7.taxon	biology_ecology	Trophic relationships. Masaris carli is probably monolectic on Dodartia orientalis L. (Mazaceae). Detailed analysis of its flower preferences and flower-visiting behavior will be present in a separate paper (Mauss et al., in preparation). Flower-visiting records (all in Kazakhstan): Caragana halodendron (Pall.) Dum. Cours. (≡ Halimodendron halodendron (Pall.) Voss, Fabaceae); Dodartia orientalis (Mazaceae); Tamarix sp. (Tamaricaceae) (Popov 1948; Fateryga & Proshchalykin 2024; Mauss et al., in preparation).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4B1DE257A1070824CC.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ Туркестан. / Ходжентский / округ. [Turkestan, Khodzhent Okrug] 11. VI. 27. / Н. Умнов [N. Umnov] // Masaris ♀ / gussakovskii Typ / G. Kostylev det. 1932 // к. Гуссаковского [collection of Gussakovskij] // Holotype ♀ / Masaris gussakovskii / Kostylev, 1935 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000318 [pale yellow label] ” [ZISP]. PARATYPE: KAZAKHSTAN. Jambyl Province: Semirechye [Jetisu], Konstantinovsk on Chu River, 12 – 25. VII. 1905, 1 ♀, leg. Matisen [ZISP]. ADDITIONAL MATERIAL: KAZAKHSTAN. Turkestan Province: Kushata, 19 – 21. VI. 1934, 1 ♀, leg. A. Rodd [ZISP]. Additional literature records. KAZAKHSTAN. Turkestan Province: Darbaza, 40 km N Tashkent (Gusenleitner 2002).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4B1DE257A1070824CC.taxon	distribution	Distribution. Tajikistan, Kazakhstan (Jambyl and Turkestan provinces).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCB6D4B1DE257A1070824CC.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2509E0708219C.taxon	description	(Fig. 12 A, C, E, F)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2509E0708219C.taxon	materials_examined	Material examined. HOLOTYPE: ♂, labeled “ [golden disc] // Вершина / Ак-Таша [Top of Ak-Tash] 8000 ʹ. [8000 feet = 2438 m] / 26. VI- 22. Н-К. [N. Kuznetzov] // Тип [Type] / Saryara lon- / gicornis Kuzn. = / = Masaris long / (Kuzn) Kost. / 23 VI 41 [Kostylev’s signature] // Masaris / longicornis / N. Kuzn / Typus // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotype ♂ / Saryara longicornis / Kuznetzov, 1923 [red label] ” [ZMMU]. ADDITIONAL MATERIAL: TAJIKISTAN: Kabadian [Qabodiyon], 21 – 22. VI. 1913, 1 ♀, 1 ♂, leg. A. Golbek [ZISP]. UZBEKISTAN: Ak-Tash, 1 ♀, leg. E. Smirnov [ZMMU]; ibid., 9. VII. 1923, 1 ♂, leg. A. Zhelochovtsev [ZMMU]. KYRGYZSTAN: Sufi-Kurgan, 23. VII. 1979, 1 ♂, leg. Yu. Pesenko [FSCV]; Alai Mts., Gulcha Ravine, Katta-Karakol, 12. VII. 2000, 1 ♂, leg. V. Dubatolov [ISEN]; ibid., 13. VII. 2000, 1 ♀, leg. V. Dubatolov [ISEN]; Kasgharia, lower reaches of Kalta-Bulak River (right inflow of Koksu River), 19. VII. 2003, 4 ♀, leg. V. Dubatolov, E. Nikolayeva, V. Zinchenko, R. Dudko [ISEN]. Additional literature records. KYRGYZSTAN: Transalai Mountain Range, E part, Kaltabulak River gorge, 39 ° 39 ′ N, 73 ° 50 ′ E, 2950 m (Gusenleitner 2002). KAZAKHSTAN. Almaty Province: Ketmen, Mt. S Kirgyzay, 43 ° 17 ′ N, 79 ° 31 ′ E, 2200 m (Gusenleitner 2002).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2509E0708219C.taxon	distribution	Distribution. Afghanistan, Tajikistan, Uzbekistan, Kyrgyzstan, Kazakhstan (Almaty Province).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2509E0708219C.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2546E01FA2235.taxon	materials_examined	(type locality: “ ТЯнь-Шань: южное побережье оЗ. Иссык-Куль, Малый Джаргылчак (около Актырека), сухие холмы со степной растительностью, на цветках Thymus ” [Kyrgyzstan, Tian Shan, south coast of Issyk-Kul Lake, Malyy Dzhargylchak (near Aktyrek), dry hills with steppe vegetation, on flowers of Thymus]), holotype, ♀, ZMMU (examined). Material examined. HOLOTYPE: ♀, labeled “ ИссЫК-КУЛЬ / М. Джаргылчак / Д. ПанфиЛОв [Issyk-Kul Lake, Malyy Dzhargylchak, D. Panfilov] / 26 VI 54 // Holotypus ♀ 1968 / Masaris / tianshanicus Panf. [red label] // Zool. Mus. / Moscow [red font] ” [ZMMU]. PARATYPES: KYRGYZSTAN: Issyk-Kul Lake, Malyy Dzhargylchak, 24. VI. 1954, 2 ♀, leg. D. Panfilov [ZMMU]; ibid., 26. VI. 1954, 1 ♀, leg. D. Panfilov [ZMMU].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2546E01FA2235.taxon	distribution	Distribution. Kyrgyzstan.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2546E01FA2235.taxon	biology_ecology	Trophic relationships. According to Panfilov (1968), the type series was collected at flowers of Thymus sp. (Lamiaceae). Probably broadly oligolectic (sensu Müller & Kuhlmann 2008) on Lamiaceae.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCA6D4B1DE2546E01FA2235.taxon	discussion	Remarks. The taxonomic independence of this species from M. longicornis is doubtful. Males of M. tianshanicus should be collected and compared with those of M. longicornis. Until this is done, M. tianshanicus is provisionally continued to be treated as a separate species closely related to M. longicornis (Fateryga 2025).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCD6D4D1DE2519A0708228B.taxon	description	(Figs 1, 11 A)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCD6D4D1DE2519A0708228B.taxon	description	Description of male (hitherto unknown). Body length (from head to apical margin of tergum II) 4 mm (total body length about 5 mm); forewing length 3.5 mm. Head in frontal view about 1.25 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus with shallowly concave disc, about 1.1 × as wide as long; its apical emargination rather shallow, about 0.2 × as deep as wide, taking about 0.6 of clypeal width, apical teeth nearly rectangular. Distance between lateral ocellus and occiput about 0.8 × as distance between lateral ocelli and about 1.25 × as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.8 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.65 × as width of compound eye at ocular sinus. Occipital carina complete and weakly lamellate, not forming angles. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina weakly lamellate, distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, strongly and evenly convex, broadly rounded posteriorly. Tegula with broadly rounded outer margin produced posterior to blunt medio-posterior angle. Epipleural and episternal sulci weak. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina weak. Dorsal carina of propodeum complete, roundly produced dorso-laterally; lateral carina of propodeum complete, ventro-laterally also produced as nearly right rounded angle. Terga I – VI and sterna II – VI with transculent apical lamella. Tergum VII apically with two rounded lobes and deep emargination between them. Sternum VII + VIII rather flat, apically tridentate, central tooth broader than lateral ones. Genitalia as in Fig. 11 A. Foretibia not modified. Disc of clypeus shining, sparsely punctate, interspaces exceeding puncture diameter, very weakly reticulately microsculptured; punctation becoming denser laterally, where interspaces less than puncture diameter. Punctures on frons as coarse as those on disc of clypeus, interspaces also exceeding puncture diameter, shining, with weak reticulate microsculpture. Sculpture becoming denser and duller on vertex behind ocelli where interspaces much narrower than puncture diameter. Gena rather dull, with fine and very sparse punctures and indistinct microsculpture. Pronotum and scutellum punctate coarser than clypeus and frons, interspaces usually about as puncture diameter, shining and completely smooth. Punctures on scutum even slightly coarser than those on pronotum and scutellum, interspaces also about as puncture diameter, shining and completely smooth. Tegula with weak, indistinct microsculpture and few rather fine punctures on posterior half. Dorsal and ventral mesepisterna and mesepimeron sculptured much denser than pronotum and scutellum, interspaces duller and less than puncture diameter. Posterior surface of metanotum shining, with smoothened micropunctures. Metapleuron and lateral surface of propodeum rather dull, with shagreened reticulate sculpture. Dorso-lateral surface of propodeum with shallow irregular punctures, interspaces shining, reaching puncture diameter. Posterior surface of propodeum shining, densely punctate, interspaces mostly less than puncture diameter. Metasomal terga dull, with reticulate microsculpture and shallow dense punctures; both punctures and interspaces increasing in size towards apical margins of terga; largest punctures on terga much finer than those on scutum and scutellum. Metasomal sterna I – VI with fine shallow punctures, interspaces reaching puncture diameter, rather shining but with distinct reticulate microsculpture. Punctures on sternum VII much coarser than those on preceding sterna, interspaces mostly less than puncture diameter, shining, with less distinct microsculpture. Mandible, labrum, clypeus, frons, pronotum, and scutum with very short whitish setae about as long as 1 / 3 of diameter of median ocellus. Some setae on vertex about as long as 1 / 2 of diameter of median ocellus. Compound eye bare. Scutellum and mesopleuron without visible setae. Propodeum, metasoma, and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with short erect stiff setae approximately as long as 2 / 3 of diameter of median ocellus, those setae on mid and hind legs much shorter. Basal color black except reddish tergum VII and sternum VII. Following parts whitish to pale yellow: mandible except black base, labrum, clypeus, frons, large spot on gena, small lateral spots on vertex, pronotum, tegula, small postero-medial spot on scutum, nearly whole scutellum except black band antero-medially, posterior (lower) half of metanotum, dorsal mesepisternum, fused spots on ventral mesepisternum and mesepimeron, large lateral spots on propodeum, apical bands on terga I – VI, apical spots on tergum VII and sternum VII, small lateral spots on sternum VII, legs approximately from middle of femora. Antenna deep yellow. Wings transparent; veins and pretostigma pale brown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCD6D4D1DE2519A0708228B.taxon	materials_examined	Material examined. LECTOTYPE (designated by Fateryga (2025: 228 )): ♀, labeled “ [golden disc] // Tschikisch- / ljar. Pom. [Pomeranzev] // Jugurtia? / chlorotica ♀ / F. Morawitz // к. Ф МОРавица [collection of F. Morawitz] // Lectotype ♀ / Jugurtia chlorotica / Morawitz, 1888 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000330 [pale yellow label] ” [ZISP]. ADDITIONAL MATERIAL: TURKMENISTAN: Usun-Ada [Uzun-Ada], 1 ♀ [ZISP]. UZBEKISTAN: Qashqadaryo Province, Muborak District, 5 km N Muborak, 39 ° 18 ′ 31 ″ N, 65 ° 08 ′ 20 ″ E, 15. V. 2015, 1 ♀, 1 ♂, leg. M. Mokrousov, M. Proshchalykin, K. Samartsev [CAFK].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCD6D4D1DE2519A0708228B.taxon	distribution	Distribution. Turkmenistan, Uzbekistan.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCD6D4D1DE2519A0708228B.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4E1DE251D20708267C.taxon	description	(Figs 11 B, 13 A)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4E1DE251D20708267C.taxon	materials_examined	Material examined. LECTOTYPE (designated by Fateryga & Buyanjargal (2022: 13 )): ♀, labeled “ [golden disc] // Имам-баба [Imam-Baba] / 15 – 24. V. 934 / А. и А. Шестаковы [A. and A. Shestakovs] // Quartinia / funebris n. sp / G. Kostylev det. 1933 // Lectotypus ♀ / Quartinia funebris / Kostylev, 1935 / des. Fateryga, 2021 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000331 [pale yellow label] ” [ZISP]. PARALECTOTYPES: TURKMENISTAN: Imam-Baba, 15 – 24. V. 1932, 2 ♀, leg. A. Shestakov [ZMMU]; ibid., 15 – 24. V. 1932, 1 ♀, 1 ♂, leg. A. Shestakov, A. Shestakova [ZISP]; ibid., 15 – 24. V. 1934, 9 ♀, 2 ♂, leg. A. Shestakov, A. Shestakova [ZISP]. ADDITIONAL MATERIAL: TURKMENISTAN: Imam-Baba, 1932, 1 ♀, leg. A. Shestakov [ZISP]; ibid., 19. V. 1932, 2 ♀, 1 ♂, leg. Kuzjakin [ZMMU]; 25 km E Ashgabat, 20. VI. 1976, 2 ♀, leg. N. Kurzenko [FSCV]; Vicinity of Ashgabat, 21. VI. 1976, 4 ♀, leg. N. Kurzenko [FSCV]; 30 km W Kazandzik [Bereket], Akhcha-Kuyma, 4. VI. 1988, 1 ♀, leg. N. Kurzenko [FSCV]; ibid., 5. VI. 1988, 1 ♀, leg. N. Kurzenko [FSCV]. KAZAKHSTAN. Kyzylorda Province: 20 km S Yany-Kurgan [Zhanakorgan], Kyzylkum Desert, 14 – 15. VII. 1990, 1 ♀, leg. V. Kazenas [FSCV].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4E1DE251D20708267C.taxon	distribution	Distribution. Turkmenistan, Kazakhstan (Kyzylorda Province).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4E1DE251D20708267C.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	description	(Fig. 2)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	diagnosis	Diagnosis. The species is unique among other Central Asian members of the genus Quartinia in having the outer margin of the tegula not produced posterior to roundly-acute posterior end and the posterior end not produced medio-apically (Fig. 2 A), completely dull frons and scutum, with a strong reticulate microsculpture between macropunctures (Fig. 2 A, C), and predominately yellow coloration (Fig. 2 A – D).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	description	Description. Female. Body length (from head to apical margin of tergum II) 2.5 mm (total body length about 3.5 mm); forewing length 2 mm. Head in frontal view about 1.3 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus moderately and rather evenly convex, about 1.45 × as wide as long; its apical emargination shallow, about 0.2 × as deep as wide, taking about 0.5 of clypeal width, apical teeth blunt. Distance between lateral ocellus and occiput about 0.75 × as distance between lateral ocelli and about 1.1 × as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.15 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.5 × as width of compound eye at ocular sinus. Occipital carina smoothened at lower part of gena and not forming angle. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina indistinct. Scutellum separated from mesoscutum by transverse suture, slightly convex, roundly truncate posteriorly. Tegula with posterior end not produced; outer margin of tegula not produced posterior to roundly-acute posterior end. Epipleural and episternal sulci distinct. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal and lateral carinae of propodeum smoothened. Terga I – V and sterna II – V with transculent apical lamella. Clypeus dull, reticulately microsculptured, without distinct punctures. Frons and vertex sparsely punctate with very shallow punctures, interspaces exceeding puncture diameter, dull, reticulately microsculptured. Gena without visible punctures, similarly to clypeus. Pronotum, scutum, and scutellum sparsely punctate similarly to clypeus but punctures slightly deeper, interspaces also exceeding puncture diameter, dull, reticulately microsculptured. Other parts of mesosoma and whole metasoma mostly without punctures, reticulately microsculptured. Setation weakly developed. Mandible covered with thin erect whitish setae about as long as diameter of median ocellus. Clypeus with erect whitish setae reaching in length 2 / 3 of diameter of median ocellus; frons and vertex with erect whitish setae reaching in length 1 / 3 of diameter of median ocellus. Compound eye bare. Mesosoma without visible setae. Metasoma and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with erect stiff setae approximately as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Coloration predominately yellow except the following: compound eye and ocelli dark, propleuron and lower part of ventral mesepisternum brown, dorsal side of antennal club, large spot on frons, ventral side of pronotum, most part of scutum, most part of ventral mesepisternum, mesepimeron, metapleuron, lateral surface of propodeum, and basal parts of metasomal terga light reddish-brown to ferruginous. Wings transparent; veins and pretostigma brownish-yellow. Male. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ окр. Урги / с. МонголиЯ / КОЗЛОв [Vicinity of Urga (currently Ulaanbaatar), N Mongolia, Kozlov] 1 – 4. VI. 909 // Holotypus ♀ / Quartinia kozlovi / Fateryga [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000374 [pale yellow label] ” [ZISP].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	etymology	Etymology. The new species is named after Petr K. Kozlov (1863 – 1935), a famous Russian and Soviet traveler and explorer of Central Asia, who collected the holotype.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	distribution	Distribution. Mongolia.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFCF6D4F1DE252C507082724.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	description	(Figs 3, 4, 11 C)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	diagnosis	Diagnosis. The species is closely related to Quartinia mongolica (Morawitz, 1889) but differs from it by a distinctly larger size (total body length 4.5 – 5.5 mm versus 3.5 – 4 mm in Q. mongolica), predominately black coloration with a pale yellow pattern usually outlined by ferruginous and a reddish to reddish-brown apical tergum in both sexes (Figs 3 A, B, D, 4 A, C, D) (versus a black coloration with an extensive orange-yellow pattern and a reddish-yellow apical tergum in Q. mongolica, Fig. 13 B), and a less broad cupula in relation to the remaining parts of the male genitalia (Fig. 11 C) (versus a broader cupula in Q. mongolica, Fig. 11 D).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	description	Description. Female. Body length (from head to apical margin of tergum II) 4 – 4.5 mm (total body length about 5 – 5.5 mm); forewing length 3.5 mm. Head in frontal view about 1.3 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus with concave disc, about 1.25 × as wide as long; its apical emargination rather deep, about 0.4 × as deep as wide, taking about 0.5 of clypeal width, apical teeth broadly rounded and narrowly lamellate. Distance between lateral ocellus and occiput about 1.15 × as distance between lateral ocelli and about 1.3 × as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.6 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.75 × as width of compound eye at ocular sinus. Occipital carina complete, forming indistinct obtuse angle just below ocular sinus. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina weakly lamellate, distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, slightly convex, broadly rounded posteriorly. Tegula with broadly rounded outer margin produced posterior to blunt medio-posterior angle. Epipleural and episternal sulci weak. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal carina of propodeum complete, produced dorso-laterally as nearly right rounded angle; lateral carina of propodeum complete, ventro-laterally also produced as nearly right rounded angle. Terga I – V and sterna II – V with transculent apical lamella. Disc of clypeus rather shining, sparsely punctate, interspaces much exceeding puncture diameter on median line, weakly reticulately microsculptured; punctation becoming denser laterally, where interspaces less than puncture diameter. Punctures on frons as coarse as those on disc of clypeus, interspaces usually exceeding puncture diameter, rather shining but with distinct reticulate microsculpture. Sculpture becoming denser and duller on vertex behind ocelli where interspaces much narrower than puncture diameter. Gena shining, with fine and very sparse punctures and indistinct microsculpture. Pronotum punctate coarser than clypeus and frons, interspaces usually exceeding puncture diameter, rather shining but with distinct reticulate microsculpture. Punctures on scutum and scutellum as coarse as those on pronotum but interspaces much larger, several times exceeding puncture diameter and more shining, with weak microsculpture. Tegula with weak reticulate microsculpture and few fine punctures on posterior half. Dorsal and ventral mesepisterna and mesepimeron sculptured denser and slightly coarser than pronotum. Posterior surface of metanotum shining, with smoothened micropunctures. Metapleuron and lateral surface of propodeum rather dull, with shagreened reticulate sculpture. Dorso-lateral surface of propodeum with shallow cell-like sculpture. Posterior surface of propodeum shining, with shallow dense punctures, interspaces less than puncture diameter. Metasomal terga dull, with reticulate microsculpture and shallow dense punctures; both punctures and interspaces increasing in size towards apical margins of terga; largest punctures on terga much finer than those on scutum and scutellum. Metasomal sterna I – V regularly punctuate with fine shallow punctures, interspaces exceeding puncture diameter, rather shining, with weak reticulate microsculpture. Punctures on sternum VI denser than those on preceding sterna, interspaces not exceeding puncture diameter, duller. Mandible and labrum covered with thin erect pale setae slightly exceeding in length diameter of median ocellus. Clypeus, frons, vertex, and pronotum with short pale setae several times less in length than diameter of median ocellus. Compound eye bare. Scutum, scutellum, and mesopleuron without visible setae. Propodeum, metasoma, and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with short erect stiff setae approximately as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Basal color black. Following parts pale yellow outlined by ferruginous: elongate medial spot on clypeus (reduced in one specimen), spot on upper part of gena (reduced in two specimens), antenna except brown dorsal sides of scape and club, anterior band on pronotum, tegula, C-shaped spot on posterior half of scutellum, thin apical band on scutellum, metanonum, apical bands on terga I – V, legs from tops of femora. Distal half of mandible, labrum, metasomal tergum VI, and sternum VI reddish to reddish-brown. Wings transparent; veins and pretostigma pale brown. Male. Body length (from head to apical margin of tergum II) 3.5 mm (total body length about 4.5 mm); forewing length 3 mm. Structure resembles that in female except the following: head in frontal view about 1.35 × as wide as long, clypeus about 1.2 × as wide as long; its apical emargination shallow, about 0.2 × as deep as wide, apical teeth blunt but not broadly rounded. Distance between lateral ocellus and occiput about as distance between lateral ocelli and about 1.65 × as distance between lateral ocellus and compound eye. Width of gena in front of ocular sinus about 0.5 × as width of compound eye at ocular sinus. Tergum VII apically with two rounded lobes and deep emargination between them. Sternum VII + VIII rather flat, apically tridentate, central tooth broader and longer than lateral ones. Genitalia as in Fig. 11 C. Foretibia not modified. Sculpture and setation as in female except that clypeus more finely and sparsely punctate, interspaces reaching several puncture diameters on whole disc. Sternum VI sculptured as previous sterna; sternum VII + VIII sculptured as sternum VI in female. Coloration resembles that in female but clypeus almost completely whitish-yellow and spot on gena absent. Metasomal tergum VI with apical band as on preceding terga; sternum VI as preceding sterna; tergum VII and sternum VII + VIII reddish.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ МОНГОЛИЯ, Кобдоский / аймак, р. Бодончин-Гол / 12 км ЮЗ АлтаЯ (нижн.) / КоЗлов [Mongolia, Khovd Province, Bodonchin-Gol River, 12 km SW Altai] 22. УП. 970 // Q. funebris Kost / det. Abasheev R. // ♀ Quartinia mongolica / (Morawitz, 1889) / det. Fateryga, 2022 // Holotypus ♀ / Quartinia mikhaili / Fateryga [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000375 [pale yellow label] ” [ZISP]. PARATYPES: KAZAKHSTAN. Kyzylorda Province: 12 km SEE Dyurmen-Tyube Station, sands, on Convolvulus, 5. VI. 1979, 1 ♀, leg. Yu. Pesenko [ZISP]. MONGOLIA: Khovd Province, Bulgan Sum, Bulgan-Gol River, oasis, 4 – 5. VII. 1966, 1 ♀, leg. B. Namhaidorj [IBUM]; Khovd Province, Altai Sum, 5 km SW Altai, lower reaches of Bodonchin-Gol River, 11. VIII. 1968, 2 ♀, leg. M. Kozlov [ZISP]; Khovd Province, Altai Sum, 12 km SW Altai, Bodonchin-Gol River, 22. VII. 1970, 1 ♂ leg. M. Kozlov [ZISP].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	etymology	Etymology. The new species is named in memory of Mikhail A. Kozlov (1936 – 2006), an outstanding Soviet and Russian entomologist, who collected most specimens of the type series.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	distribution	Distribution. Kazakhstan (Kyzylorda Province), Mongolia.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC16D421DE251D204CB2240.taxon	biology_ecology	Trophic relationships. The specimen from Kazakhstan was collected at Convolvulus sp. (Convolvulaceae), according to its label.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D431DE251D207082714.taxon	description	(Figs 11 D, 13 B)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D431DE251D207082714.taxon	materials_examined	Material examined. LECTOTYPE (designated by Fateryga & Buyanjargal (2022: 11 )): ♀, labeled “ [golden disc] // Mongol. mer. / Tdschin-Ta-ssu. [China: Gansu] // к. Ф МОРавица [collection of F. Morawitz] // Celonites? / mongolicus ♀ / F. Morawitz // Lectotypus ♀ / Celonites mongolicus / Morawitz, 1889 / des. Fateryga, 2021 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000332 [pale yellow label] ” [ZISP]. PARALECTOTYPES: CHINA: Gansu, Tdschin-Ta-ssu [Dschin-Tasu], 1 ♂ [ZISP]; ibid., 11. VII, 1 ♀, leg. G. Potanin [ZISP].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D431DE251D207082714.taxon	distribution	Distribution. China (Gansu).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D431DE251D207082714.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D451DE253DD070824CC.taxon	description	(Figs 5, 11 F)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D451DE253DD070824CC.taxon	description	Description of male (hitherto unknown). Body length (from head to apical margin of tergum II) 3 mm (total body length about 4 mm); forewing length 2.5 mm. Head in frontal view about 1.25 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus strongly evenly convex, about 1.3 × as wide as long; its apical emargination rather shallow, about 0.25 × as deep as wide, taking about 0.55 of clypeal width, apical teeth rounded and lamellate. Distance between lateral ocellus and occiput about 0.9 × as distance between lateral ocelli and about 0.7 × as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.5 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.7 × as width of compound eye at ocular sinus. Occipital carina complete, forming indistinct obtuse angle at lower part of gena. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina indistinct. Scutellum separated from mesoscutum by transverse suture, strongly and evenly convex, broadly rounded posteriorly. Tegula with strongly produced posterior end and nearly straight inner margin. Epipleural and episternal sulci distinct. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal carina of propodeum complete, rounded dorso-laterally; lateral carina of propodeum complete, ventro-laterally also produced as nearly right rounded angle. Terga I – VI and sterna II – VI with transculent apical lamella. Tergum VII apically with two rounded lobes and moderately deep emargination between them. Sternum VII + VIII rather flat, apically trilobed, central lobe broader and longer than lateral ones. Genitalia as in Fig. 11 F. Foretibia not modified, just slightly thickened. Clypeus dull, with shallow and very dense indistinct punctures. Frons and vertex very densely punctate with rather fine but distinct punctures, interspaces less than puncture diameter, with strong reticulate microsculpture. Gena with similar but somewhat coarser and sparser punctures than those on frons and vertex. Pronotum and anterior part of scutum punctate coarser than frons and vertex, interspaces less than puncture diameter, moderately dull, with reticulate microsculpture. Punctures and interspaces becoming even larger on posterior part of scutum and scutellum where interspaces exceeding puncture diameter, moderately dull, with distinct reticulate microsculpture. Tegula with weak, indistinct microsculpture and few very fine indistinct punctures on posterior half. Dorsal and ventral mesepisterna and mesepimeron sculptured distinctly denser and coarser than pronotum and anterior part of scutum, interspaces duller and much less than puncture diameter. Posterior surface of metanotum dull, mostly with microsculpture. Metapleuron and lateral surface of propodeum rather dull, with shagreened sculpture. Dorso-lateral surface of propodeum with fine cell-like irregular sculpture. Posterior surface of propodeum moderately dull, with shallow dense punctures, interspaces less than puncture diameter. Metasomal terga dull, mostly with reticulate microsculpture, fine dense and shallow punctures visible mostly on terga I and II. Metasomal sterna moderately shining, with fine reticulate microsculpture, only sternum VII with regular dense shallow punctures and interspaces about as puncture diameter. Mandible and labrum covered with thin erect pale setae about as long as diameter of median ocellus. Clypeus, frons, vertex, and pronotum with short pale setae about three times shorter than diameter of median ocellus. Compound eye bare. Scutum, scutellum, and mesopleuron without visible setae. Propodeum, metasoma, and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with short erect stiff setae approximately as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Basal color black to dark brown. Following parts pale yellow: mandible except reddish-brown teeth, labrum, clypeus, antenna except most part of club, anterior band on pronotum, tegula (except small brownish antero-central part), narrow apical band on scutellum, apical bands on terga I – VII, legs from tops of femora. Posterior part of sternum VII reddish-brown. Wings moderately fuscous; veins and pretostigma brown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D451DE253DD070824CC.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ ущ. Кондара [Kondara Canyon] / 7 - VIII 37 / В. Гуссаковск. [V. Gussakovskij] // Quartinia / popovi Kost. ♀ / G Kostylev det. 1938 // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotype ♀ / Quartinia popovi / Richards, 1962 [red label] ” [ZMMU]. ADDITIONAL MATERIAL: TAJIKISTAN: SE of Kulyab [Kulob], Shuroabid [Shurobod], 2150 m, 27. VIII. 1933, 1 ♀, leg. V. Popov [ZISP]; 35 km N Stalinabad [Dushanbe], Kvak Place, 2000 m, 5. VIII. 1937, 9 ♀, leg. V. Gussakovskij [ZISP]; ibid., 27. VIII. 1937, 6 ♀, leg. V. Gussakovskij [ZISP]; Kvak, 21. VII. 1937, 2 ♀, leg. V. Gussakovskij [ZISP]; ibid., 17. VIII. 1937, 9 ♀, leg. V. Gussakovskij [ZISP]; Gissar-Gebirge, 2000 m, 17 – 27. VIII. 1937, 1 ♀ [ZMMU]; Kvak Place, 20. VII. 1938, 2 ♀, leg. V. Gussakovskij [ZISP]; Khorog-Dasht [Dasht-Khorog], 3. VIII. 1939, 1 ♀, leg. A. Saakyan-Baranova [ZISP]; Gafilabad [Gofilobod], 2500 m, 12. VIII. 1940, 2 ♀, leg. V. Gussakovskij [ZISP]; ibid., 14. VIII. 1940, 6 ♀, leg. V. Gussakovskij [ZISP]; ibid., 23. VIII. 1940, 2 ♀, leg. V. Gussakovskij [ZISP]; Kok-Kul Place, 3000 m, 20 – 22. VIII. 1940, 1 ♀, leg. V. Gussakovskij [ZISP]; Kukhilal, Panj River, 2400 m, 18. VII. 1956, 1 ♀, leg. V. Tobias [ZISP]. UZBEKISTAN: N slopes of Gissar Range, Khan-Takhta Place near Yakkabag, V. 1933, 1 ♀, leg. Veltishchev [ZISP]; N slope of Gissar Range, Mt. Maydanak, 2800 m, 31. VIII. 1933, 1 ♀, leg. Veltishchev [ZISP]. KYRGYZSTAN: Transalai Mountain Range, Berk-Suu Ravine, 39 ° 29 ′ N, 72 ° 02 ′ E, ~ 2600 m, 19. VII. 1998, 1 ♀, leg. D. Milko [CSZT]; Transalai Mountain Range, W part, Berksu River gorge, 39 ° 28 ′ N, 72 ° 01 ′ E, 2600 m, 19. VII. 1998, 2 ♂, leg. I. Makogonova [CSZT].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D451DE253DD070824CC.taxon	distribution	Distribution. Tajikistan, * Uzbekistan, Kyrgyzstan.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC26D451DE253DD070824CC.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	materials_examined	(Figs 6, 11 E)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	description	Description of male (hitherto unknown). Body length (from head to apical margin of tergum II) 1.5 mm (total body length about 2 mm); forewing length 1.5 mm. Head in frontal view about 1.3 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus slightly evenly convex, about 1.5 × as wide as long; its apical emargination very shallow, taking about 0.45 of clypeal width, apical teeth blunt. Distance between lateral ocellus and occiput about 0.9 × as distance between lateral ocelli and about 1.5 × as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 1.6 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.4 × as width of compound eye at ocular sinus. Occipital carina complete, forming indistinct obtuse angle at lower part of gena. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina indistinct. Scutellum separated from mesoscutum by transverse suture, moderately convex, broadly rounded posteriorly. Tegula with broadly rounded outer margin produced posterior to blunt medio-posterior angle. Epipleural and episternal sulci distinct. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal and lateral carinae of propodeum mostly rounded. Terga I – VI and sterna II – VI with transculent apical lamella. Tergum VII apically with two rounded lobes and moderately deep emargination between them. Sternum VII + VIII rather flat, apically trilobed, central lobe broader than lateral ones. Genitalia as in Fig. 11 E. Structure of foretibia unknown (not present in studied specimen). Clypeus dull, with shallow and indistinct punctures. Frons, vertex, and gena finely and sparsely punctate, interspaces moderately shining, distinctly reticulately microsculptured. Pronotum, scutum, scutellum, dorsal and ventral mesepisterna, and mesepimeron also finely and sparsely punctate but punctures slightly larger than those on frons, vertex, and gena, interspaces also moderately shining, distinctly reticulately microsculptured. Other parts of mesosoma and metasoma reticulately microsculptured, without distinct punctures. Setation weakly developed. Mandible, labrum, clypeus, frons, vertex, and pronotum with minute erect setae much shorter than diameter of median ocellus. Other parts of body mostly without visible setae. Legs from coxae to femora mostly with short appressed setae. Mid and hind tibiae and tarsi with very short setae. Basal color reddish-brown to dark brown. Following parts pale yellow to yellow: mandible except reddish-brown base, labrum, clypeus, most part of pronotum, tegula (except small brownish antero-central part), large central spot on scutellum, narrow apical band on scutellum, apical bands on terga I – VI, large apical spot on tergum VII, legs from tops of femora. Antenna deep yellow. Wings transparent; veins and pretostigma brown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ [golden disc] // Красноводск [Krasnovodsk (currently Türkmenbaşy)] / 18. VI. 928 / В. Гуссаковский [V. Gussakovskij] // Quartinia ♀ / pusilla, Typ / G. Kostylev det. 1933 // Quartinia / pusilla Kost / Тип. Колл. В. / Гуссаковского [Type. Collection of V. Gussakovskij] / 1941 23 VI [Kostylev’s signature] // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotype ♀ / Quartinia pusilla / Kostylev, 1935 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000379 [pale yellow label] ” [ZISP]. ADDITIONAL MATERIAL: TURKMENISTAN: Repetek, on Alhagi camelorum, 21. VII. 1937, 1 ♀, leg. G. Kostylev [ZMMU]; ibid., on Alhagi camelorum, 22. VII. 1937, 1 ♀, leg. G. Kostylev [ZMMU]. KAZAKHSTAN. Kyzylorda Province: 20 km S Yany-Kurgan [Zhanakorgan], Kyzylkum Desert, 14 – 15. VII. 1990, 12 ♀, 1 ♂, leg. V. Kazenas [FSCV].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	distribution	Distribution. Turkmenistan, Kazakhstan (Kyzylorda Province).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	biology_ecology	Trophic relationships. The specimens from Repetek were collected at Alhagi maurorum Medik. (= A. camelorum Fisch., Fabaceae), according to their labels.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC46D471DE2509E01AC25BC.taxon	discussion	Remarks. After the publication of my paper (Fateryga 2025), curators of both ZISP and ZMMU decided to return the holotype of this species from ZMMU to ZISP, since ZISP was indicated as the depository in the original publication by Kostylev (1935), while the holotype was actually deposited in ZMMU until present.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC66D581DE2500E069625BC.taxon	description	(Figs 7, 11 G, 13 C – G)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC66D581DE2500E069625BC.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ [golden disc] // Тартугай [Tartugay] / 3 - 15. VI. 1929 / А. Шестакова [A. Shestakova] // Quartinia ♀ / shestakovi Typ / G. Kostylev det. 1932 // Holotype ♀ / Quartinia shestakovi / Kostylev, 1935 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000333 [pale yellow label] ” [ZISP]. PARATYPES: UZBEKISTAN: NW Bukhara, Yargak, 26. V. 1928, 1 ♂, leg. L. Zimin [ZISP]; NW Bukhara, Kumak, 14. VI. 1929, 16 ♀, 10 ♂, leg. L. Zimin [16 ♀, 9 ♂, ZISP; 1 ♂, ZMMU]; 50 km N Bukhara, Bag-Abzal, 5. IX. 1931, 2 ♀, leg. A. Zhelochovtsev [ZMMU]; ibid., 10. IX. 1931, 3 ♀, leg. A. Zhelochovtsev [ZMMU]. ADDITIONAL MATERIAL: TAJIKISTAN: Dzhilikul [Jilikul], Staraya Pristan, 8. VII. 1941, 1 ♂, leg. V. Gussakovskij [ZISP]; ibid., 12. VII. 1941, 1 ♀, leg. V. Gussakovskij [ZISP]; ibid., 13. VII. 1941, 2 ♂, leg. V. Gussakovskij [ZISP]; ibid., 17. VII. 1941, 11 ♀, 5 ♂, leg. V. Gussakovskij [ZISP]; ibid., 18. VII. 1941, 3 ♀, 3 ♂, leg. V. Gussakovskij [ZISP]; ibid., 20. VII. 1941, 11 ♀, 3 ♂, leg. V. Gussakovskij [ZISP]; ibid., 21. VII. 1941, 8 ♀, 1 ♂, leg. V. Gussakovskij [ZISP]; ibid., 22. VII. 1941, 1 ♂, leg. V. Gussakovskij [ZISP]; Staraya Pristan, Vakhsh River, on Halocharis hispida, 12. IX. 1948, 1 ♀, leg. V. Popov [ZISP]. UZBEKISTAN: Vicinity of Zarafshon [River], Kumak, 2. VI. 1929, 1 ♂, leg. L. Zimin [ZISP]; Dzhuma [Juma] Station, Uzbek alfalfa brigade of ZISP, on Horaninovia ulicina, 23. VII. 1937, 3 ♀, 1 ♂, leg. V. Popov [ZISP]; ibid., on Horaninovia ulicina, 25. VII. 1937, 12 ♀, leg. V. Popov [ZISP]; Kurgan-Tyube [Qoʻrgʻontepa], Fergana alfalfa brigade of ZISP, on Horaninovia ulicina, 16. VII. 1938, 1 ♀, leg. V. Popov [ZISP]; ibid., on Horaninovia ulicina, 25. VII. 1938, 1 ♀, leg. V. Popov [ZISP]; Kokand Sands, 40 ° 35 ′ 49 ″ N, 70 ° 48 ′ 22 ″ E, on Limonium otolepis, 18. V. 2023, 1 ♀, 1 ♂, leg. A. Fateryga [CAFK]. KAZAKHSTAN. Akmola Province: 30 km E Kokchetav [Kokshetau], 16. VII. 1977, 1 ♂, leg. V. Kazanas [FSCV].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC66D581DE2500E069625BC.taxon	distribution	Distribution. Tajikistan, Uzbekistan, Kazakhstan (Akmola and Kyzylorda provinces).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC66D581DE2500E069625BC.taxon	biology_ecology	Trophic relationships. According to Popov (1948), this species is oligolectic on Chenopodiaceae and was recorded at flowers of Horaninovia ulicina Fisch. & C. A. Mey. in Uzbekistan and “ Salsola sp. ” in Tajikistan, but in the latter case he probably meant Halocharis hispida (Schrenk) Bunge mentioned on one label. However, I observed a pair of Q. shestakovi at flowers of Limonium otolepis (Schrenk) Kuntze (Plumbaginaceae) in Uzbekistan, and at least the male consumed pollen from them (Fig. 7). Therefore, the species is probably polylectic.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFC66D581DE2500E069625BC.taxon	discussion	Remarks. The specimen from the Akmola Province is lacking both the head and the metasoma. Its identification is preliminary and requires further confirmation. The mesosoma is smaller than that in other specimens of Q. shestakovi but still larger than that of the next species described below.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	description	(Figs 8, 9, 11 H)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	diagnosis	Diagnosis. The species is closely related to Quartinia shestakovi but differs from it by a remarkably smaller size (total body length about 3 mm versus 4 – 5 mm in Q. shestakovi) and larger reticulately microsculptured interspaces between macropunctures on the frons, several times exceeding the puncture diameter (Figs 8 B, 9 B) (versus smaller interspaces equal to the puncture diameter or not much larger in Q. shestakovi, Fig. 13 C, F).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	description	Description. Female. Body length (from head to apical margin of tergum II) 2.5 mm (total body length about 3 mm); forewing length 2 mm. Head in frontal view about 1.15 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus moderately and rather evenly convex, about 1.25 × as wide as long; its apical emargination shallow, about 0.15 × as deep as wide, taking about 0.5 of clypeal width, apical teeth rounded. Distance between lateral ocellus and occiput about 1.1 × as both distance between lateral ocelli and between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.6 × as long as wide in dorsal view. Width of gena in front of ocular sinus about 0.5 × as width of compound eye at ocular sinus. Occipital carina complete, evenly curved, not forming angles. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, slightly convex, broadly rounded posteriorly. Tegula with strongly produced posterior end and strongly emarginate inner margin, so that posterior end of tegula produced distinctly medially. Epipleural and episternal sulci distinct. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal carina of propodeum complete, roundly produced dorso-laterally; lateral carina of propodeum complete, ventro-laterally produced as nearly right rounded angle. Terga I – V and sterna II – V with transculent apical lamella. Clypeus dull, finely and densely punctate, interspaces about as puncture diameter, reticulately microsculptured. Punctures on frons coarser and sparser than those on clypeus, interspaces exceeding puncture diameter, more shining than those on clypeus but also with distinct reticulate microsculpture. Sculpture becoming denser on vertex behind ocelli and gena where interspaces generally not exceeding puncture diameter. Pronotum and scutum punctate coarser than frons, interspaces exceeding puncture diameter, rather shining but with distinct reticulate microsculpture. Scutellum punctate similarly to pronotum and scutum but punctures sparser. Tegula almost smooth, without punctures and with weak reticulate microsculpture. Dorsal and ventral mesepisterna and mesepimeron sculptured similarly to pronotum and scutum. Posterior surface of metanotum, metapleuron, as well as lateral and posterior surfaces of propodeum rather dull, with shagreened miscrosculpture. Dorso-lateral surface of propodeum with few punctures in addition to miscrosculpture. Metasomal terga moderately shining, with reticulate microsculpture and shallow punctures on posterior halves; punctures much finer than those on pronotum and scutum. Metasomal sterna punctuate similarly to posterior halves of terga. Mandible and labrum covered with erect pale setae about as long as diameter of median ocellus. Clypeus, frons, vertex, and pronotum with short pale setae about three times shorter than diameter of median ocellus. Compound eye bare. Scutum, scutellum, and mesopleuron without visible setae. Propodeum, metasoma, and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with short erect stiff setae approximately as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Basal color reddish-brown. Antenna brownish-yellow to yellow. Anterior band on pronotum, tegula, C-shaped spot on posterior half of scutellum, thin apical band on scutellum, lateral spots on propodeum (often reduced), apical band on tergum I and sometimes also on tergum II yellow. Legs from tops of femora also more or less yellow. Wings transparent; veins and pterostigma brown. One female specimen has aberrant coloration, typical of male (see below and Fig. 8 F). Male. Structure and size resemble those in female except the following: head in frontal view about 1.2 × as wide as long, clypeus about 1.2 × as wide as long, apical teeth of clypeus rather blunt than rounded. Distance between lateral ocellus and occiput about as distance between lateral ocelli and about 1.1 × as distance between lateral ocellus and compound eye. Tergum VII apically with two rounded lobes and narrow, moderately deep emargination between them. Sternum VII + VIII rather flat, apically trilobed, central lobe broader than lateral ones and apically rounded to truncate. Genitalia as in Fig. 11 H. Foretibia not modified. Sculpture and setation resemble those in female. Coloration remarkably different from that in female. Basal color pale brown except that upper part of frons, vertex, and scutum rather reddish-brown to black. Following parts pale yellow: mandible, labrum, clypeus, antenna except brownish-yellow distal half of club, spot on upper part of gena, pronotum, tegula, large crescent spot on scutellum, thin apical band on scutellum, dorsal mesepisternum, lateral spots on propodeum, apical bands on terga I – VII. Legs from tops of femora also more or less pale yellow. Wings transparent; veins and pterostigma pale brown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	materials_examined	Material examined. HOLOTYPE: ♀, labeled “ СтараЯ Пристань / р. Вахш, Таджик. / ПОПОв [Staraya Pristan, Vakhsh River, Tajikistan, Popov] 12. IX 948 / Halocharis hispida [on underside] // Holotypus ♀ / Quartinia sovietica / Fateryga [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000376 [pale yellow label] ” [ZISP]. PARATYPES: TAJIKISTAN: Staraya Pristan, Vakhsh River, on Halocharis hispida, 12. IX. 1948, 9 ♀, leg. V. Popov [ZISP]; ibid., on Halocharis hispida, 12. IX. 1948, 16 ♀, leg. V. Rudolf [ZISP]; Dzhilikul [Jilikul], Staraya Pristan, 14. VII. 1941, 1 ♀, leg. V. Gussakovskij [ZISP]; ibid., 20. VII. 1941, 1 ♀, leg. V. Gussakovskij [ZISP]; ibid., 21. VII. 1941, 11 ♀, 5 ♂, leg. V. Gussakovskij [ZISP]; ibid., 22. VII. 1941, 38 ♀, 25 ♂, leg. V. Gussakovskij [ZISP].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	etymology	Etymology. The specific name is an adjective meaning “ Soviet ” as a reference to the USSR, in recognizing a great contribution of the Soviet hymenopterists, such as Georg (Yuriy A.) Kostylev (1889 – 1942), Vladimir B. Popov (1902 – 1960), Vsevolod V. Gussakovskij (1904 – 1948), etc., to the study of the Central Asian fauna (particularly the pollen wasps).	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	distribution	Distribution. Tajikistan.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFD96D5A1DE2500E01F227B4.taxon	biology_ecology	Trophic relationships. According to Popov (1948), this species was recorded at flowers of “ Salsola sp. ” in Tajikistan, but he probably meant Halocharis hispida (Chenopodiaceae) mentioned on labels.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	description	(Figs 10, 11 I)	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	description	Description of female (hitherto unknown). Body length (from head to apical margin of tergum II) 5 mm (total body length about 6.5 mm); forewing length 4 mm. Head in frontal view about 1.2 × as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus with concave disc, about 1.25 × as wide as long; its apical emargination rather shallow, about 0.2 × as deep as wide, taking about 0.55 of clypeal width, apical teeth narrowly rounded and lamellate. Distance between lateral ocellus and occiput about 1.15 × as distance between lateral ocelli and about as distance between lateral ocellus and compound eye. Antenna with articles 8 – 12 forming club about 2.8 × as long as wide in dorsal view. Width of gena in front of ocular sinus nearly about as width of compound eye at ocular sinus. Occipital carina complete and strongly lamellate, forming blunt angle in front of ocular sinus. Pronotum with anterior side roundly elevated to dorsal surface. Antero-lateral sides of pronotum broadly rounded in dorsal view; in lateral view, pronotal carina weakly lamellate, distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, slightly convex, broadly rounded posteriorly. Tegula with broadly rounded outer margin produced posterior to blunt medio-posterior angle. Epipleural and episternal sulci weak. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal carina of propodeum complete, rounded dorsally; lateral carina of propodeum complete, ventro-laterally produced as nearly right rounded angle. Terga I – V and sterna II – V with transculent apical lamella. Disc of clypeus shining, sparsely and coarsely punctate, interspaces much exceeding puncture diameter on median line, almost smooth; punctation becoming denser laterally, where interspaces less than puncture diameter. Punctures on frons as coarse as those on disc of clypeus, interspaces usually exceeding puncture diameter, shining but with very weakly visible reticulate microsculpture. Sculpture becoming denser and duller on vertex behind ocelli where interspaces much narrower than puncture diameter. Gena shining, with fine and very sparse punctures and indistinct microsculpture. Pronotum punctate coarser than clypeus and frons, interspaces usually exceeding puncture diameter, shining, almost smooth. Punctures on scutum and scutellum as coarse as those on pronotum but interspaces much larger, several times exceeding puncture diameter, also shining and almost smooth. Tegula with weak reticulate microsculpture and few fine punctures on posterior half. Dorsal and ventral mesepisterna and mesepimeron sculptured much denser than pronotum, interspaces less than puncture diameter, with weak microsculpture. Posterior surface of metanotum shining, with smoothened micropunctures. Metapleuron and lateral surface of propodeum rather dull, longitudinally wrinkled. Dorso-lateral surface of propodeum sculptured similarly to pronotum. Posterior surface of propodeum densely punctate and longitudinally wrinkled, interspaces less than puncture diameter, shining. Metasomal terga rather dull, with reticulate microsculpture and shallow dense punctures; both punctures and interspaces increasing in size towards apical margins of terga; largest punctures on terga much finer than those on scutum and scutellum. Metasomal sterna I – V regularly punctuate with moderately coarse shallow punctures, interspaces about as puncture diameter, rather shining but with distinct reticulate microsculpture. Punctures on sternum VI much coarser than those on preceding sterna, interspaces also about as puncture diameter. Mandible and labrum covered with thin erect pale setae about as long as diameter of median ocellus. Clypeus, frons, vertex, and pronotum with short pale setae about three times shorter than diameter of median ocellus. Compound eye bare. Scutum, scutellum, and mesopleuron without visible setae. Propodeum, metasoma, and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with short erect stiff setae approximately as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Basal color black. Following parts pale yellow outlined by ferruginous: anterior band on pronotum, tegula, crescent spot on scutellum, thin apical band on scutellum, small spots on dorsal and lateral carinae of propodeum, apical bands on terga I – VI, legs from tops of femora. Mandible, labrum, metasomal sterna, and legs until tops of femora reddish-brown to brown. Antenna reddish-brown except dark brown dorsal side of club. Wings transparent; veins and pretostigma brown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	materials_examined	Material examined. LECTOTYPE (designated by Fateryga (2025: 233 )): ♂, labeled “ [golden disc] // Аман-Кутан, Самарк [Aman-Kutan, Samarkand] / р., [district?] 1500 м. у. м. [m a. s. l.] 4 - VII / В Гуссаков. [V. Gussakovskij] 1932 // Quartinia ♂ / uzbeka, Typ / G. Kostylev det. 1933 // Quartinia / uzbeka Kost / Тип. [Type] ♂ / 23 VI 41 [Kostylev’s signature] // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Lectotype ♂ / Quartinia uzbeka / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS _ HYM _ 0000380 [pale yellow label] ” [ZISP]. PARALECTOTYPES: UZBEKISTAN: Samarkand, Aman-Kutan, 1500 m, 4. VII. 1932, 2 ♂, leg. V. Gussakovskij [ZMMU]. ADDITIONAL MATERIAL: TURKMENISTAN: 5 km N Kushka, 26. V. 1985, 1 ♀, 1 ♂, leg. V. Makarkin [FSCV]. UZBEKISTAN: Samarkand, Aman-Kutan, 1500 m, 21. VII. 1931, 1 ♂, leg. V. Gussakovskij [ZISP].	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	distribution	Distribution. Turkmenistan, Uzbekistan.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	biology_ecology	Trophic relationships. Unknown.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
D70F637FFFDB6D5D1DE257E701B027B4.taxon	discussion	Remarks. After the publication of my paper (Fateryga 2025), curators of both ZISP and ZMMU decided to return the lectotype of this species from ZMMU to ZISP, since ZISP was indicated as the depository in the original publication by Kostylev (1935), while the lectotype was actually deposited in ZMMU until present.	en	Fateryga, Alexander V. (2025): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution I. Genera Masaris Fabricius, 1793 and Quartinia André, 1884. Zootaxa 5659 (2): 151-178, DOI: 10.11646/zootaxa.5659.2.1, URL: https://doi.org/10.11646/zootaxa.5659.2.1
