taxonID	type	description	language	source
DB749D3BB71AFF9DFF4B60525B0EFA75.taxon	type_taxon	Type species: Aranea pigra Walckenaer, 1802	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF9DFF4B60525B0EFA75.taxon	diagnosis	Diagnosis. Species of Tmarus can be separated from other Thomisidae by a dorsoposterior tubercle or hump on the opisthosoma, which is more prominent in females (Figs 3 C, 6 C), an elevated ocular area, relatively wide clypeus gradually sloping towards the front (Ileperuma Arachchi & Benjamin 2019).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF9DFF4B60525B0EFA75.taxon	description	Description. Carapace with long setae; tubercles of lateral eyes developed and close to each other, lateral eyes larger than median eyes; abdomen pyriform, the posterior end occasionally extending beyond spinnerets; bulb simple and without apophysis, embolus commonly short and thick; epigynum with a broad epigynal hood or a vertical epigynal ridge, copulatory ducts short and thick, spermathecae small, global, oval or reniform (Ono 1988, Yang et al. 2005, Sarkar et al. 2022).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	description	Figures 1 – 3, 7	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	materials_examined	Type material. Holotype: ♂, CHINA: Hunan Province: Changsha City: Yuelu Mountain, 28.113290 ° N, 112.555391 ° E, 291 m, 26 June 2018 (YLS- 18 - 0606). Paratypes: 1 ♀, same locality, 28.114580 ° N, 112.562274 ° E, 251 m, 26 July 2024 (YLS- 24, matured on 15 December 2024); 1 ♀, Yuelu Mountain, 28.111247 ° N, 112.562897 ° E, 262 m, 13 July 2017 (YLS- 17 - 0705); 5 ♂ 6 ♀, Yuelu Mountain, 28.111247 ° N, 112.562897 ° E, 262 m, 8 July 2018 (YLS- 18 - 0702); 1 ♂ 4 ♀, Yuelu Mountain, 28.113193 ° N, 112.564248 ° E, 149 m, 6 July 2018 (YLS- 18 - 0701); 10 ♂ 2 ♀, Yuelu Mountain, 28.113193 ° N, 112.564248 ° E, 149 m, 15 July 2018 (YLS- 18 - 0704); 1 ♂, Yuelu Mountain, 28.111256 ° N, 112.561998 ° E, 235 m, 10 August 2018 (YLS- 18 - 0803); 2 ♀, Taozi Lake, 28.105490 ° N, 112.575310 ° E, 50 m, 13 August 2018 (YLS- 18 - 0805); 1 ♀, Yuelu Mountain, 28.118760 ° N, 112.562336 ° E, 232 m, 20 August 2018 (YLS- 18 - 0806); 1 ♀, Yuelu Mountain, 28.118760 ° N, 112.562336 ° E, 232 m, 14 September 2018 (YLS- 18 - 0904). Specimen (YLS- 24) was collected by Yuchen Zhou, Gang Tang, Shulin Wang and Wenlong Yan, other specimens were collected by Guchun Zhou and Xuanwei Zhou.	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	etymology	Etymology. The specific name is an adjective related to the type locality	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	diagnosis	Diagnosis. Males of this new species are characterized by their slender embolus and a horned RTA. They can be distinguished from the similar T. taibaiensis Song & Wang, 1994 (see Zhang & Zhang 2023: 85, figs 43 – 50) by: (1) RTA tips fused (Figs 2 B-C) (RTA deeply bifurcated in T. taibaiensis); (2) cymbium longer than wide in ventral view (Fig. 2 C) (vs. as wide as long in T. taibaiensis); (3) embolus originating at 11 o’clock position of tegulum in ventral view (Fig. 2 C) (vs. 12 o’clock in T. taibaiensis); (4) embolus about half as long as tegulum in retrolateral view (Fig. 2 B) (vs. 2 / 3 as long in T. taibaiensis). Females are characterized by short copulatory ducts, peanut-shaped spermathecae, absence of a distinct hood and the epigynal plate being longer than wide. In the similar T. taibaiensis the epigynal plate is as long as wide and the copulatory openings are located anteriorly (versus medially in T. yueluensis sp. nov.). The females of this new species also resemble those of T. piger (Walckenaer, 1802) (see Song & Zhu 1997: 51, fig. 28 A – D; Baba & Tanikawa 2015: 92), but can be easily distinguished by copulatory openings facing anteromedially and located medially (Fig. 3 D) (vs. facing anterolaterally and located posteriorly in T. piger).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	description	Description. Male (holotype) (Figs 1 A – C): Total length 5.00; carapace 2.06 long, 2.01 wide; abdomen 2.93 long, 1.58 wide. Carapace almost as long as wide, yellowish brown, with a longitudinal white band medially, irregular lighter stripes laterally and symmetric setae; cervical and radial grooves visible, fovea unconspicuous. Eye tubercles grayish white, AER and PER recurved in dorsal view. Eye sizes and interdistances: AME 0.07, ALE 0.16, PME 0.10, PLE 0.19, AME – AME 0.20, AME – ALE 0.16, PME – PME 0.25, PME – PLE 0.38. MOA length 0.31 with front width 0.34 and back width 0.43. Clypeus height 0.38. Legs yellow. Leg measurements: I, 14.16 (4.24, 4.86, 3.51, 1.55); II, 14.20 (4.24, 4.88, 3.50, 1.58); III, 5.90 (2.02, 2.22, 1.02, 0.64); IV, 5.83 (2.19, 2.04, 1.02, 0.58). Leg formula: 2134. Abdomen longer than wide, widest at the posterior 1 / 3 part, dorsum yellowish white, with four pairs of transverse light stripes and symmetric setae, posterior part wrinkled, caudal protuberance inconspicuous; venter yellowish white, with a longitudinal brown band medially. Spinneret dark brown. Palp (Figs 2 A – E). Tibia with two apophyses: VTA thumb-shaped with blunt tip. RTA stronger, halberd-shaped, basal part almost 1 / 2 width of tibia, terminal part bifurcated, densely covered with tiny teeth. Tegulum oval and tabular. Embolus slender with pointed tip, originates at 11 o’clock position of tegulum and extended downwards, terminal end curved ventrally in retrolateral view. Female (paratype YLS- 24) (Figs 3 A – C): Total length 4.69; carapace 1.98 long, 1.76 wide; abdomen 2.75 long, 1.74 wide. Eye sizes and interdistances: AME 0.11, ALE 0.18, PME 0.10, PLE 0.17. AME – AME 0.19, AME – ALE 0.15, PME – PME 0.26, PME – PLE 0.40. MOA length 0.47 with front width 0.40 and back width 0.47. Clypeus height 0.30. Leg measurements: I, 7.80 (2.26, 2.82, 1.73, 0.99); II, 8.68 (2.64, 3.20, 1.86, 0.98); III, 4.59 (1.62, 1.76, 0.77, 0.44); IV, 5.09 (1.90, 1.55, 0.96, 0.68). Leg formula: 2143. Coloration pattern similar as in male. Epigyne (Figs 3 D, E). Epigynal plate longer than wide; copulatory openings located in the median-lateral sides of epigyne; copulatory ducts thick and short; spermatheca peanut-shaped; fertilization ducts short.	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	distribution	Distribution. Changsha City, Hunan Province, and Sanming City, Fujian Province, China (Fig. 7).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71AFF99FF4B61A75AB7FD89.taxon	discussion	Remarks. Re-examination of the type series of T. longqicus Song & Zhu, 1993 by the type-bearing institution (Institute of Zoology, Chinese Academy of Sciences, Beijing, China) revealed that the allotype males of T. longqicus are identical with males of T. yueluensis sp. nov. However, the holotype female of T. longqicus is clearly distinct from T. yueluensis sp. nov.: epigynal plate wider than long and copulatory openings located anteriorly (versus epigynal plate longer than wide and copulatory openings located medially in T. yueluensis sp. nov.). We have collected both male and female specimens of T. yueluensis sp. nov. two times from the same locality at the same dates. Males matched with females on morphological characteristics (females appeared similar to males, Figs 1, 3), and DNA barcoding data (showing a genetic distance of 0 % between the male and female voucher specimens). We therefore conclude that the allo- and paratype males of T. longqicus were mistakenly paired with the female holotype. The real male of T. longqicus is still unknown.	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	description	Figures 4 – 7	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	materials_examined	Type material. Holotype: ♂, CHINA: Hunan Province: Zhangjiajie City: Suoxiyu Tujia Township, Tianjiawan Village, 29.325725 ° N, 110.607042 ° E, 454 m, 26 May 2017, Guchun Zhou, Tian Tian and Zongguang Huang leg. (HNU-HN-IV- 1710). Paratypes: 1 ♂ 4 ♀, same data as holotype (HNU-HN-IV- 1710). 3 ♀, Suoxiyu Tujia Township, Wenfeng Village, 29.331437 ° N, 110.599815 ° E, 457 m, 4 April 2016, Wang Liu, Ceng Zeng and Tian Tian leg. (HNU-HN-IV- 1601).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	etymology	Etymology. The specific epithet is derived from the Latin “ rostratus ”, meaning beaked / curved, referring to the coracoid shape of the RTA, adjective.	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	diagnosis	Diagnosis. The males of this new species resemble those of T. piger (Walckenaer, 1802) (see Song & Zhu 1997: 51, figs 28 A – D; Baba & Tanikawa 2015: 92) and T. punctatissimus (Simon, 1870) (see Bosmans & Hervé 2015: 38, figs 1 A – G) in having a similar course of sperm duct, embolus and RTA shape, but can be distinguished by: (1) the distal end of the VTA is curved in an arc in ventral view (Fig. 5 C) (vs. broadly round in T. piger and T. punctatissimus); (2) RTA extends upwards, with coracoid apex in dorsal view (Fig. 5 B) (vs. dorsal curved in T. piger and T. punctatissimus); (3) the embolus duck-head-shaped in ventral view (Fig. 5 C) (vs. hook-shaped in T. piger and champagne bottle-like in T. punctatissimus). The females of this new species resemble those of T. piger (Walckenaer, 1802) (see Song & Zhu 1997: 51, fig. 28 A – B; Baba & Tanikawa 2015: 92) in having two posterior copulatory openings and short copulatory ducts, but can be distinguished by: (1) copulatory openings facing anteromedially (Fig. 6 D) (vs. facing anterolaterally in T. piger); (2) spermatheca stomach-shaped and lateral to copulatory ducts (Fig. 6 D) (vs. peanut-shaped and posterior to copulatory ducts in T. piger).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	description	Description. Male (holotype) (Fig. 4): Total length 4.28; carapace 1.88 long, 1.75 wide; abdomen 2.40 long, 1.18 wide. Carapace almost as long as wide, reddish brown, with irregular lighter stripes laterally and symmetric setae; cervical and radial grooves visible, fovea unconspicuous. Eye tubercles grey, AER and PER recurved in dorsal view. Eye sizes and interdistances: AME 0.07, ALE 0.13, PME 0.10, PLE 0.14, AME – AME 0.25, AME – ALE 0.26, PME – PME 0.32, PME – PLE 0.34. MOA length 0.21 with front width 0.16 and back width 0.21. Clypeus height 0.29. Legs yellow. Leg measurements: I, 9.93 (2.81, 3.53, 2.33, 1.26); II, 10.46 (2.87, 3.76, 2.49, 1.34); III, 4.73 (1.34, 2.01, 0.79, 0.59); IV, 4.94 (1.66, 1.87, 0.78, 0.63). Leg formula: 2143. Abdomen longer than wide, dorsum yellowish white, mottled with brown patches and symmetric setae, posterior part wrinkled, caudal protuberance inconspicuous; venter with a wide longitudinal greyish yellow band medially and a pair of thin longitudinal white band laterally. Spinneret yellowish brown. Palp (Fig. 5). Tibia with two apophyses: VTA hook-shaped with blunt tip, curved prolaterally; RTA longer and thicker than VTA, with pointed and curved tip. Tegulum oval and tabular. Embolus thicker proximally and slender distally, originates at 1 o’clock position of tegulum. Female (one paratype of HNU-HN-IV- 1710) (Figs 6 A – C): Total length 6.92; carapace 2.36 long, 2.23 wide; abdomen 4.52 long, 2.62 wide. Eye sizes and interdistances: AME 0.09, ALE 0.15, PME 0.16, PLE 0.14. AME – AME 0.31, AME – ALE 0.35, PME – PME 0.42, PME – PLE 0.50. MOA length 0.24 with front width 0.22 and back width 0.26. Clypeus height 0.33. Leg measurements: I, 8.54 (2.29, 3.78, 1.61, 0.86); II, 8.66 (2.41, 3.83, 1.69, 0.73); III, 5.03 (1.26, 2.14, 0.86, 0.77); IV, 5.21 (1.48, 2.14, 0.83, 0.76). Leg formula: 2143. Otherwise colour and patterns same as in male. Epigyne (Figs 6 D, E). Epigynal plate slightly wider than long; copulatory openings located posteriorly; copulatory ducts thick and short; spermatheca stomach-shaped; fertilization ducts short.	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
DB749D3BB71EFF94FF4B66FB5A76FC4D.taxon	distribution	Distribution. Known only from the type locality (Fig. 7).	en	Zhou, Yu-Chen, Qin, Xin-Yu, Zhou, Gu-Chun, Peng, Xian-Jin, Liu, Ping (2025): Two new species of the genus Tmarus Simon, 1875 (Araneae: Thomisidae) from China. Zootaxa 5666 (1): 105-114, DOI: 10.11646/zootaxa.5666.1.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
