taxonID	type	description	language	source
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	type_taxon	Type species Morogorius pallidus Verhoeff, 1941 (Tanzania) by monotypy.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	diagnosis	History and diagnosis The history of Morogorius is brief. In the original description, it was placed in subfamily Cordyloporinae Brölemann, 1916 (now: Prepodesminae) and was diagnosed as follows (translated from Verhoeff 1941: 249): “ This genus is based on the absence of dorsal processes, on lack of denticulation on the paranota as well as on the hind margin of the diplosomites between paranota and sterna, and on characters of the gonopods ” and “ Different from the related genera Cordyloporus, Paracordyloporus, Scolopopleura and Graphidochirus in the gonopods, in part also in body characters ” (Verhoeff 1941: 249 – 250). Chamberlin (1952) didn’t mention Morogorius, nor did Demange & Mauriès (1975). Hoffman (1977) restudied and re-drew the gonopods of the type species and compared it to his Tanzaniella howelli Hoffman, 1977. That’s all. A useful differential diagnosis of Morogorius is thus not available, as is the case for very many millipede genera. Considering the very large number of genera in Chelodesmidae, in combination with the lack of useful diagnoses of the family Chelodesmidae and the subfamily Prepodesminae, such a diagnosis cannot be presented here. However, Morogorius differs from other Tanzanian genera of Chelodesmidae in the characters presented in Table 2. The homology of the largest gonopodal process in Morphotelus and Mesodesmus is uncertain, therefore the process may preliminary be termed “ solenophore ”.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFBFFFD8B532A4C6FAE37F0F.taxon	description	General description of Morogorius (males) SIZE. Length 21 – 31 mm, max. width 3.6 – 5.7 mm. Collum wider than head but narrower than ring 2, roughly parallel-sided from ring 3 to midbody, thereafter moderately tapering. COLOUR. Overall colour (reddish) brown, in part (species-specific) with some or all paranota contrastingly pale (Fig. 2 A – B). HEAD (Fig. 3 A – B). Densely setose below and between antennal sockets, labrogenal offset not very pronounced, epicranial groove distinct, interantennal space narrow, 0.5 – 0.6 × length of antennomere 2. Antennae reaching back to ring 6 when stretched. COLLUM (Fig. 3 A, C). In the shape of an isosceles curved trapezoid with the longer margin anteriorly, lateral corners smoothly rounded, surface granular, especially laterally. A row of 2 – 4 setae close to anterior margin. BODY RINGS (Figs 3 C – H, 7, 10 B – F). Prozonites smooth, with simple, cellular microsculpture (Fig. 3 D). Metazonites flat, granular dorsally and laterally, 1.4 – 1.8 × as broad as prozonites, with 1 + 1 small, thin setae anteriorly (setae apparently absent in M. pugio Enghoff sp. nov.) and a prominent transverse sulcus on rings 5 – 18. Paranota horizontal, those of rings 2 – 3 very slightly projecting forwards, following paranota rectangular but from some point (ring 5 – 15 according to species) with posterior corners increasingly projecting backward. Paranotal microsculpture (Fig. 10 D) formed by longitudinally stretched, narrow cytoscutes of more or less irregular shape. Ozopore formula normal (5, 7, 9 – 10, 12 – 13, 15 – 19), ozopores circular, delimited by smooth ring in peritremata on edge of paranota, ca midway between anterior and posterior margin. Sterna (Fig. 4 E) broad, with a transverse impression and a row of fine setae near anterior margin. Spiracles (only studied in M. divisus Enghoff sp. nov.): anterior spiracle on each diploring pear-shaped, posterior spiracle subcircular, both with plugs showing a very pronounced cellular pattern (Fig. 3 G). Limbus (Fig. 4 F) with straight margin, consisting of a densely, finely striate marginal zone preceded by a row of rectangular cells. LEGS (Figs 4 C – D, 10 F). Slender, without modifications, length 1.2 – 1.6 × maximum body width, length not increasing towards hind end, TELSON (Fig. 4 A – B). Preanal ring with several setae along dorsal part of posterior margin; epiproct large, with prominent lateral setiferous tubercles; spinnerets arranged in an almost quadratic trapezoid on a smooth terminal swelling, simple, flanked by 1 + 1 globular, sometimes partly collapsed structures. Anal valves (paraprocts, pp) unmodified, each with 2 setae, dorsal seta (ds) on marginal rim, ventral seta (vs) more lateral, Subanal scale (hypoproct, hp) semicircular, unmodified, with 1 + 1 marginal setae. GONAPOPHYSES. Short, triangular. GONOPOD APERTURE. Transversely oval, more than twice as broad as long, rim simple. GONOPODS (Figs 5 – 6, 8 – 9, 11 – 12). No sternal remnant. Coxa (cx) cylindrical, short, length ≈ diameter (perhaps slightly longer in M. pallidus), with a conical process (cxp) distally on the anterior side (absent to barely discernible in M. divisus sp. nov.); two long setae on anterior surface basal to process, a field of up to ca a dozen setae (ls) on lateral surface (except in M. louishanseni sp. nov. and perhaps M. pallidus) and numerous setae on meso-posterior surface. Cannula (ca) stout. Prefemoral part (prf) bent at right angles relative to cx, cylindrical: length 2 – 2 ½ × diameter (maybe slightly less in M. pallidus), numerous long setae on ventral surface. Efferent canal running straight on meso-dorsal side of prf, at level of base of parasolenomere bending laterad into solenomere; prefemoral process (prp) originating dorso-laterally from main body of prf, at least as long as solenomere and parasolenomere, basally slender, apically more or less expanded, of species-specific shape and curving over solenomere and parasolenomere; prf distally delimited by cingulum (ci), cingulum very distinct on ventral side of gonopod (except in M. kitungulu Enghoff sp. nov. where it is less distinct). Acropodite: main body strongly reduced, indiscernible, giving rise to solenomere and parasolenomere. Solenomere (slm) slightly shorter than prp (apparently same length in M. pallidus according to the original decription of that species), several times as long as broad, of species-specific shape, either regularly tapering or abruptly narrowing at ⅔ of its length. Parasolenomere (ps) originating mesal to slm, of species-specific shape, more or less shorter than slm, sometimes simple, sometimes deeply divided into two branches.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	description	urn: lsid: zoobank. org: act: 48 AFBD 40 - 57 E 0 - 4 F 1 B- 9 F 58 - ABF 792 FFE 13 E Figs 1, 2 A, 3 – 6	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	diagnosis	Diagnosis Differs from other species of Morogorius by the deep division of the parasolenomere (ps) into two long, subequal lobes (Fig. 6 A) and by the very poorly developed coxal process (Fig. 5 C).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	etymology	Etymology Named after the deeply divided parasolenomere. Adjective.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	materials_examined	Material examined (total 11 ♂♂, 11 ♀♀) Holotype TANZANIA • ♂; Iringa Region, Kilolo District, Udzungwa Scarp Nature Reserve, Uhafiwa; 8 ° 31 ′ 58.404 ″ S, 35 ° 51 ′ 28.368 ″ E; 1375 m a. s. l.; 25 Jan. 2022. A. Ngute, R. Malanda, W. Mhagawale, H. Mnendendo, A. Mpoto and A. Marshall leg.; FoRCE Plot 106, open canopy; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184703. Paratypes TANZANIA – Iringa Region, Kilolo District, Udzungwa Scarp Nature Reserve • 8 ♂♂, 7 ♀♀; same data as for holotype; NHMD 1184699, NHMD 1184701, NHMD 1184702 • 2 ♂♂, 2 ♀♀; same data as for holotype; Uhafiwa; 8 ° 31 ′ 49.296 ″ S, 35 ° 51 ′ 20.772 ″ E; 1354 m a. s. l.; 24 Jan. 2022; FoRCE Plot 107, open canopy; NHMD 1184704 to NHMD 1184706 • 2 ♀♀; Chita Juu; 8 ° 32 ′ 23.388 ″ S, 35 ° 51 ′ 36 ″ E; 1387 m a. s. l.; 1 Feb. 2022; A. Ngute, R. Malanda, W. Mhagawale, H. Mnendendo, A. Mpoto and A. Marshall leg.; FoRCE Plot 109, open canopy; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184708, NHMD 1184709.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	description	Description (males) SIZE. Length 24 – 27 mm, max. width 4.4 – 4.8 mm. COLOUR (Fig. 2 A). After 2 years in alcohol head, collum and most of dorsum reddish-brown. Paranota of rings 2, 5, 7, 9 – 10, 12 – 15 and 15 – 19 contrastingly white, like epiproct and paraprocts. Antennae, venter and legs light brown; antennomeres 1 and 5 – 7 whitish. COLLUM (Fig. 3 A, C) with a row of 4 setae close to anterior margin. BODY RINGS (Fig. 3 C, E – H). Metazonites 1.7 – 1.8 × as broad as prozonites, with 1 + 1 small, thin setae anteriorly and a prominent transverse sulcus from ring until ring 17 or 18, granulation regular also behind sulcus. Paranota rectangular until ring 13 or 14, thereafter posterior corners projecting increasingly backwards, triangular, those of rings 16 – 18 with a few denticles at base of mesal margin. Anterior spiracle (asp) on each body ring pear-shaped, posterior spiracle (pop) subcircular, both with plugs showing a very pronounced cellular pattern (Fig. 3 G). Ozopores (Fig. 3 E, H) in smooth peritremata (pe) on edge of paranota, ca midway between anterior and posterior margin. Sides of rings with a row of slightly larger tubercles along posterior margin. LEGS (Fig. 4 C – D). Length 1.3 × maximum body width. GONOPODS (Figs 5 – 6). Coxa (cx) with a barely discernible apical process (cxp), with two long setae (as) a on anterior surface field of ca a dozen setae (ls) on lateral surface. Prefemoral part (prf) ca 2 × as long as broad. Prefemoral process (prp) closely appressed to and distally curving over solenomere; prp slender in basal 2 / 3, distally expanded and ending in two tines. Solenomere (slm) long, slender, taeniate with parallel margins, at ca 4 / 5 of its length abruptly narrowed from ventral side; apical 1 / 5 thin, tapering. Parasolenomere (ps) deeply divided into two branches; ventral branch ca 3 / 4 as long as solenomere, slender; dorsal branch slightly shorter and broader, apically strongly tapering.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB3FFDDB53EA41FFDF97C53.taxon	distribution	Distribution and habitat Only known from three sites in the Udzungwa Scarp Nature Reserve (Fig. 1). Collected under open canopy at 1354 – 1387 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	description	urn: lsid: zoobank. org: act: 4 F 551606 - 55 DA- 40 F 7 - 818 B-C 21 F 9 BA 84545 Figs 1, 2 B, 7 – 8	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	diagnosis	Diagnosis Differs from other species of Morogorius by the shape of the parasolenomere: broad base, lateral shoulder and tapering apical part (Fig. 8 C – D).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	etymology	Etymology Named after Louis Hansen, collector of many new millipede species in the Udzungwa Mts. Noun in genitive.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	materials_examined	Material examined (total 31 ♂♂, 10 ♀♀) Holotype TANZANIA • ♂; Iringa Region, Mahenge Distr., West Kilombero Scarp Forest Reserve, Nyumbanitu Mts, S Udekwa Village; 07 ° 48 ′ S, 36 ° 21 ′ E; 1500 m a. s. l.; Dec. 1993; J. O. Svendsen leg.; NHMD 1184745. Paratypes TANZANIA • 9 ♂♂, 5 ♀♀; same data as for holotype; NHMD 1184713 • 4 ♂♂, 2 ♀♀; same data as for holotype; 1700 m a. s. l.; NHMD 1184577 • 1 ♂, 2 ♀♀; Iringa Region, Udzungwa Mts, Ndundulu Forest, Waller’s camp; 07 ° 46 ′ S, 36 ° 29 ′ E; 1550 m a. s. l.; Dec. 2006; L. A. Hansen and local assistants leg.; tropical semi-evergreen forest; NHMD 1184700 • 2 ♂♂, 1 ♀; same data as for preceding; 1 – 12 Jan. 2007; NHMD 1184712 • 14 ♂♂; Udzungwa Mts, Kitungulu Forest Reserve; 08 ° 09 ′ S, 36 ° 05 ′ E; 1500 m a. s. l.; Jan. 1996; M. Andersen, P. Gravlund and A. Jakobsen leg.; NHMD 1184563.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	description	Description (males) SIZE. Length 29 – 30 mm, max. width 3.6 – 3.8 mm. COLOUR (Fig. 2 B). After 17 – 28 years in alcohol head light brown, collum light brown with broad whitish margin. Dorsal side of rings with posterior half of each ring whitish to very light brown, anterior half of each ring brownish; paranota whitish, no difference between poriferous and non-poriferous rings. Antennae, legs, venter and telson very light brown to whitish. COLLUM. With a row of 4 setae close to anterior margin. BODY RINGS (Fig. 7). Metazonites 1.4 – 1.6 × as broad as prozonites, with 1 + 1 small, thin setae anteriorly and a prominent transverse sulcus on rings 5 – 17, granulation irregular, with a tendency to longitudinal striolation behind sulcus. Paranota already from ring 5 with posterior corners projecting increasingly backwards, first as rounded lobes, further back as pointed triangles, those of ring 16 with a few denticles at base of mesal margin. Ozopores in smooth peritremata on edge of paranota, ca midway between anterior and posterior margin. Sides of rings with a row of slightly larger tubercles along posterior margin. LEGS. Length 1.5 – 1.6 × maximum body width. GONOPODS (Fig. 8). Coxa (cx) with a stout cone-shaped apical process (cxp), two long setae (as) at base of process, no setae on lateral surface. Prefemoral part (prf) ca 2 × as long as broad. Prefemoral process (prp) closely appressed to and distally curving over solenomere, prp at base with triangular expansion, distally expanding, its tip curved back on itself. Solenomere (slm) long, slender, regularly tapering. Parasolenomere (ps) slightly shorter than solenomere, lamellate, basal part broad, lateral margin with marked ‘ shoulder’, ps thereafter more or less regularly tapering.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFB6FFC0B562A7D2FABD792C.taxon	distribution	Distribution and habitat Known from three sites in the Udzungwa Mountains: Nyumbanitu Mountains, Ndundulu Forest and Kitungulu Forest Reserve (Fig. 1). Collected in tropical semi-evergreen forest at 1500 – 1550 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	description	urn: lsid: zoobank. org: act: 66768505 - 0 FD 5 - 4 E 1 B-AC 6 F-D 763 ADD 4639 E Figs 1, 9	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	diagnosis	Diagnosis Differs from other species of Morogorius by the simple thin, straight, pointed parasolenomere with a small needlelike side branch (Fig. 9 D – E).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	etymology	Etymology Named after the parasolenomere which reminds of a dagger (Latin ‘ pugio ’). Noun in apposition.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	materials_examined	Material examined (total 1 ♂) Holotype TANZANIA • ♂; Morogoro Region, Udzungwa Mts Natl. Park, Kidatu; 07 ° 40 ′ 42.1 ″ S, 36 ° 55 ′ 06.8 ″ E; 1482 m a. s. l.; 21 Apr. 2014; J. Malumbres-Olarte leg.; Plot 6, pitfall trap, sample code 6 PT 8; NHMD 1184710.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	description	Description (male) SIZE. Length ca 31 mm, max. width 5.7 mm. COLOUR. After 10 years in alcohol head medium brown, collum and dorsal side of body rings dark; paranota 2, 3, 5, 7, 9 – 10, 12 – 13 and 15 – 19 contrastingly whitish, paranota 4, 6 and 8 of a slightly lighter hue than main part of dorsum, sides of body rings medium brown; telson light brown, epiproct whitish; antennae and venter light brown; legs whitish brown. COLLUM. With pair of setae close to anterior margin. BODY RINGS. Metazonites 1.4 × as broad as prozonites, apparently without setae; with a prominent transverse sulcus on rings 5 – 17. Granulation irregular, but with a tendency to longitudinal striolation behind sulcus. Paranota from ring with 7 posterior corners projecting slightly backwards, first as rounded lobes, further back more triangular. Sides of rings with uniform tuberculation, no larger tubercles along posterior margin. LEGS. Length 1.2 × maximum body width. GONOPODS (Fig. 9). Coxa (cx) with a strong conical apical process (cxp), one long seta (as) on anterior surface basal to process, a field of setae (ls) on lateral surface. Prefemoral part (prf) ca 2 × as long as broad. Prefemoral process (prp) closely appressed to and distally curving over solenomere. Solenomere (slm) long, slender, taeniate, pointed. Parasolenomere (ps) much shorter than solenomere, thin, straight, pointed, with small needlelike side branch (nb) basally.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFABFFC2B537A27DFE9F7863.taxon	distribution	Distribution and habitat Known only from one site in the Udzungwa Mountains National Park (Fig. 1). Collected in a pitfall trap at 1482 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	description	urn: lsid: zoobank. org: act: 052 AECF 5 - 0923 - 403 F-AEC 2 - F 2 E 1 F 918 DB 22 Figs 1, 10 – 11	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	diagnosis	Diagnosis Differs from other species of Morogorius, except M. cochlear sp. nov., by the short, compact parasolenomere. Differs from M. cochlear by the pointed shape of the parasolenomere, the slenderer prefemoral process and the larger coxal process (Fig. 11, compare with Fig. 12).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	etymology	Etymology Named after the type locality. Noun in apposition.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	materials_examined	Material examined (total 1 ♂) Holotype TANZANIA • ♂; Udzungwa Mts, Kitungulu Forest Reserve; 08 ° 09 ′ S, 36 ° 05 ′ E; 1500 m a. s. l.; Jan. 1996; M. Andersen, P. Gravlund and A. Jakobsen leg.; NHMD 1184573.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	description	Description (male) SIZE. Length ca 27 mm, max. width 4.6 mm. COLOUR. After 28 years in alcohol head brownish; dorsal side of all rings including collum brownish with contrasting whitish paranota on all rings, post-sulcus area of rings also whitish; telson anteriorly brownish, posteriorly whitish; antennae, legs and ventral side light brown. COLLUM. With two setae close to anterior margin. BODY RINGS (Fig. 10 A – C). Metazonites ca 1.6 times as broad as prozonites, with 1 + 1 small, thin setae anteriorly, a prominent transverse sulcus on rings 5 – 18, granulated with irregular shallow, longitudinal striae behind transverse sulcus. Paranota rectangular until ring 14, thereafter posterior corners projecting increasingly backwards, triangular. Ozopores in smooth peritremata on edge of paranota, ca midway between anterior and posterior margin (Fig. 10 B – C). Sides of rings with uniform tuberculation, no larger tubercles along posterior margin. LEGS (Fig. 10 F). Length 1.2 × maximum body width. GONOPODS (Fig. 11). Coxa (cx) with a long, stout, conical apical process (cxp) on anterior side, two long setae (as) on anterior surface basal to process, a field of ca a dozen setae (ls) on lateral surface. Prefemoral part (prf) ca 2 ½ × as long as broad, distally delimited by distinct cingulum (ci) on ventral side. Efferent duct running straight on meso-dorsal side of prefemoral part, at level of base of parasolenomere bending laterad into hollowed side of acropodite. Prefemoral process (prp) with slender club-shaped outline, apical part sub-rhomboid, dorsal surface concave (accommodating solenomere), apically forming hood over tip of solenomere; a few denticles on distal margin and subdistally on dorsal margin. Solenomere (slm) shorter (ca 0.8 ×) than prp, stout and slightly tapering in basal ⅔, then abruptly narrower, forming slender hook, the solenomere s. str. (sslm); a slightly serrate high, short ridge (s r) facing concave side of sslm. Parasolenomere (ps) originating mesal to slm, much shorter than slm, stout, pointed.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA9FFC5B550A323FEBA7863.taxon	distribution	Distribution and habitat Known only from one site in the Kitungulu Forest Reserve, Udzungwa Mountains (Fig. 1). Collected at 1300 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	description	urn: lsid: zoobank. org: act: 5 E 6446 F 3 - AF 09 - 4 A 1 F- 8716 - E 117 ECC 2 D 99 A Figs 1, 12	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	diagnosis	Diagnosis Differs from other species of Morogorius, except M. kitungulu sp. nov., by the short, compact parasolenomere. Differs from M. kitungulu by the rounded shape of the parasolenomere, which has a retrose pointed process, the broader, spoonlike prefemoral process and the smaller coxal process (Fig. 12, compare with Fig. 11).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	etymology	Etymology Named after the prefemoral process which is in the shape of a spoon (Latin ‘ cochlear ’). Noun in apposition.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	materials_examined	Material examined (total 1 ♂) Holotype TANZANIA • ♂; Morogoro Region, Udzungwa Mountains National Park, Kidatu; 07 ° 41 ′ 06.2 ″ S, 36 ° 54 ′ 52.4 ″ E; 1527 m a. s. l.; 23 Oct. 2014; J. Malumbres-Olarte leg.; Plot 9, pitfall trap; NHMD 1184574.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	description	Description (male) SIZE. Length ca 25 mm, max. width 5.0 mm. COLOUR. After 10 years in alcohol head brownish; dorsal side of all rings including collum brownish with whitish paranota on all rings; antennae light brown; telson, legs and ventral side whitish. COLLUM. With three setae close to anterior margin (a fourth setae has probably been present, too). BODY RINGS. Ca 1.6 times as broad as prozonites, with 1 + 1 small, thin setae anteriorly, a prominent transverse sulcus on rings 5 – 18, granulated with irregular shallow, longitudinal striae behind transverse sulcus. Paranota rectangular until ring 13, thereafter posterior corners projecting increasingly backwards, triangular. Ozopores in smooth peritremata on edge of paranota, ca midway between anterior and posterior margin. Sides of rings with uniform tuberculation, no larger tubercles along posterior margin. LEGS. Length ≈ 1.3 × maximum body width. GONOPODS (Fig. 12). Coxa (cx) with a pointed, slightly curved apical process (cxp), two long setae (as) on anterior surface basal to process, and a field of ca a dozen setae (ls) on lateral surface. Prefemoral part (prf) ca 2 ½ × as long as broad, distally delimited by distinct cingulum (ci) on ventral side. Prefemoral process (prp) with club-shaped outline, apical part almost circular, dorsal surface concave (accommodating solenomere), apically forming hood over tip of solenomere; distal margin with stout, triangular, dorsad canopy (tr). Solenomere (slm) shorter (ca 0.8 ×) than prp, stout and slightly tapering in basal ⅔, then abruptly narrower, forming slender hook, the solenomere s. str. (sslm). Parasolenomere (ps) originating mesal to slm, much shorter than slm, stout, subrectangular, with short, pointed retrorse process (psp) dorsally.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	discussion	Remarks Very similar to M. kitungulu sp. nov., sharing, i. a., the general shape of the solenomere and the short, stout parasolenomere. Differs from M. kitungulu in the detailed shape of all three processes.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFAEFFC7B548A323FE9F78F6.taxon	distribution	Distribution and habitat Known only from one site in the Udzungwa Mountains National Park (Fig. 1). Collected in a pitfall trap at 1527 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC7B53CA3B4FAF07C95.taxon	diagnosis	Diagnosis Differs from other species of Morogorius by the deep division of the parasolenomere into a long slender branch and a considerably shorter daggerlike branch, in combination with a well-developed ‘ shoulder’ of the solenomere (Verhoeff 1941: fig. 21; Hoffman 1977: fig. 11).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC7B53CA3B4FAF07C95.taxon	materials_examined	Material examined (total 1 ♀) Paratype TANZANIA • 1 ♀; “ Tanganyika ”; “ 1123 ”; ZMB 2696.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC7B53CA3B4FAF07C95.taxon	description	Descriptive notes Body length ca 23 mm; max. width 3.0 mm. 20 ‘ segments’, i. e., 18 podous + 1 apodous ring + telson. A side-by side comparison with a female of Morogorius divisus Enghoff sp. nov. shows the following differences: – the posterior margin of the collum is straight, not ‘ trapezoid’ (cp. Fig. 3 C); – the peritremata are clearly, not indistinctly set off from the lateral margin of the paranota (cp. Fig. 3 E). Details of spinnerets not visible (no SEM).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC7B53CA3B4FAF07C95.taxon	discussion	Remarks Maybe this female is not conspecific with the holotype male and maybe even belong to another genus? In any case, there is no doubt that the gonopods of M. pallidus as described by Verhoeff (1941) and Hoffman (1977) are ‘ congeneric’ with those of the new species of Morogorius described above.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC8B532A794FE3F796B.taxon	type_taxon	Type species Tanzaniella howelli Hoffman, 1977 (Tanzania) by monotypy.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC8B532A794FE3F796B.taxon	diagnosis	Diagnosis Tanzaniella differs from other Tanzanian genera of Chelodesmidae as shown in Table 2, notably by the solenomere which is very much broader than and by far more conspicuous than the other terminal telopodital process.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFACFFC8B532A794FE3F796B.taxon	discussion	Remarks Hoffman (1977) compared the type and only species of Tanzaniella with the type (and then only) species of Morogorius, viz., M. pallidus Verhoeff, 1941. He noticed several specific similarities including a broadened solenomere and realized that “ eventual annectant forms may be found that connect the two nominal genera ” (Hoffman 1977: 76). Although five additional species of Morogorius have been described here, T. howelli still stands out by the extreme development of the solenomere, compared with a very thin prefemoral process (tibiotarsus of Hoffman 1977). For the time being the two genera are therefore kept separate.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	description	Figs 1, 2 C – D, 13 – 14	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	diagnosis	Diagnosis Redundant, genus monotypic.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	materials_examined	Material examined (total 34 ♂♂, 42 ♀♀, 13 juv.) TANZANIA – Morogoro Region, Kilombero District, Magombera Nature Reserve • 1 ♀, 1 juv.; 7 ° 48 ′ 46.332 ″ S, 36 ° 58 ′ 32.2314 ″ E; 273 m a. s. l.; 26 Dec. 2019; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 2; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184714 • 1 ♀; 7 ° 48 ′ 53.946 ″ S, 36 ° 59 ′ 20.025 ″ E; 267 m a. s. l.; 13 Jan. 2019; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 7; NHMD 1184715 • 3 ♂♂, 2 ♀♀; 7 ° 49 ′ 28.0344 ″ S, 36 ° 58 ′ 55.236 ″ E; 271 m a. s. l.; 31 Jan. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 13; NHMD 1184716 • 2 ♂♂, 4 ♀♀; 7 ° 48 ′ 53.154 ″ S, 36 ° 58 ′ 2.7948 ″ E; 282 m a. s. l.; 5 Mar. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 22; NHMD 1184717 • 3 ♂♂, 7 ♀♀; 7 ° 49 ′ 3.6114 ″ S, 36 ° 57 ′ 33.8754 ″ E; 280 m a. s. l.; 16 Mar. 2020; A. Ngute and A. R. Marshall leg.; closed forest, Micro-FoRCE plot 27; NHMD 1184698 • 1 ♂, 2 ♀♀, 4 juvs; 7 ° 48 ′ 45.8634 ″ S, 36 ° 58 ′ 38.4954 ″ E; 274 m a. s. l.; 27 Feb. 2018; A R. Marshall leg.; open forest, Half-FoRCE plot 33; NHMD 1184718 • 1 ♂, 1 juv.; 7 ° 48 ′ 34.0554 ″ E, 36 ° 59 ′ 4.4514 ″ E; 281 m a. s. l.; 2 Feb. 2018; A. R. Marshall leg.; open forest, Half-FoRCE plot 36; NHMD 1184719. – Morogoro Region, Kilombero District, Udzungwa Mountains National Park, Mwanihana • 1 ♀; Njokamoni; 7 ° 50 ′ 31.8474 ″ S, 36 ° 52 ′ 47.1354 ″ E; 401 m a. s. l.; 21 Dec. 2021; A. Ngute, R. Malanda, W. Mhagawale and A. Marshall leg.; open canopy, ForCE Plot 11; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184720 • 1 ♂, 2 ♀♀, 1 juv.; same data as for preceding; 7 ° 50 ′ 33.1434 ″ S, 36 ° 52 ′ 54.912 ″ E; 357 m a. s. l.; 22 Dec. 2021; open canopy, ForCE Plot 12; NHMD 1184721 • 1 ♂, 1 ♀; Sanje Kati camp and plot; 7 ° 45 ′ 47.6 ″ S, 36 ° 53 ′ 10.4 ″ E; 850 m a. s. l.; 20 – 23 Jan. 2014; T. Pape and N. Scharff leg.; in copula, hand-collected; NHMD 1184730 • 1 ♂; Sanje Chini camp; 7 ° 46 ′ 24.6 ″ S, 36 ° 53 ′ 47.7 ″ E; 598 m a. s. l.; 17 – 19 Jan. 2014; T. Pape and N. Scharff leg.; hand-collected; NHMD 1184731 • 1 ♂, 3 ♀♀, 5 juvs; Sanje; 7 ° 46 ′ 43.3914 ″ S, 36 ° 54 ′ 4.32 ″ E; 504 m a. s. l.; 26 Feb. 2022; A. Ngute, E. Kivambe, R. Malanda, H. Mnendendo, W. Mhagawale, M. Mpoto and A. Marshall leg.; open canopy, ForCE Plot 24; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184722 • 4 ♂♂, 1 ♀; Sanje; 7 ° 46 ′ 52.14 ″ S, 36 ° 54 ′ 11.7714 ″ E; 421 m a. s. l.; 20 Feb. 2022; A. Ngute, E. Kivambe, R. Malanda, W. Mhagawale, H. Mnendendo and A. Marshall leg.; open canopy. ForCE Plot 92: COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184661. – Morogoro Region, Kilombero District, Udzungwa Mountains National Park • 8 ♂♂, 11 ♀♀, 1 juv.; 7 ° 49 ′ 13.2198 ″ S, 36 ° 53 ′ 33.4608 ″ E; 334 m a. s. l.; 25 Feb. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 17; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184723 • 1 ♂; 7 ° 50 ′ 14.5314 ″ S, 36 ° 53 ′ 7.7634 ″ E; 358 m a. s. l.; 13 Feb. 2020; A. Ngute and A. R. Marshall leg.; closed forest, Micro-FoRCE plot 18; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184724 • 5 ♂♂; 7 ° 49 ′ 30.8424 ″ S, 36 ° 53 ′ 29.1798 ″ E; 340 m a. s. l.; 21 Feb. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 19; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184725 • 1 ♂; 7 ° 49 ′ 46.5348 ″ S, 36 ° 53 ′ 27.873 ″ E; 347 m a. s. l.; 27 Feb. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 20; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184726 • 2 ♀♀; 7 ° 47 ′ 24.216 ″ S, 36 ° 54 ′ 7.128 ″ E; 334 m a. s. l.; 17 Feb. 2020; A. Ngute and A. R. Marshall leg.; open forest Micro-FoRCE plot 25; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184727 • 1 ♂, 1 ♀; 7 ° 47 ′ 43.1088 ″ S, 36 ° 53 ′ 57.6132 ″ E; 331 m a. s. l.; 19 Feb. 2020; A. Ngute and A. R. Marshall leg.; open forest, Micro-FoRCE plot 28; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184728 • 3 ♀♀, 1 juv.; 7 ° 47 ′ 1.1394 ″ S, 36 ° 54 ′ 13.8594 ″ E; 381 m a. s. l.; 15 Feb. 2020; A. Ngute and A. R. Marshall leg.; closed forest, Micro-FoRCE plot 30; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184729.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	description	Descriptive notes The original description of T. howelli, including the accompanying drawings (Hoffman 1977), are fully satisfactory for recognizing the species. Study of the abundant material from the Udzungwa Mts allows, however, some additions to the description. SIZE. Max. body width of males 4.3 – 4.7 mm (Hoffman 1977: 4.6 mm). COLOUR (Fig. 2 C – D). Overall colour of freshly preserved specimens (Fig. 2 C) reddish brown, with contrasting yellowish paraterga. After four years in alcohol as described by Hoffman (1977), except that the colour of “ middorsum of metaterga, most of prozona, and sides of metazona ” is dark piceous brown in males, and medium brown in females, rather than “ piceous black ”. COLLUM (Fig. 13 A). Same shape as in Morogorius spp.: an isosceles curved trapezoid with the longer margin anteriorly. A few setae close to anterior margin. BODY RINGS (Fig. 13 B – D). Metazonites regularly granulotuberculate dorsally. Microsculpture of paranota (Fig. 13 C) as in Morogorius spp.: formed by longitudinally stretched, narrow cytoscutes of more or less irregular shape. Limbus (Fig. 13 D) as in Morogorius spp. TELSON (Fig. 13 E). Spinnerets arranged in an almost quadratic trapezoid on a smooth terminal swelling of the epiproct, simple, flanked by 1 + 1 globular, sometimes partly collapsed swellings. GONOPODS (Fig. 14) (interpreted in the light of Morogorius spp.). Prefemoral part (prf) longer than in Morogorius spp. Prefemoral process (prp) (“ tibiotarsus ” of Hoffman 1977) a narrow band curving over convex surface of solenomere, apically with two small tines. Solenomere (slm) (solenomerite and acropodite of Hoffman 1977) a very large and broad sheet, dorsally deeply concave. Parasolenomere (ps) a thin, straight rod.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	distribution	Distribution and habitat The type locality is “ Kisaware, 6.63 ° S, 39.05 ° E, near Dar es Salaam. ”. This is probably a typo for Kisarawe, a district in Tanzania’s Pwani Region. Also known from the Magombera Nature Reserve and from several sites in the Udzungwa Mountains National Park. Collected in open and closed forest at 267 – 504 m a. s. l., and a single outlier at 850 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA3FFCCB538A238FB2B7B66.taxon	discussion	Remarks The relatively high abundance of T. howelli in the Udzungwa Mts contrasts with the situation at the type locality near Dar es Salaam where, according to Hoffman (1977: 81), “ The species does not appear to be abundant (or conspicuous), as two specimens only have been found by Dr Howell over a period of “ several years of collecting in the region of Dar es Salaam ”. See also Discussion below.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFCCB55BA027FEEB7992.taxon	description	urn: lsid: zoobank. org: act: 2 F 0 BBC 98 - DA 48 - 46 A 0 - 80 FA-A 5 C 375507 E 14	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFCCB55BA027FEEB7992.taxon	type_taxon	Type species Dionaeadesmus force Enghoff gen. et sp. nov.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFCCB55BA027FEEB7992.taxon	diagnosis	Diagnosis Dionaeadesmus gen. nov. differs from other Tanzanian genera of Chelodesmidae as shown in Table 2, notably by the large spiky postfemoral process (prp, Figs 16 E – F, 17 A – D).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFCCB55BA027FEEB7992.taxon	etymology	Etymology Named after the Venus fly-trap Dionaea muscipula J. Ellis because the pair of heavily spiked gonopod prefemoral processes may remind of the spike-edged leaves of this carnivorous plant.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	description	urn: lsid: zoobank. org: act: EDE 39835 - BCF 3 - 4 A 30 - A 5 EC- 3 FF 129 D 9 CB 29	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	materials_examined	Figs 1, 2 E, 15 – 17	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	diagnosis	Diagnosis Redundant, genus monotypic.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	etymology	Etymology Named after the FoRCE project (see Material and methods). Noun in apposition.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	materials_examined	Material examined (total 6 ♂♂, 9 ♀♀) Holotype TANZANIA • ♂; Iringa Region, Kilolo District, Kilombero Nature Reserve, Ndundulu, Luala; 7 ° 46 ′ 00.4 ″ S, 36 ° 29 ′ 33.2 ″ E; 1903 m a. s. l.; 26 Oct. 2021; A. Ngute, R. Malanda, W. Mhagawale, A. Mpoto and A. Marshall leg.; FoRCE Plot 70, closed canopy, litter sifting; COLL. NHMD – ACC. NO. 2022 - EN- 003; NHMD 1184575. Paratypes TANZANIA • 2 ♂♂, 4 ♀♀; same data as for holotype; Ndundulu, Chawemba; 7 ° 44 ′ 54.4 ″ S, 36 ° 29 ′ 02.0 ″ E; 2256 m a. s. l.; 8 Nov. 2021; FoRCE Plot 81, open canopy; NHMD 1184562 • 1 ♂, 1 ♀; Iringa Region, Mahenge District, West Kilombero Scarp Forest Reserve, Nyumbanitu Mts, S Udekwa Village; 07 ° 48 ′ S, 36 ° 21 ′ E; 2300 m a. s. l.; Dec. 1993; J. O. Svendsen leg.; NHMD 1184576 • 2 ♂♂, 4 ♀♀; same data as for preceding; 1700 m a. s. l.; L. L. Sørensen and J. O. Svendsen leg.; NHMD 1184572.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	description	Specimens from Luala are very fragile. Description (males) SIZE. Length 22 – 25 mm, max. width 4.6 mm. COLOUR. Specimens from Luala after 3 years in alcohol completely bleached, whitish. Specimens from Nyumbanitu Mts (Fig. 2 E) after 31 years in alcohol: head and dorsum medium brown, lateral lobes of collum narrowly whitish, paranota of rings 2 – 3 and of poriferous rings broadly whitish, paranota of non-poriferous rings same colour as rest of ring; antennae, venter, legs and telson light brownish. HEAD (Fig. 15 A – B). Labrogenal offset distinct, otherwise as Morogorius. Antennae reaching back to ring 5 when stretched. COLLUM (Fig. 15 B). As in Morogorius: an isosceles curved trapezoid with the longer margin anteriorly, lateral corners smoothly rounded, surface granular, especially laterally. Two setae close to anterior margin. BODY RINGS (Fig. 15 C – E). Prozonites smooth, with simple, cellular microsculpture. Metazonites flat, granular dorsally and laterally, ~ 1.4 × as broad as prozonites, apparently without setae, with a prominent transverse sulcus on rings 5 – 17. Paranota horizontal, those of rings 2 – 3 slightly projecting forwards, rectangular until ring 12 – 14, thereafter posterior corners projecting increasingly backwards, triangular. Paranotal microsculpture (Fig. 16 C) formed by cytoscutes of irregular shape, narrow or amoeba-like branched. Ozopore formula normal (5, 7, 9 – 10, 12 – 3, 15 – 19), ozopores circular, delimited by smooth ring in smooth peritremata on edge of paranota, ca midway between anterior and posterior margin; peritremata protruding from lateral margin of paranota. Sterna broad, with a transverse impression interrupted in the middle and extensive setation along anterior and posterior margins. Sides of body rings (Fig. 15 D) with a row of small tubercles along posterior margin. Limbus (Fig. 16 D) with straight margin, consisting of a row of rectangular cells with a dense fringe of threadlike ridges ending in free processes. LEGS (Fig. 15 G). Rather stout, without modifications, length ~ 1.3 × maximum body width, length not increasing towards hind end, relative (percentual) length of podomeres (coxa-claw): 8 / 14 / 23 / 14 / 14 / 24 / 2. T ELSON (Fig. 16 A – B). Preanal ring with several setae along dorsal part of posterior margin; epiproct large, with prominent lateral setiferous tubercles; spinnerets arranged in a trapezoid on a smooth terminal swelling, simple, flanked by 1 + 1 globular, swellings. Anal valves (paraprocts) unmodified, each with 2 setae, dorsalmost seta on marginal rim, ventralmost seta more lateral, Subanal scale (hypoproct) semicircular, unmodified, with 1 + 1 marginal setae. GONAPOPHYSES. Unconspicuous. GONOPOD APERTURE. Transversely oval, more than twice as broad as long, rim simple. GONOPODS (Figs 16 E – F, 17). No sternal remnant. Coxa (cx) cylindrical, length slightly> diameter, with a strong conical process (cxp) distally on the anterior side; two long setae (as) on anterior surface basal to process, a few setae on lateral surface, and a field of ~ 15 – 20 setae on meso-posterior surface. Cannula (ca) stout. Prefemoral part (prf) continuing in same axis as cx, conical, short, length> 1 ½ × basal diameter, with numerous long setae on ventral and mesal surfaces and a small tooth (prt) distomesally. Efferent canal running on meso-dorsal side of prf, at level of prefemoral tooth (prt) bending laterad into solenomere. A huge prefemoral process (prp) originating dorso-laterally from prf, together with solenomere (slm) forming right angles with main body of prf, longer than solenomere (slm) and accommodating slm in mesal concavity; overall shape of prp like a twisted spoon with a stout shaft, apically bending mesad over tip of slm and ending in a point; dorsal margin of prp with numerous (~ 20) spikelike processes (msp); mesal surface with a spiked ridge (spr) and additional scattered spikes. Solenomere (slm) in the shape of a ribbon with rolled-in edges, apically dividing into a slender, pointed, curved, ridged branch (the solenomere s. str., sslm) and a broader, simple, pointed branch which may be regarded as a solenophore (sph). A small rectangular process near the base of slm may be regarded as a parasolenomere (ps).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	distribution	Distribution and habitat Only known from two sites in the Udzungwa Mountains: Nyumbanitu Mountains and Ndundulu Forest (Fig. 1). Collected at 1700 – 2300 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FFA7FFF1B577A291FE94793D.taxon	discussion	Remarks A structure similar to the complicated prefemoral process (prp) of Dionaeadesmus force gen. et sp. nov. occurs in Prepodesmus laciniosus (Attems, 1937), of uncertain generic affiliation (Demange & Mauriès 1975) from Congo. This species, however, differs markedly from D. force in having a very short, simple, pointed solenomere (Attems 1937: “ Rinnenast ”) and a two-branched solenophore (Attems 1937: “ Tibiotarsus ”), and the two taxa are probably not closely related. See also Chamberlin (1952) (as Morphotelus l.).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF1B534A20DFED57C6F.taxon	type_taxon	Type species Callistocilla beatrix Hoffman, 1977. KENYA, Tiwi S of Mombasa.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF1B534A20DFED57C6F.taxon	discussion	Remarks Hoffman (1977: 71) wrote “ This genus is based upon one of the most disjunct and interesting chelodesmoids so far known to me ” and “ Actually, had the specimen been labelled with a Brasilian locality I should have experienced less difficulty in reconciling it with existing group ”. These words, coming from the all-time number one authority on the huge family Chelodesmidae, testify to the isolated position of Callistocilla. As in so many other millipede groups, a phylogenetic analysis including molecular data is loudly called for. Here, the genus as such will not be discussed further, but notes on the Udzungwan species C. dolorotrix are given. The gonopod terminology of Hoffman (2005) will be employed.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	description	Figs 1, 2 F, 18 – 19, 20 A – B	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	diagnosis	Diagnosis Differing from congeners by the presence of ozopores on body rings 5, 7, 9 – 10, 12 – 13 and 15 – 19, i. e., the by far commonest ‘ pore formula’ in Polydesmida (vs ozopores absent in congeners), as well as by the spatulate shape of the gonopodal prefemoral process and the reflexed distal part of the parasolenomere (vs prefemoral process differently shaped and parasolenomere not reflexed in congeners).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	materials_examined	Material examined (total 12 ♂♂, 9 ♀♀, 2 juv.) TANZANIA – Morogoro Region, Kilombero District, Udzungwa Mountains National Park • 1 ♀; Njokamoni; 7 ° 50 ′ 31.9 ″ S, 36 ° 52 ′ 471 ″ E; 401 m a. s. l.; 21 Dec. 2021; A. Ngute, R. Malanda, W. Mhagawale and A. Marshall leg.; FoRCE Plot 11, closed canopy; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184735 • 1 ♂, 1 ♀; Sanje; 7 ° 46 ′ 43.4 ″ S, 36 ° 54 ′ 04.3 ″ E; 504 m a. s. l.; 26 Feb. 2022; A. Ngute, E. Kivambe, R. Malanda, H. Mnendendo, W. Mhagawale, M. Mpoto and A. Marshall leg.; FoRCE Plot 24, open canopy; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184736 • 3 ♂♂, 1 ♀; same data as for preceding; 7 ° 46 ′ 47.6 ″ S, 36 ° 54 ′ 07.6 ″ E; 495 m a. s. l.; 23 Feb. 2022; FoRCE Plot 25, open canopy; NHMD 1184737 • 2 ♀♀; same data as for preceding; 7 ° 46 ′ 52.1 ″ S, 36 ° 54 ′ 11.8 ″ E; 421 m a. s. l.; 20 Feb. 2022; A. Ngute, E. Kivambe, R. Malanda, W. Mhagawale, H. Mnendendo and A. Marshall leg.; FoRCE Plot 92, open canopy; NHMD 1184738 • 2 ♂♂, 1 ♀, 1 juv.; same data as for preceding; Sanje; 7 ° 47 ′ 03.0 ″ S, 36 ° 54 ′ 05.8 ″ E; 410 m a. s. l.; 17 Feb. 2022; A. Ngute, E. Kivambe, W. Mhagawale, H. Mnendendo, M. Mpoto and A. Marshall leg.; FoRCE plot 93, open canopy; NHMD 1184739 • 1 subad. ♀; Udzungwa Mountains National Park; 07 ° 50 ′ 56.4 ″ S, 36 ° 53 ′ 00.4 ″ E; 325 m a. s. l.; 2 Mar. 2020; A. Ngute and A. R. Marshall leg.; micro-FoRCE plot 26, open forest; COLL. NHMD. ACC. NO. 2020 - EN- 002; NHMD 1184740. – Iringa Region, Kilolo District, Udzungwa Scarp Nature Reserve • 2 ♂♂; Uhafiwa; 8 ° 31 ′ 58.4 ″ S, 35 ° 51 ′ 28.4 ″ E; 1375 m a. s. l.; 25 Jan. 2022; A. Ngute, R. Malanda, W. Mhagawale, H. Mnendendo, A. Mpoto and A. Marshall leg.; FoRCE Plot 106, open canopy; COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184741 • 3 ♂♂, 3 ♀♀; Chita Juu; 8 ° 32 ′ 23.4 ″ S, 35 ° 51 ′ 36.0 ″ E; 1387 m a. s. l.; 1 Feb. 2022; A. Ngute, R. Malanda, W. Mhagawale, H. Mnendendo, A. Mpoto and A. Marshall leg.; FoRCE Plot 109, open canopy, COLL. NHMD - ACC. NO. 2022 - EN- 003; NHMD 1184742 • 1 ♂; 11 km SE of Masisiwe, Kihanga Stream, 8 ° 22 ′ 5.7 ″ S, 35 ° 58 ′ 41.6 ″ E, 1800 m a. s. l.; 17 – 27 May 1997; E. Milungu leg.; understorey, NHMD 1184743.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	distribution	Distribution and habitat Known from two sites in the Udzungwa Mountains National Park (Fig. 1) – collected at 325 – 504 m a. s. l., and three sites in the Udzungwa Scarp Nature Reserve – collected at 1373 – 1800 m a. s. l.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9AFFF2B54BA73FFD4E7DE6.taxon	discussion	Remarks The examined specimens agree with the description by Hoffman (2005). However, there are a few differences between specimens from Udzungwa Mountains National Park (topotypes and near topotypes) and specimens from Udzungwa Scarp Nature Reserve, see Table 3 and Figs 18 – 20). The specimens from the latter place were collected at much higher altitudes than those from the former. The microsculpture of the paranota (Fig. 18 E) is remarkable: the cuticular microscutes (“ cytoscutes ”) are rounded-hexagonal, ca 0.01 mm in diameter, and densely beset with tiny denticles, not only along one edge, as is very common in myriapods, but all over the surface; 74 – 83 denticles on each of the three central scutes on Fig. 18 E. Similar microsculpture was also seen in C. beatrix Hoffman, 1977 (Fig. 20 F) and in an undescribed congener from Kanga Montauns (Fig. 20 E). See Discussion. The spinnerets were not described by Hoffman (2005). The four simple, setiform sensilla are placed in a quadrat at the tip of the epiproct. On Fig. 18 C there is what looks somewhat, but not quite like the socket of a fifth spinneret; this structure was seen only on one of two specimens studied with scanning electron microscopy and may be insignificant. Unlike the other Udzungwa chelodesmids, C. dolorotrix has no globular swelling lateral to the spinnerets. The gonopods (Figs 19, 20 B) fully agree with Hoffman’s description and drawings. In males from Udzungwa Mountains National Park, the proximal edge of the solenomere is flexed at right angles with main plane of the solenomere (Fig. 20 B).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B539A24EFB5C7E8D.taxon	description	Fig. 20 F	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B539A24EFB5C7E8D.taxon	materials_examined	Material examined (total 1 ♂) Holotype KENYA • ♂; Tiwi S of Mombasa; 18 May 1968; Schiøtz leg.; ZMUC 00101515.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B539A24EFB5C7E8D.taxon	discussion	Remarks A small piece of one body ring was removed from the unique holotype and mounted for SEM. The paranotal microsculpture (Fig. 20 F) is similar to that seeen in C. dolotrotrix (Fig. 18 E)	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B5CDA59DFE6E7D1B.taxon	description	Fig. 20 C – E	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B5CDA59DFE6E7D1B.taxon	materials_examined	Material examined (total 1 ♂, 2 ♀♀) TANZANIA • 1 ♂, 2 ♀♀; Morogoro Region, Kanga Mts, Kanga Forest Reserve; 400 – 500 m a. s. l.; 22 – 25 Nov. 1984; N. Scharff leg.; lowland rain forest; NHMD 1184571.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B5CDA59DFE6E7D1B.taxon	discussion	Remarks These specimens agree closely with the original description of C. beatrix (Hoffman 1977), except for the presence of very small ozopores (Fig. 20 C – D) and tiny small gonopod details. This species, too, shares the peculiar paranotal microsculpture seen in C. dolorotrix and C. beatrix (Fig. 20 E).	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
D924C438FF9DFFF6B5CDA59DFE6E7D1B.taxon	diagnosis	Identification of Chelodesmidae from the Udzungwa Mountains As mentioned in the introduction, no useful diagnosis of the family Chelodesmidae is available. For identification of chelodesmids from the Udzungwa Mts it is therefore necessary to single out differences between this family and each other polydesmidan family occurring in the Udzungwas. Table 4 provides this information.	en	Enghoff, Henrik, Ngute, Alain Senghor K., Mnendendo, Hamidu R., Kivambe, Exaud E., Mhagawale, Waziri, Malanda, Robin C., Mpoto, Athuman, Marshall, Andrew R. (2025): A mountain of millipedes XII. The Chelodesmidae of the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida). European Journal of Taxonomy 997: 210-255, DOI: 10.5852/ejt.2025.997.2935, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2935/13261
