taxonID	type	description	language	source
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	description	(Figs 1 – 5, 17, 18, 21)	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	materials_examined	Type material. Holotype (Figs 1, 17): male, “ 15 – 18. III. 2011, | SE Kazakhstan, | Altyn-Emel Range, | Sholak Mts, 750 m, | 43 ˚ 54 ' 47.04 '' N | 77 ˚ 47 ' 16.45 '' E | local collector leg. ” / “ Slide | AV 9145 ♂ | A. Volynkin ” (CAV). Paratypes. KAZAKHSTAN: 51 males, same data as in holotype (CAV); 2 females, 26. III. 2011, Toraygyr Mts, 1380 m, 43 ° 19 ' 7.3 '' N 78 ° 33 ' 45.4 '' E, local collector leg. (CAV); 1 female, 24. III. 2011, Arkharly Pass, 1115 m, 44 ° 14 ' 20.6 '' N 77 ° 43 ' 16.1 '' E, local collector leg. (CAV); 2 females, 5. IV. 2009, foothills of Syrdarya Karatau Mts, 20 km WSW of Cholakkurgan, 741 m, 43 ° 42 ' 47.69 '' N 68 ° 54 ' 36.95 '' E, local collector leg. (CAV); 1 male, 1 female, Ustyurt Plateau, 19 km N of Beineu town, 45 ˚ 30 ' N 55 ˚ 15 ' E, 4 – 5. IV. 2013, P. Gorbunov [leg.] (MDS); 10 males, 19. III. 2024, SE Kazakhstan, Zhetysu Region, Sholak Mts, 750 m, 43 ° 54 ' 57.6 '' N, 77 ° 47 ' 47.8 '' E, S. V. Titov leg. (STP); KYRGYZSTAN: 1 male, 2 females, Moldo-Too Mt. Range, 15 km NW of Kazarman, Naryn River vall., 41 ˚ 30 ' 46 '' N 73 ˚ 55 ' 38 '' E, 1220 m, 21 – 22. IV. 2023, Pavel Gorbunov leg. (BBM); UZBEKISTAN: 1 male, Tashkent Reg., Korzhin-Tau Range, Aktash, 31. III. 1962, V. Ganson leg., gen. [in a vial] No. 1364, ex coll. A. V. Nekrasov, gen. slide No.: AV 9141 ♂ (ZISP).	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	diagnosis	Diagnosis. Perigrapha belyalovi sp. n. (Figs 1 – 5) is externally reminiscent of the sympatric P. centralasiae (Figs 9 – 16) and the Tajikistani P. heidi (Figs 6 – 8) and can be distinguished from the former species by the larger size and somewhat longer rami of the male antenna. Compared to P. heidi, the new species has a somewhat less tapered forewing apex. In the male genitalia, P. belyalovi sp. n. (Figs 17, 18) differs clearly from both similar congeners (Figs 19 – 24) in the shorter and broader uncus and the markedly broader cucullus with a larger and apically pointed dorsal lobe in the genital capsule, and the longer cluster consisting of more robust spine-like cornuti in the vesica. Additionally, in the new species, the ventral lobes of the cuculli are shorter and less asymmetrical than in P. centralasiae, and more asymmetrical than in P. heidi and apically pointed, whereas in the latter they are apically rounded. In the female genitalia, P. belyalovi sp. n. (Fig. 25) is distinct from P. centralasiae (Fig. 27) in the somewhat longer ductus bursae with a twisted anterior end (it is even in the congener), the larger corpus bursae with a larger right lateral signum, the longer and reniform posterior sclerotised plate of the corpus bursae (it is trapezoidal in P. centralasiae), and the longer appendix bursae with a broader distal section and a more heavily sclerotised basal rugose cluster. Compared to P. heidi (Fig. 26), the new species has a broader ductus bursae, a markedly broader posterior sclerotised plate of the corpus bursae, a rounded lateral right signum (it is elliptical in P. heidi), and a considerably broader and more heavily gelatinous appendix bursae with a sclerotised lateral wall of its distal dilated section.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	description	Description. External morphology of adults (Figs 1 – 5). Forewing length 16.5 – 19.0 mm in males and 18.0 – 19.0 mm in females. Antenna bipectinate in male and filiform in female. Body reddishbrown with admixture of pale ochreous brown. Forewing with elongate apex and oblique tornus. Forewing ground colour pale ochreous brown with suffusion of dark brown; medial area darker, reddishbrown. Subbasal line dark brown, sinuous, diffuse. Antemedial line double, straight medially and zigzagged inwards posteriorly, stretching obliquely. Orbicular, suborbicular and reniform markings large, pale ochreous brown with slight brown suffusion, fused with pale ochreous brown Cu vein, encircled by blackish suffusion. Postmedial line double and thin, question mark-like curved, slightly dentate between veins. Subterminal line pale ochreous, thin, irregular. Terminal line black, interrupted into small spots between veins. Forewing cilia ochreous brown with black speckles between vein tips. Hindwing dark grey with dark ochreous brown suffusion along veins. Hindwing cilia pale ochreous brown. Male genitalia (Figs 17, 18). Uncus short, dorso-ventrally flattened, somewhat distally dilated, apically rounded. Tegumen arms narrow and moderately sclerotised. Vinculum as long as tegumen, heavily sclerotised, V-shaped but with rounded ventral end. Valva distally tapered and downcurved. Dorsal margin strongly convex antemedially, while it’s medial and postmedial sections almost straight. Cucullus broad, trapezoidal, with triangular and setose dorsal lobe and narrower, falcate and upcurved ventral lobe. Ampulla slender, digitiform, almost straight, directed distally and somewhat protruding beyond ventral margin of valva. Clasper falcate, with dilated dorsal section bearing short, tubercle-like ampulla. Sacculus proximally almost as broad as valva, distally tapered and reaching clasper. Juxta shield-shaped with elongate dorsal section. Anellus setose. Phallus long, cylindrical, with short and rounded coecum, and short ribbon-like ventral carina protruding into base of vesica. Vesica tubular, proximally straight and distally helicoid, with broad but short diverticulum bearing elongate cluster of numerous spine-like cornuti. Female genitalia (Fig. 25). Ovipositor short and conical. Papilla analis short, trapezoidal, weakly setose. Apophyses thin and elongate, anterior one somewhat longer and thicker than posterior one. Ostium bursae broad. Ductus bursae elongate and heavily sclerotised, dorso-ventrally flattened, its posterior half posteriorly dilated while anterior one somewhat anteriorly tapered and twisted. Corpus bursae pyriform, its posterior narrow end rugose and with longitudinal reniform sclerotised plate stretching perpendicularly the appendix bursae base. Anterior section of corpus bursae membranous and bearing four small, elliptical signa of different sizes. Appendix bursae arising posterolaterally on left side, elongate and helicoid, gelatinous, with two longitudinal sclerotised rugose clusters basally and medially, its distal section dilated and heavily gelatinous with weak sclerotisation laterally.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	etymology	Etymology. The new species is dedicated to the memory of the late Oleg V. Belyalov (Almaty, Kazakhstan), close friend of Sergey Titov, excellent wildlife photographer, documentary filmmaker and cameraman, botanist and zoologist, and populariser of the knowledge of the wildlife of Kazakhstan.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C115FFB873BAF99FFF58FEF6.taxon	distribution	Distribution and bionomics. The new species is currently known from Southeast and South Kazakhstan, Uzbekistan and Kyrgyzstan. In Southeast Kazakhstan, P. belyalovi sp. n. is found occurring sympatrically with P. centralasiae in low, dry mountains with stony steppe slopes (Fig. 28) dominated by Artemisia L. spp. and with scarce low shrubs of Spiraea hypericifolia L., Rosa platyacantha Schrenk, R. alberti Regel, Lonicera microphylla Roem. & Schult., Caragana balchaschensis (Kom.) Pojark., Cerasus tianshanica Pojark., Salix songarica Andersson, S. tenuijulis Ledeb., and S. wilhelmsiana M. Bieb. (Goloskokov 1984). It was not found in more flat landscapes such as Balkhash-Alakol Depression, where P. centralasiae also occurs (e. g., in Tamgaly Tas rock massif NE from Konaev City) (Figs 11, 12).	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB573BAFF47FDB0FB71.taxon	description	(Figs 6 – 8, 19, 26)	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB573BAFF47FDB0FB71.taxon	materials_examined	Material examined. Paratype: male, USSR, Tajikistan, 39 ° N 68 ° 40 ' E, Hissar Mts, Romit, 1100 m, 21. IV. 1991, U. Jūrivete leg., gen. slide No.: 4062 (prepared by Hreblay) (MH / HNHM). Non-type: 2 males, 1 female, Tajikistan, Hissar Mts, Varzob Range, Takob valley at Poshum village, 1700 m, 38 ° 52 ' 20 '' N 69 ° 55 ' 47 '' E, 11 – 12. IV. 2015, B. Benedek leg., gen. slide No.: JB 2377 ♀ (prepared by Babics) (BBM).	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB573BAFF47FDB0FB71.taxon	diagnosis	Diagnosis. The forewing length is 18.0 – 19.0 mm in both sexes. Perigrapha heidi is morphologically most similar to P. belyalovi sp. n. and the detailed comparison is provided above in the diagnosis of the latter species. It is also largely reminiscent of P. centralasiae but in most cases can be distinguished by its somewhat larger size and slightly longer rami of male antenna. However, as in Tajikistan some specimens of P. centralasiae are as large as P. heidi, the reliable identification requires examination of the genitalia structures. In the male genital capsule, P. heidi differs from P. centralasiae in the more distally dilated uncus, the longer and somewhat narrower ‘ neck’ of the cucullus, the more symmetrical cuculli with shorter, distally broader, and apically rounded ventral lobes (they are apically pointed in P. centralasiae), and the somewhat larger left dorsal lobe of the cucullus. The phalli of two species are alike. The vesica of P. heidi is somewhat longer and more helicoid than in P. centralasiae. In the female genitalia, P. heidi is distinct from P. centralasiae in the longer and posteriorly narrower ductus bursae, the markedly larger corpus bursae, the considerably narrower posterior sclerotised plate of the corpus bursae, and the somewhat longer and distally narrower appendix bursae. Additionally, the lateral right signum of P. heidi is elliptical and large whereas in is very small and rounded in P. centralasiae.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB573BAFF47FDB0FB71.taxon	distribution	Distribution. The species is currently known only from Hissar Range in Tajikistan (Hreblay 1996; Benedek et al. 2021).	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB773BAFB52FC96FA69.taxon	description	(Figs 9 – 16, 20 – 24, 27)	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB773BAFB52FC96FA69.taxon	materials_examined	Type material examined. Photographs of the holotype (Fig. 21): male, “ Asia centr. | Aschckab [ad] | 1898 ” / pink label “ Type | v. centralasiae Bart. | ♂ ” / red black- framed label “ Type ” / “ Zool. Mus. | Berlin ” / “ Genit. praep. | Kononenko, 1991 ” / “ Perigrapha centralasiae | Bartel, 1906 | Holotypus, [slide] NO: 4067 | det. M. Hreblay ” (MfN); photographs of the holotype of duktana (Fig. 22): male, [Tajikistan or Uzbekistan, Zeravshan Range] “ Duktan | Sarafschan | Juni ” / pale brick red label “ Type | Perigrapha | duktana | ♂ Draudt. ” / “ 129. ” / “ P. centralasiae Bart. | = ductana Drdt | syn. n. | Kononenko, 1991 ” / “ Genit. praep. | Kononenko, 1991 ” / “ Perigrapha duktana | Draudt, 1934 | Holotypus | [slide] NO: 4066 | det. M. Hreblay ” (MfN). Additional material examined: series of both sexes from Turkmenistan, Tajikistan, Kyrgyzstan and South and Southeast Kazakhstan (HNHM, BBM, CAV, STP, ZISP).	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB773BAFB52FC96FA69.taxon	discussion	Remark. The species is highly polymorphic in its forewing colouration and size. The populations of P. centralasiae from Hissar Range in Tajikistan (Figs 15, 16, 20), where P. heidi is also found, are larger, equal in size to the latter species and therefore the reliable identification requires an examination of the genitalia structures.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB773BAFB52FC96FA69.taxon	distribution	Distribution. The species is widespread in Central Asia and currently known from Turkmenistan, Tajikistan, Kyrgyzstan, eastern Uzbekistan, and southern Kazakhstan. Acknowledgements Sergey Titov expresses his sincere thanks to Alexey Matov (ZISP) for his kind assistance during the work at ZISP collection, and Andrey Shapovalov (Institute of Zoology, Almaty, Kazakhstan) for organising the collecting trip in Southeast Kazakhstan in the early spring of 2024. The authors are indebted to Balázs Benedek (Mohács, Hungary) for a valuable discussion, data from his rich research collection and photos of the type specimens of Perigrapha centralasiae provided. The study of Sergey Titov and Ruslan Rakhimov was funded by Science Committee of the Ministry of Education and Science of the Republic of Kazakhstan (Program No. BR 21882199, “ Inventory of wild animals of arid territories of the Balkhash- Alakol basin with an assessment of threats to their conservation and sustainable use. ”). Anton Volynkin and Balázs Tóth, while performing this work, received no specific grant from any funding agencies.	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
DA6C8799C11CFFB773BAFB52FC96FA69.taxon	description	Benedek, B., Babics, J. & Bálint, Zs. (2021) Faunistical records and annotations for a better knowledge of the Tajikistani moth and butterfly fauna (Lepidoptera: Noctuoidea, Papilionoidea). Caucasian entomological bulletin, 17 (1), 61 – 76. http: // dx. doi. org / 10.23885 / 181433262021171 - 6176 Benedek, B. & Ronkay, L. (2001) Two new Noctuidae species from Iran (Lepidoptera). Folia Entomologica Hungarica, 62, 191 – 198. [http: // publication. nhmus. hu / pdf / folentom / FoliaEntHung _ 2001 _ Vol _ 62 _ 191. pdf] Draudt, M. (1931 – 1938) Noctuidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Supplement 3. Alfred Kernen, Stuttgart, pp. 1 – 337, pls. 1 – 26. [In German] [https: // www. biodiversitylibrary. org / page / 56099446] Goloskokov, V. P. (1984) Flora of the Dzhungar Alatau. Nauka, Alma-Ata, 224 pp. [In Russian] Hacker, H. (1996) Ergänzungen zu “ Die Noctuidae Vorderasiens ” und neuere Forschungsergebnisse zur Fauna der Türkei II (Lepidoptera). Esperiana, 4, 273 – 330. [In German] Hreblay, M. (1996) Neue paläarktische Taxa aus der Gattung Perigrapha Lederer, 1857 (Lepidoptera, Noctuidae). Esperiana, 4, 65 – 94. [In German] Hreblay, M. & Ronkay, L. (1997) New Noctuidae (Lepidoptera) species from Taiwan and the adjacent areas. Acta Zoologica Academiae Scientiarum Hungaricae, 43 (1), 21 – 83. Hreblay, M. & Ronkay, L. (1999) Neue trifide Noctuidae aus dem himalayanischen Raum und der südostasiatischen Region (Lepidoptera: Noctuidae). Esperiana, 7, 485 – 620. [In German] Kononenko, V. S. (2010) Micronoctuidae, Noctuidae: Rivulinae – Agaristinae (Lepidoptera). Noctuidae Sibiricae. Vol. 2. Entomological Press, Sorø, 475 pp. Korb, S. K. (2022) Perigrapha heidi Hreblay, 1996, a little-known species new to the faunas of Kyrgyzstan and Kazakhstan, with molecular-genetic evidence of its species status (Lepidoptera: Noctuidae). Russian Entomological Journal, 31 (1), 71 – 75. https: // dx. doi. org / 10.15298 / rusentj. 31.1.14 Lafontaine, J. D. & Mikkola, K. (1987) Lock-and-key system in the inner genitalia of Noctuidae (Lepidoptera) as taxonomic character. Entomologiske Meddelelser, 55, 161 – 167. [In Swedish with English abstract] Ronkay, L., Yela, J. L. & Hreblay, M. (2001) Hadeninae II. Noctuidae Europaeae, 5. Entomological Press, Sorø, 452 pp. Ronkay, G., Ronkay, L., Gyulai, P. & Hacker, H. (2010) New Orthosiini (Lepidoptera, Noctuidae, Hadeninae) species and genera from the wide sense Himalayan region. Esperiana, 15, 127 – 221. Saldaitis, A., Ivinskis, P. & Borth, R. (2012) Two new Perigrapha species from China (Lepidoptera, Noctuidae). Zootaxa, 3426, 64 – 68. https: // doi. org / 10.11646 / zootaxa. 3426.1.4 Volynkin, A. V. (2024) On the terminology of the genitalia structures of lichen moths (Lepidoptera: Erebidae: Arctiinae: Lithosiini) with some references to Noctuidae. Ecologica Montenegrina, 73, 176 – 207. https: // dx. doi. org / 10.37828 / em. 2024.73.18 Volynkin, A. V. & Matov, A. Yu. (2014) A new Perigrapha Lederer, 1857 from Western China (Lepidoptera: Noctuidae, Hadeninae). Journal of Asia-Pacific Entomology, 17, 609 – 612. http: // dx. doi. org / 10.1016 / j. aspen. 2014.05.011 Volynkin, A. V., Titov, S. V. & Knyazev, S. A. (2014) A new Perigrapha Lederer, 1857 from South Kazakhstan (Lepidoptera, Noctuidae). Zootaxa, 3856 (2), 292 – 296. http: // dx. doi. org / 10.11646 / zootaxa. 3856.2.8	en	Titov, Sergey V., Volynkin, Anton V., Tóth, Balázs, Rakhimov, Ruslan D. (2025): Perigrapha belyalovi, a new species from Central Asia (Lepidoptera: Noctuidae: Noctuinae). Ecologica Montenegrina 83: 1-11, DOI: 10.37828/em.2025.83.1, URL: https://doi.org/10.37828/em.2025.83.1
