identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
DA3E8786F50616532DCCF976FDB983F9.text	DA3E8786F50616532DCCF976FDB983F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheleocloeon clavifolium Kluge 2016	<div><p>Cheleocloeon clavifolium Kluge 2016</p><p>Centroptilum clavifolium Kluge 2016: 149 (larva, subimago, ♂ and ♀ imago, eggs).</p><p>Material. ZAMBIA: Holotype and paratypes (see Kluge 2016). TANZANIA: Morogoro Region, river Molukwi ( right tributary of river Mkondoa upstream Kilosa), coll. N. Kluge &amp; L. Sheyko, 18.VIII.2017: 2 L-S-I♂, 1 L-S♂, 2 L-S-I♀, 3 larvae.</p><p>Distribution. Tanzania and Zambia.</p></div>	https://treatment.plazi.org/id/DA3E8786F50616532DCCF976FDB983F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2025): Two new species of the genus Cheleocloeon Wuillot & Gillies 1993 (Ephemeroptera: Baetidae) from Tanzania and corrections to the generic diagnosis. Zootaxa 5660 (3): 330-350, DOI: 10.11646/zootaxa.5660.3.2, URL: https://doi.org/10.11646/zootaxa.5660.3.2
DA3E8786F50616532DCCFB1AFC70825B.text	DA3E8786F50616532DCCFB1AFC70825B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheleocloeon Wuillot & Gillies 1993	<div><p>Genus Cheleocloeon Wuillot &amp; Gillies 1993</p><p>(Figs 1–82)</p><p>Type species: Cheleocloeon yolandae Wuillot (in Wuillot &amp; Gillies) 1993.</p><p>Diagnosis of Cheleocloeon was given in the previous paper (Kluge 2016). Characters of the new species Ch. psammonella sp. nov., partly disagree with this diagnosis, so the generic diagnosis should be modified as following.</p><p>Point (3): Labrum. Distal margin of labrum varies from usual for most Baetidae (Fig. 8) to concave (Fig. 56).</p><p>Point (5): Maxillae. Maxillary canines vary from long and slender (Fig. 11) to fused forming short, flat, wide and blunt plate (Figs 62–63).</p><p>Point (7): Labium. Glossae and paraglossae vary from pointed, both with a regular dorso-lateral row of long setae (Figs 12–13) to blunt, with numerous irregularly situated long setae on apex of paraglossa (Fig. 64).</p><p>Points (1)–(2), (4), (6), (8)–(24) of the diagnosis are unchanged.</p></div>	https://treatment.plazi.org/id/DA3E8786F50616532DCCFB1AFC70825B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2025): Two new species of the genus Cheleocloeon Wuillot & Gillies 1993 (Ephemeroptera: Baetidae) from Tanzania and corrections to the generic diagnosis. Zootaxa 5660 (3): 330-350, DOI: 10.11646/zootaxa.5660.3.2, URL: https://doi.org/10.11646/zootaxa.5660.3.2
DA3E8786F50516592DCCFF27FF06827F.text	DA3E8786F50516592DCCFF27FF06827F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheleocloeon acutipalpum Kluge 2025	<div><p>Cheleocloeon acutipalpum sp. nov.</p><p>(Figs 1–49)</p><p>Material. Holotype: L-S-I ♂ {specimen [XXXVIII] (15) 2016} TANZANIA, Tanga Region, Amani, Usambara Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=38.633335&amp;materialsCitation.latitude=-5.1" title="Search Plazi for locations around (long 38.633335/lat -5.1)">river Sigi</a>, 5°06’S, 38°38’E, 1.IX.2016, coll. N. Kluge &amp; L. Sheyko. Paratypes: TANZANIA: the same locality and collectors, 19.VIII–2.IX.2016: 1 L-S-I ♂, 1 L-S ♂, 11 larvae; Mbeya Region, Matema, Livingstone Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.033333&amp;materialsCitation.latitude=-9.483334" title="Search Plazi for locations around (long 34.033333/lat -9.483334)">river Mwalalo</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.033333&amp;materialsCitation.latitude=-9.483334" title="Search Plazi for locations around (long 34.033333/lat -9.483334)">tributary of Lake Nyasa</a>), 9°29′S, 34°02′E, 6–14.VIII.2016, coll. N. Kluge &amp; L. Sheyko: 4 L-S-I♂, 2 L-S♂, 5 L-S-I♀, 2 L-S♀, 7 larvae ; Morogoro Region, Uluguru Mountains: river Morogoro in Morogoro, 31.VII.2017, coll. N. Kluge &amp; L. Sheyko: 4 larvae ; basin of river Ruvu near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.75&amp;materialsCitation.latitude=-6.9" title="Search Plazi for locations around (long 37.75/lat -6.9)">Kinole</a>, 6°54′S, 37°45′E, 27–30.VII.2017, coll. N. Kluge &amp; L. Sheyko: 2 L-S-I♀, 2 larvae .</p><p>Etymology. The adjective acutipalpum refers to acute apex of the disto-median projection of 2nd segment of labial palp.</p><p>Descriptions</p><p>Larva. CUTICULAR COLORATION: Cuticle with contrasting ornament of brown and colorless areas. Frons close to frontal suture darkened, vertex darkened with composite blanks corresponding to attachments of mandibularcranial adductors (Fig. 2). Pronotum and mesonotum with composite ornament of brown and colorless areas; cuticle of fore protoptera brown, distally lighter, with darker brown stripes along convex longitudinal veins and colorless stripes along concave longitudinal veins (Figs 3, 6). Metanotum brown with pair of transverse blanks; hind protoptera of male brown, distally lighter (Figs 4–5). Thoracic pleura brown with colorless blanks, thoracic sterna colorless (Fig. 5). Femora colorless, with or without diffusive brown longitudinal macula in distal part; tibiae and tarsi either nearly colorless, or diffusively darkened on outer side, or brown with colorless patella-tibial suture (Figs 5, 15–17). Abdominal terga with composite ornament of brown and colorless areas; terga I, III, VI and IX darker than neighboring ones (Figs 1, 7). Sterna I–VIII mostly colorless; in posterior part of abdomen, anterior margin of each sternum more or less darkened with brown; sternum IX darkened with brown. Caudalii often darkest just distad of middle and at apex, with lighter area between these dark areas (Figs 1,7).</p><p>HYPODERMAL COLORATION. Either not expressed, or abdominal terga with reddish or brown median and lateral maculae as in imago (Fig. 23).</p><p>SHAPE AND SETATION: Labrum with anterior margin convex, with median incision (Fig. 8). Mandibles— Figs 9–10. Maxillae non-modified (Fig. 11). Glossae and paraglossae non-modified (Figs 12–13). Disto-median projection of 2nd segment of labial palp (characteristic for Cheleocloeon) sharply projected, apically pointed (Fig. 14).</p><p>Legs slender (Figs 15–17). Claw with pair of longitudinal rows of minute denticles near base, continued by pair of longitudinal rows of few (2–3) wide, very short, truncated, colorless denticles in proximal 1/3 of claw (Fig. 18).</p><p>Male larva with hind protoptera (Fig. 4); female larva with vestiges of hind protoptera rather wide and projected posteriad of posterior margin of metanotum (Fig. 5).</p><p>Abdominal terga I–X with dense scales in wide sockets; narrower and longer scales located near posterior margin, some attached on posterior margin between denticles (Fig. 21). Posterior margin of tergum I with smaller denticles; posterior margin of terga II–X bordered by dark brown band and bear regular row of dark brown pointed denticles (Fig. 21). Posterior margins of abdominal sterna I–IV smooth; posterior margins of sterna V–IX with regular row of denticles, smaller than denticles on terga, without scales between them. Paraproct with pointed denticles (Fig. 20). All tergalii I–VII not wide and widest in distal half; tergalius I narrowest, as long as next ones (Figs 24–30). Lateral side of each cercus and dorsal side of paracercus with several pointed denticles on posterior margin of each 4th segment (Fig. 22). Distal part of cercus with longitudinal row of secondary swimming setae on lateral side (besides primary swimming setae on medial side) (as in Fig. 71).</p><p>Pose of subimaginal gonostyli under larval cuticle. « Cloeon - type » (as in Fig. 79), as in other Cheleocloeon (see Kluge 2016: character 22 on p. 144).</p><p>Subimago. CUTICULAR COLORATION. Head colorless. Pronotum light brown with colorless blanks. Mesonotum very light brownish, nearly colorless, with contrasting dark brown anterior part of lateroparapsidal suture, antelateroparapsidal suture, prelateroscutum and parascutellum (Fig. 46). Thoracic pleura and sterna with contrasting dark brown, light brownish and colorless areas (Fig. 45). Legs colorless. Abdominal terga and sterna light brownish, each tergum with pair of darker sublateral spots. Cerci colorless.</p><p>TEXTURE. All tarsal segments of all legs covered with pointed microlepides.</p><p>Imago, male (Figs 31–37). Head brown. Turbinate eyes nearly cylindrical, slightly widened apically; stem ochre; facetted surface orange, bordered by narrow contrasting dark brown ring. Thorax uniformly brown on dorsal, lateral and ventral sides. Fore leg with femur brown or ochre-brownish, tibia and tarsus lighter; middle and hind legs either uniformly light ochre (Figs 33–34), or with brownish spot near apex of femur (Fig. 37). Middle and hind legs with apical spine of 1st+2nd and 3rd tarsal segments. Fore wing colorless, with veins light ochre, base proximad of costal brace with brown; pterostigma with several simple, oblique, incomplete veins. Hind wing present, narrow, with hooked costal projection (Fig. 35). Abdominal terga from ochre to dark brown, with darker reddish unpaired and paired maculae (Fig. 31–32). Abdominal sterna ochre. Cerci uniformly pale ochre.</p><p>Genital structure as characteristic for Cheleocloeon (see Kluge 2016), with following peculiarities (Figs 38–42): Unpaired sclerite between unistyligers absent. Unistyligers with distal margin widened medially to form a pair of projections directed toward one another (Figs 38, 40, 42). 1st segment of gonostylus basally on median side either with well outlined sclerotized concave area (Figs 38, 40), or without it (Fig. 42).</p><p>Imago, female (Fig. 44). Head and thorax lighter than in male, brownish-ochre. Abdominal terga either uniformly colored, or with reddish maculae as in male. Hind wings absent. Fore leg with apical spines on 2nd and 3rd tarsal segments. Middle and hind legs as in male.</p><p>Egg. Oval; chorion with even and regular net-like relief (Figs 48–49).</p><p>Dimension. Fore wing length (and approximated body length) 5 mm.</p><p>Distribution. Known from Tanzania: Livingstone Mountains (near Matema), Uluguru Mountains, Usambara Mountains.</p><p>Comparison. By the pointed projection of 2nd segment of labial palp (Fig. 14), larva of Ch. acutipalpum sp. nov. resembles Ch. yolandae Wuillot (in Wuillot &amp; Gillies) 1993, Ch. carinatum Wuillot (in Wuillot &amp; Gillies) 1993 and Ch. littorale McCafferty 2001; among them, Ch. yolandae and Ch. carinatum differ from other species of Cheleocloeon by lacking hind wings in both sexes. Larva of Ch. acutipalpum sp. nov. well differs from larvae of Ch. yolandae and Ch. carinatum by tergalius of the 1st pair, which is not longer than others (Fig. 24).</p><p>Ch. littorale was described as larvae from Lake Nyasa, i.e. in the same area as Ch. acutipalpum sp. nov. According to the original description, larva of Ch. littorale has «Claws ... with inner marginal row of 12–14 ... denticles ... extending from near the base of the claw to approximately two-thirds length of claw» (McCafferty 2001: 66 and Fig. 7). In contrast to this, Ch. acutipalpum sp. nov. has very small denticles only on the proximal part of claw (Fig. 18). Projection of 2nd segment of labial palp is pointed also in the undescribed species from Uganda reported as « Cheleocloeon sp. 3 » (Kluge 2016); this species differs from others by unusually stout legs and peculiar shape of right prostheca.</p><p>Male imago of Ch. acutipalpum sp. nov. differs from other species by prominent median projections of unistyligers (Figs 38, 40, 42).</p><p>Comment. A part of material on the new species Ch. acutipalpum sp. nov. was collected in the Upper Sigi River in Usambara Mountains, in the place from which the species originally named Cheleocloeon sigiense Gillies 2001 was described. As shown by Kluge (2016) and Kluge et al. (2018), this species does not belong to Cheleocloeon, but belongs to the genus Susua in Protopatellata. The new species Ch. acutipalpum sp. nov. differs from Susua sigiense (Gillies 2001) by many characters at larval and winged stages, so that M.T. Gillies could not confuse these two species. The species described here as Ch. acutipalpum sp. nov., was not reported under any name in the list of mayfly species from the Upper Sigi River, where Susua sigiense was reported as « Dabulamanzia sp. nov. » (Gillies 1999).</p></div>	https://treatment.plazi.org/id/DA3E8786F50516592DCCFF27FF06827F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2025): Two new species of the genus Cheleocloeon Wuillot & Gillies 1993 (Ephemeroptera: Baetidae) from Tanzania and corrections to the generic diagnosis. Zootaxa 5660 (3): 330-350, DOI: 10.11646/zootaxa.5660.3.2, URL: https://doi.org/10.11646/zootaxa.5660.3.2
DA3E8786F50C16412DCCF912FECF878B.text	DA3E8786F50C16412DCCF912FECF878B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheleocloeon psammonella Kluge 2025	<div><p>Cheleocloeon psammonella sp. nov.</p><p>(Figs 50–82)</p><p>Material. Holotype: L-S/I ♂ {specimen [X] (13) 2017}, TANZANIA, Iringa Region, Uszungwa Mountains, river Msosa (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.533333&amp;materialsCitation.latitude=-7.55" title="Search Plazi for locations around (long 36.533333/lat -7.55)">tributary of Great Ruaha</a>), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.533333&amp;materialsCitation.latitude=-7.55" title="Search Plazi for locations around (long 36.533333/lat -7.55)">Msosa Camp</a> (9 km S Mbuyuni), 7°33′S, 36°32′E, 7.VIII.2017, coll. N. Kluge and L. Sheyko.</p><p>Etymology. The feminine noun psammonella repeats the genus-group name Psammonella Glazaczow (in Jacob &amp; Glazaczow) 1987, which was derived from Greek ψαμμη (sand) referring to the preferred larval substratum. The name refers to the convergent similarity between this species of Cheleocloeon and the subgenus Psammonella, belonging to Procloeon Bengtsson 1915 .</p><p>Descriptions</p><p>Larva. CUTICULAR COLORATION: Cuticle with brown ornament on colorless background. Dorsal eyes (of mature male larva) brown, occipit between them partly brown; other parts of head cuticle colorless (Fig. 52). Pronotum and mesonotum with brown and colorless areas; cuticle of fore protoptera light ocher-brownish with incomplete brown stripes along some convex veins and with colorless stripes along distal parts of concave veins (Fig. 51). Metanotum brown; metapleura partly brown (Fig. 54), other thoracic pleura and sterna colorless. Femora colorless, with diffusive brown longitudinal macula in distal part; tibiae and tarsi nearly colorless, diffusively darkened on outer side (Figs 53–55). Abdominal terga with composite ornament of brown and colorless areas; terga I and VI darkest; tergum III darker than tergum II; anterior and posterior margins of terga bordered by dark brown; sterna colorless; segment X (tergum and paraprocts) light (Fig. 50). Cerci uniformly light brownish; paracercus lighter, with posterior margins of segments darkened with brown (Fig. 50, 72).</p><p>HYPODERMAL COLORATION. Unknown.</p><p>SHAPE AND SETATION: Labrum with anterior margin concave (Fig. 56). Incisors of mandibles wide and short, with most distal denticles shortest and bent ventrally (Figs 57–58).</p><p>Maxilla of peculiar structure, different from that of other Cheleocloeon (compare: Kluge 2016: fig. 2) (Figs 59, 61–63): Maxillary canines fused together forming short and flat apical plate with smooth outer margin; this apical plate borders bases of medio-ventral setal row from distal and ventral sides. Medio-ventral setal row terminates close to the apical plate; its 4 most distal setae are flattened and widened distally, much longer than the apical plate and arched backward from it; several previous setae are also arched at the same direction (Figs 61–62). Mediodorsal setal row contains 3 distinguishable dentisetae (Fig. 61).</p><p>Labium of peculiar structure, different from that of other Cheleocloeon (compare: Kluge 2016: figs 4–5) (Figs 60, 64–65). Glossae shortened and blunt apically. Paraglossae blunt apically, with enlarged number of long setae at apex. Disto-median projection of 2nd segment of labial palp (characteristic for Cheleocloeon) sharply projected, apically narrowly rounded, but not pointed (Fig. 66).</p><p>Legs slender. Claw long and slender, slightly arched, completely lacking denticles (Fig. 55).</p><p>Hind protoptera absent in male (Fig. 54) (the same should be in female).</p><p>Abdominal terga I–X with dense scales in wide sockets (Fig. 70). Posterior margin of tergum I with few irregular denticles; posterior margin of terga II–X with regular row of dark brown pointed denticles alternated with scales (Fig. 70). Posterior margins of abdominal sterna I–IV smooth; posterior margins of sterna V–IX with regular row of colorless triangular denticles, smaller than denticles on terga, without scales between them. Paraproct with pointed denticles (Fig. 69). All tergalii I–VII narrow and widest in distal half; tergalius I narrowest, shorter than next ones (Figs 73–78). Cerci without enlarged denticles on lateral sides; paracercus with several pointed enlarged denticles on dorsal side of posterior margin of each 4th segment (Fig. 72). Distal part of cercus with longitudinal row secondary swimming setae on lateral side (besides primary swimming setae on medial side) (Fig. 71).</p><p>Pose of subimaginal gonostyli under larval cuticle (retained in examined winged specimen). « Cloeon - type » (Fig. 79), as in other Cheleocloeon (see Kluge 2016: character 22 on p. 144).</p><p>Subimago. CUTICULAR COLORATION. Mesonotum light brownish, with darker brown areas (Fig. 82).</p><p>TEXTURE. All tarsal segments of all legs covered with pointed microlepides.</p><p>Imago, male (enclosed in subimaginal cuticle) (Fig. 81). Head ochre-brown. Turbinate eyes yellow. Thorax brown, equally dark on dorsal, ventral and lateral sides. Hind wings absent. Legs uniformly pale ochre. Abdominal terga light brownish-ochre, with poorly expressed reddish markings along median line, on posterior margins of terga and on sides of terga III and V; sterna uniformly light ochre. Shape of gonostyli unknown (not spread after molt from larva); gonovectes dark brown, structure of gonovectes typical for Cheleocloeon (Fig. 79–80). Cerci uniformly pale ochre.</p><p>Imago, female. Unknown.</p><p>Egg. Unknown.</p><p>Dimension. Fore wing length (and body length) about 3 mm.</p><p>Distribution. Known from one place in Tanzania.</p><p>Systematic position. The new species Ch. psammonella sp. nov. undoubtedly belongs to Cheleocloeon . Its shape of labial palp is typical for Cheleocloeon, i.e. with disto-median projection of the 2nd segment greatly projected medially and bent backward (Fig. 60); male imaginal gonovectes, which are developed in the specimen examined (Fig. 80), have structure characteristic for Cheleocloeon .</p><p>Comparison. Larva of Cheleocloeon psammonella sp. nov. differs from all other species of Cheleocloeon by the following features of the mouthparts’ structure: anterior margin of labrum is concave (Fig. 56) (rather than convex with median concavity in other species— Fig. 8); maxillary canines and setae of the medio-ventral row are modified forming a scraper (Figs 62–63) (rather than pointed maxillary canines and slender setae in other species— Fig.11); glossae and paraglossae are more blunt, and glossae are shorter than in all other species of Cheleocloeon (Figs 60, 64–65). These mouthparts modifications in Cheleocloeon psammonella sp. nov. resemble the modifications characteristic for the taxa Pseudocentroptiloides Jacob (in Jacob &amp; Glazaczow) 1987 and Psammonella Glazaczow (in Jacob &amp; Glazaczow) 1987, which belong to the genus Procloeon Bengtsson 1915 . Other representatives of Procloeon have the same structure of labrum, maxillae, glossae and paraglossae, as representatives of Cheleocloeon other than Ch. psammonella sp. nov.</p><p>Larva of Ch. psammonella sp. nov. also differs from all other species of Cheleocloeon by longer and slender legs with especially long and slender claws lacking denticles (Fig. 55) (in other species at least a few minute denticles are present—see Fig. 18 and Kluge 2016: figs 19–29).</p><p>Such leg structure is also characteristic for Pseudocentroptiloides and Psammonella. Probably, both the leg structure and the mouthparts’ structure are adaptations for inhabitancy on sand ground, convergently evolved in these taxa. The single larva of Ch. psammonella sp. nov. was collected in river Msosa, where most bottom consists of pure sand.</p></div>	https://treatment.plazi.org/id/DA3E8786F50C16412DCCF912FECF878B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2025): Two new species of the genus Cheleocloeon Wuillot & Gillies 1993 (Ephemeroptera: Baetidae) from Tanzania and corrections to the generic diagnosis. Zootaxa 5660 (3): 330-350, DOI: 10.11646/zootaxa.5660.3.2, URL: https://doi.org/10.11646/zootaxa.5660.3.2
