taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748220/files/figure.png	https://doi.org/10.5281/zenodo.15748220	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748232/files/figure.png	https://doi.org/10.5281/zenodo.15748232	Fig. 7. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184713). A. Ring 12, dorsal view. B. Rings 11–12, lateral view. Scale bars = 0.4 mm.	Fig. 7. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184713). A. Ring 12, dorsal view. B. Rings 11–12, lateral view. Scale bars = 0.4 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748240/files/figure.png	https://doi.org/10.5281/zenodo.15748240	Fig. 10. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573). A. Rings 11–12, dorsal view. B. Ring 12, right paranotum, dorsal view. C. Ring 12, right paranotum, lateral view. D. Paranotal dorsal microsculpture. E. Tip of epiproct, with spinnerets and lateral knobs. F. Midbody leg. Scale bars: A = 1 mm; B–C = 0.2 mm; D = 0.01 mm; E = 0.1 mm; F = 0.4 mm.	Fig. 10. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573). A. Rings 11–12, dorsal view. B. Ring 12, right paranotum, dorsal view. C. Ring 12, right paranotum, lateral view. D. Paranotal dorsal microsculpture. E. Tip of epiproct, with spinnerets and lateral knobs. F. Midbody leg. Scale bars: A = 1 mm; B–C = 0.2 mm; D = 0.01 mm; E = 0.1 mm; F = 0.4 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748222/files/figure.png	https://doi.org/10.5281/zenodo.15748222	Fig. 4. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Telson, ventral view. B. Tip of preanal process (epiproct) with spinnerets, posterior view. C–D. Right leg 10, anterior views. E. Sterna 11–12. F. Midbody dorsal limbus. Abbreviations: ds = dorsal seta on anal valve; ep = epiproct; gs = globular swelling; hp = subanal scale (hypoproct); pp = anal valve (paraproct); vs = ventral seta on anal valve. Scale bars: A, C–E = 0.5 mm; B = 0.05 mm; F = 0.02 mm.	Fig. 4. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Telson, ventral view. B. Tip of preanal process (epiproct) with spinnerets, posterior view. C–D. Right leg 10, anterior views. E. Sterna 11–12. F. Midbody dorsal limbus. Abbreviations: ds = dorsal seta on anal valve; ep = epiproct; gs = globular swelling; hp = subanal scale (hypoproct); pp = anal valve (paraproct); vs = ventral seta on anal valve. Scale bars: A, C–E = 0.5 mm; B = 0.05 mm; F = 0.02 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748224/files/figure.png	https://doi.org/10.5281/zenodo.15748224	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748228/files/figure.png	https://doi.org/10.5281/zenodo.15748228	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748234/files/figure.png	https://doi.org/10.5281/zenodo.15748234	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748238/files/figure.png	https://doi.org/10.5281/zenodo.15748238	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748244/files/figure.png	https://doi.org/10.5281/zenodo.15748244	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748246/files/figure.png	https://doi.org/10.5281/zenodo.15748246	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748220/files/figure.png	https://doi.org/10.5281/zenodo.15748220	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748222/files/figure.png	https://doi.org/10.5281/zenodo.15748222	Fig. 4. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Telson, ventral view. B. Tip of preanal process (epiproct) with spinnerets, posterior view. C–D. Right leg 10, anterior views. E. Sterna 11–12. F. Midbody dorsal limbus. Abbreviations: ds = dorsal seta on anal valve; ep = epiproct; gs = globular swelling; hp = subanal scale (hypoproct); pp = anal valve (paraproct); vs = ventral seta on anal valve. Scale bars: A, C–E = 0.5 mm; B = 0.05 mm; F = 0.02 mm.	Fig. 4. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Telson, ventral view. B. Tip of preanal process (epiproct) with spinnerets, posterior view. C–D. Right leg 10, anterior views. E. Sterna 11–12. F. Midbody dorsal limbus. Abbreviations: ds = dorsal seta on anal valve; ep = epiproct; gs = globular swelling; hp = subanal scale (hypoproct); pp = anal valve (paraproct); vs = ventral seta on anal valve. Scale bars: A, C–E = 0.5 mm; B = 0.05 mm; F = 0.02 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748224/files/figure.png	https://doi.org/10.5281/zenodo.15748224	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748228/files/figure.png	https://doi.org/10.5281/zenodo.15748228	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB6FFC0B562A7D2FABD792C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB6FFC0B562A7D2FABD792C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB6FFC0B562A7D2FABD792C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748232/files/figure.png	https://doi.org/10.5281/zenodo.15748232	Fig. 7. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184713). A. Ring 12, dorsal view. B. Rings 11–12, lateral view. Scale bars = 0.4 mm.	Fig. 7. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184713). A. Ring 12, dorsal view. B. Rings 11–12, lateral view. Scale bars = 0.4 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFB6FFC0B562A7D2FABD792C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748234/files/figure.png	https://doi.org/10.5281/zenodo.15748234	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFABFFC2B537A27DFE9F7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFABFFC2B537A27DFE9F7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748238/files/figure.png	https://doi.org/10.5281/zenodo.15748238	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA9FFC5B550A323FEBA7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA9FFC5B550A323FEBA7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748240/files/figure.png	https://doi.org/10.5281/zenodo.15748240	Fig. 10. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573). A. Rings 11–12, dorsal view. B. Ring 12, right paranotum, dorsal view. C. Ring 12, right paranotum, lateral view. D. Paranotal dorsal microsculpture. E. Tip of epiproct, with spinnerets and lateral knobs. F. Midbody leg. Scale bars: A = 1 mm; B–C = 0.2 mm; D = 0.01 mm; E = 0.1 mm; F = 0.4 mm.	Fig. 10. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573). A. Rings 11–12, dorsal view. B. Ring 12, right paranotum, dorsal view. C. Ring 12, right paranotum, lateral view. D. Paranotal dorsal microsculpture. E. Tip of epiproct, with spinnerets and lateral knobs. F. Midbody leg. Scale bars: A = 1 mm; B–C = 0.2 mm; D = 0.01 mm; E = 0.1 mm; F = 0.4 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA9FFC5B550A323FEBA7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748244/files/figure.png	https://doi.org/10.5281/zenodo.15748244	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA9FFC5B550A323FEBA7863.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748246/files/figure.png	https://doi.org/10.5281/zenodo.15748246	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFAEFFC7B548A323FE9F78F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFAEFFC7B548A323FE9F78F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748246/files/figure.png	https://doi.org/10.5281/zenodo.15748246	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFAEFFC7B548A323FE9F78F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748244/files/figure.png	https://doi.org/10.5281/zenodo.15748244	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFACFFC7B53CA3B4FAF07C95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748220/files/figure.png	https://doi.org/10.5281/zenodo.15748220	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	Fig. 3. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184702). A. Head, right antenna, collum and rings 2–3, oblique view.B. Clypeus and labrum. C. Head, collum and rings 2–3, dorsal view. D. Microsculpture of prozonite, ring 9. E. Ring 9, dorsal view. F. Rings 8–9, lateral view. G. Stigmata of ring 8, lateral view. H. Left paranotum and ozopore of ring 9, lateral view. Abbreviations: asp = anterior spiracle; eg = epicranial groove; lg = labrogenal offset; pe = peritreme with ozopore; pop = posterior spiracle; ts = transverse sulcus. Scale bars: A, C, E–F = 0.5 mm; B, G–H = 0.1 mm; D = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA3FFCCB538A238FB2B7B66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA3FFCCB538A238FB2B7B66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA3FFCCB538A238FB2B7B66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748250/files/figure.png	https://doi.org/10.5281/zenodo.15748250	Fig. 13. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661). A. Head, collum and rings 2–3, dorsal view.B. Three midbody rings, dorsal view.C. Paranotal microsculpture. D. Limbus. E. Tip of epiproct, with four spinnerets (one broken and lying loose) and globular swellings, posterior view. Scale bars: A–B = 1 mm; C = 0.05 mm; D = 0.01 mm; E = 0.03 mm.	Fig. 13. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661). A. Head, collum and rings 2–3, dorsal view.B. Three midbody rings, dorsal view.C. Paranotal microsculpture. D. Limbus. E. Tip of epiproct, with four spinnerets (one broken and lying loose) and globular swellings, posterior view. Scale bars: A–B = 1 mm; C = 0.05 mm; D = 0.01 mm; E = 0.03 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA3FFCCB538A238FB2B7B66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748252/files/figure.png	https://doi.org/10.5281/zenodo.15748252	Fig. 14. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661), gonopods. A. Oblique ventral view. B. Right gonopod, mesal view. C. Oblique dorsal view. Abbreviations: ca = cannula; cx = coxa; cxp = coxal process; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.1 mm.	Fig. 14. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661), gonopods. A. Oblique ventral view. B. Right gonopod, mesal view. C. Oblique dorsal view. Abbreviations: ca = cannula; cx = coxa; cxp = coxal process; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFCCB55BA027FEEB7992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748258/files/figure.png	https://doi.org/10.5281/zenodo.15748258	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFCCB55BA027FEEB7992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748260/files/figure.png	https://doi.org/10.5281/zenodo.15748260	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFF1B577A291FE94793D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFF1B577A291FE94793D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFF1B577A291FE94793D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748256/files/figure.png	https://doi.org/10.5281/zenodo.15748256	Fig. 15. Dionaeadesmus force Enghoff gen. et sp. nov. A–B. Paratype, ♂ from Ndundulu (NHMD 1184562). C–G. Paratype,♂ from Nyumbanito Mts (NHMD 1184572). A. Head, anterior view.B. Head, collum and ring 2, dorsal view. C. Ring 13, dorsal view. D. Ring 13, lateral view. E. Ring 12, posterior view. F. Sternum of ring 12. G. Legs from ring 12. Abbreviation: lg = labrogenal offset. Scale bars = A–D, F–G = 0.4 mm; E = 1 mm.	Fig. 15. Dionaeadesmus force Enghoff gen. et sp. nov. A–B. Paratype, ♂ from Ndundulu (NHMD 1184562). C–G. Paratype,♂ from Nyumbanito Mts (NHMD 1184572). A. Head, anterior view.B. Head, collum and ring 2, dorsal view. C. Ring 13, dorsal view. D. Ring 13, lateral view. E. Ring 12, posterior view. F. Sternum of ring 12. G. Legs from ring 12. Abbreviation: lg = labrogenal offset. Scale bars = A–D, F–G = 0.4 mm; E = 1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFF1B577A291FE94793D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748258/files/figure.png	https://doi.org/10.5281/zenodo.15748258	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FFA7FFF1B577A291FE94793D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748260/files/figure.png	https://doi.org/10.5281/zenodo.15748260	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748216/files/figure.png	https://doi.org/10.5281/zenodo.15748216	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	Fig. 1. Map of the Udzungwa Mts showing collecting areas for Chelodesmidae Cook, 1895. Yellow squares: Morogorius spp.D: M. divisus Enghoff sp. nov.,L: M. louishanseni Enghoff sp. nov., P: M. pugio Enghoff sp. nov. K: M. kitungulu Enghoff sp. nov., C: M. cochlear Enghoff sp. nov. Red squares: Tanzaniella howelli Hoffman, 1977. Blue squares Dionaeadesmus force Enghoff gen. et sp. nov. Purple squares: Callistocilla dolorotrix Hoffman, 2005. The position of the squares within the green circles/ ovoids does not indicate exact location. Based on Marshall et al. (2010: fig. 1).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748218/files/figure.png	https://doi.org/10.5281/zenodo.15748218	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	Fig. 2. Chelodesmids from the Udzungwa Mts. A. Morogorius divisus Enghoff sp. nov., paratype, ♀ (NHMD 1184699). B. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184563). C–D. Tanzaniella howelli Hoffman, 1977. C. Freshly preserved specimen (uncatalogued). D. Female from Mwanihana (NHMD 1184730). E. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♀ (NHMD 1184572), rings 11–13 missing. F. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). Scale bars: A–B, D–F = 2 mm; C not to scale. Photos: S.G. Selvantharan (A–B, D–F), A.R. Marshall (C).	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748262/files/figure.png	https://doi.org/10.5281/zenodo.15748262	Fig. 18. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). A. Rings 11–13, dorsal view; inset: close-up of crenulation. B. Telson and rings 18–19, dorsal view. C. Epiproct and spinnerets, posterior view. D. Limbus from ring 11. E. Microsculpture of paranotum 11. Scale bars: A–B = 0.2 mm; C = 0.05 mm; D = 0.01 mm; E = 0.005 mm.	Fig. 18. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). A. Rings 11–13, dorsal view; inset: close-up of crenulation. B. Telson and rings 18–19, dorsal view. C. Epiproct and spinnerets, posterior view. D. Limbus from ring 11. E. Microsculpture of paranotum 11. Scale bars: A–B = 0.2 mm; C = 0.05 mm; D = 0.01 mm; E = 0.005 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748265/files/figure.png	https://doi.org/10.5281/zenodo.15748265	Fig. 19. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742), right gonopod. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. E. Solenomere and process of parasolenomeree. Abbreviations: cx = coxa; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm.	Fig. 19. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742), right gonopod. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. E. Solenomere and process of parasolenomeree. Abbreviations: cx = coxa; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748267/files/figure.png	https://doi.org/10.5281/zenodo.15748267	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9DFFF6B539A24EFB5C7E8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748267/files/figure.png	https://doi.org/10.5281/zenodo.15748267	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9DFFF6B539A24EFB5C7E8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748262/files/figure.png	https://doi.org/10.5281/zenodo.15748262	Fig. 18. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). A. Rings 11–13, dorsal view; inset: close-up of crenulation. B. Telson and rings 18–19, dorsal view. C. Epiproct and spinnerets, posterior view. D. Limbus from ring 11. E. Microsculpture of paranotum 11. Scale bars: A–B = 0.2 mm; C = 0.05 mm; D = 0.01 mm; E = 0.005 mm.	Fig. 18. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742). A. Rings 11–13, dorsal view; inset: close-up of crenulation. B. Telson and rings 18–19, dorsal view. C. Epiproct and spinnerets, posterior view. D. Limbus from ring 11. E. Microsculpture of paranotum 11. Scale bars: A–B = 0.2 mm; C = 0.05 mm; D = 0.01 mm; E = 0.005 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9DFFF6B5CDA59DFE6E7D1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748267/files/figure.png	https://doi.org/10.5281/zenodo.15748267	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	Fig. 20. Callistocilla spp. A–B. C. dolorotrix Hoffman, 2005, male from Udzungwa Mountains National Park (NHMD 1184737).A. Rings 12–13, right paranota.B. Right gonopod, apical part.C–E. Callistocilla sp. 1, male from Kanga Mountains (NHMD 1184571), ring 12. C. Dorsal view. D. Right ozopore. E. Microsculpture of paranotum. F. C. beatrix Hoffman, 1977, holotype, ♂ (ZMUC 00101515), microsculpture of paranotum 13. Abbreviations: fx = flexed edge of solenomere; ps = parasolenomere; slm = solenomere. Scale bars: A, C = 0.5 mm; B = 0.05 mm; D = 0.02 mm; E–F = 0.005 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748265/files/figure.png	https://doi.org/10.5281/zenodo.15748265	Fig. 19. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742), right gonopod. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. E. Solenomere and process of parasolenomeree. Abbreviations: cx = coxa; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm.	Fig. 19. Callistocilla dolorotrix Hoffman, 2005, male from Udzungwa Scarp Nature Reserve (NHMD 1184742), right gonopod. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. E. Solenomere and process of parasolenomeree. Abbreviations: cx = coxa; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748258/files/figure.png	https://doi.org/10.5281/zenodo.15748258	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	Fig. 16. Dionaeadesmus force Enghoff gen. et sp. nov. A, E–F. Paratype, ♂ from Ndundulu (NHMD 1184562). B–D. Paratype, ♂ from Nyumbanito Mts (NHMD 1184572). A. Rings 18–19 and telson, dorsal view. B. Tip of epiproct, with four spinnerets and globular swellings, posterior view. C. Paranotal microsculpture. D. Limbus. E. Right gonopod, submesal view. F. Right gonopod, anterio-mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; cx = coxa; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; slm = solenomere; sph = solenophore(?); spr = spiked ridge. Scale bars: A = 0.4 mm; B, E–F = 0.1 mm; C–D = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748260/files/figure.png	https://doi.org/10.5281/zenodo.15748260	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	Fig. 17. Dionaeadesmus force Enghoff gen. et sp. nov., paratype, ♂ from Nyumbanito Mts (NHMD 1184572), gonopods. A. Dorsal view. B. Ventral view. C. Sublateral (right) view. D. Left telopodite. E. Left solenomere. Abbreviations: as = anterior coxal setae; cxp = coxal process; msp = marginal spikes; prf = prefemoral part; prp = prefemoral process; prt = prefemoral tooth; ps = parasolenomere; sph = solenophore(?); spr = spiked ridge; sslm = solenomere s. str. Scale bars: A–C = 0.2 mm; D = 0.1 mm; E = 0.05 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748252/files/figure.png	https://doi.org/10.5281/zenodo.15748252	Fig. 14. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661), gonopods. A. Oblique ventral view. B. Right gonopod, mesal view. C. Oblique dorsal view. Abbreviations: ca = cannula; cx = coxa; cxp = coxal process; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.1 mm.	Fig. 14. Tanzaniella howelli Hoffman, 1977, male from Udzungwa Mountains National Park (NHMD 1184661), gonopods. A. Oblique ventral view. B. Right gonopod, mesal view. C. Oblique dorsal view. Abbreviations: ca = cannula; cx = coxa; cxp = coxal process; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748224/files/figure.png	https://doi.org/10.5281/zenodo.15748224	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 5. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184699), left gonopod, A. Ventral view. B. Dorsal view. C. Lateral view with directions indicated. D. Submesal view. E. Mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748228/files/figure.png	https://doi.org/10.5281/zenodo.15748228	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	Fig. 6. Morogorius divisus Enghoff sp. nov., paratype, ♂ (NHMD 1184701), right gonopod, prefemoral process cut at X. A. Meso-ventral view. B. Sub-anterior view. C. Tip of solenomere. D. Separated prefemoral process. Abbreviations: ps = parasolenomere; slm = solenomere. Scale bars. A = 0.2 mm; B, D = 0.1 mm; C = 0.01 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748238/files/figure.png	https://doi.org/10.5281/zenodo.15748238	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	Fig. 9. Morogorius pugio Enghoff sp. nov., holotype, ♂ (NHMD 1184710), left gonopod. A. Lateral view. B. Tip of solenomere. C. Mesal view. D. Ventral view (most prefemoral setae trimmed). E. Parasolenomere, isolated. F. Anterior view. Abbreviations: as = anterior coxal seta; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; nb = needlelike side branch; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere. Scale bars: A, C–D, F = 0.2 mm; B = 0.02 mm; E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748244/files/figure.png	https://doi.org/10.5281/zenodo.15748244	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	Fig. 11. Morogorius kitungulu Enghoff sp. nov., holotype, ♂ (NHMD 1184573), left gonopod. A. Mesal view. B. (Dorso)lateral view. C. Ventrolateral view. D. Acropodite, meso-ventral view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; slm = solenomere; sr = serrate ridge of solenomere; sslm = solenomere s. str. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748246/files/figure.png	https://doi.org/10.5281/zenodo.15748246	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	Fig. 12. Morogorius cochlear Enghoff sp. nov., holotype, ♂ (NHMD 1184574), left gonopod. A. Mesal view. B. Lateral view. C. Close-up of parasolenomere. D. Ventral view. E. Acropodite, mesal view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; ls = lateral coxal setae; prf = prefemoral part; prp = prefemoral process; ps = parasolenomere; psp = process of parasolenomere; slm = solenomere; sslm = solenomere s. str.; tr = triangular canopy of prefemoral process. Scale bars: A–B, D = 0.2 mm; C, E = 0.1 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
D924C438FF9CFFF8B7C2A4CAFA9878BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15748234/files/figure.png	https://doi.org/10.5281/zenodo.15748234	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	Fig. 8. Morogorius louishanseni Enghoff sp. nov., paratype, ♂ (NHMD 1184653), left gonopod. A. Lateral view. B. Submesal view. C. Dorsal view. D. Ventral view. E. Anterior view. Abbreviations: as = anterior coxal setae; ca = cannula; ci = cingulum; cx = coxa; cxp = coxal process; mvs = mesoventral setae; prf = prefemoral part; prp = prefemoral process; ps =parasolenomere; slm = solenomere. Scale bars = 0.2 mm.	2025-06-20	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.		Zenodo	biologists	Enghoff, Henrik;Ngute, Alain Senghor K.;Mnendendo, Hamidu R.;Kivambe, Exaud E.;Mhagawale, Waziri;Malanda, Robin C.;Mpoto, Athuman;Marshall, Andrew R.			
