taxonID	type	description	language	source
DC5287DAFF90FFD5FF01100DFDEACAAC.taxon	materials_examined	Material examined: GEORGIA • 1 ♂, 2 ♀ (ISUIE); Kobuleti; N 41.80473 °, E 41.88427 °; 92 m a. s. l.; 10 Aug. 2018; leg. GGBC team; hand collecting; humid lowlands; CaBOL IDs 1022917, 1022918, 1022916, and 1022919 (BGEIDs 0659 _ C 06, 00659 _ C 09, 00659 _ H 11, and 00660 _ A 01). Barcoding: Two nearly identical sequences were obtained from the specimens with CaBOL IDs 1022917 and 1022916 (BGE-IDs 0659 _ C 06 and 00659 _ H 11) (BOLD: AGT 2788, p - distance 0.15 %). A single sequence obtained from Georgian C. caucasicus (Šilhavý, 1966) (BOLD: AGT 2789) exhibited 9.57 % divergence (p - distance) from the COI sequences of C. minutus. There were no sequences for C. minutus and C. caucasicus in BOLD Systems, as we submitted the first ones.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF90FFD5FF01100DFDEACAAC.taxon	discussion	Remarks: In neighboring territories, C. minutus was reported from Russia (Snegovaya & Chumachenko 2011). Previously considered endemic to the Northwest Caucasus, it is now recognized as endemic to the Caucasus region based on the herein published data.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF92FFD5FF0115E6FCA6CF52.taxon	materials_examined	Material examined: GEORGIA • 1 ♂, (ISUIE); Dmanisi; N 41.363655 °, E 44.36486 °; 250 m a. s. l.; 20 Nov. 2023; leg. A. Memishishi; hand collecting; semi-arid mixed forest; CaBOL ID 1022671 (BGE-ID 00673 _ E 10). Barcoding: A single sequence was obtained from the specimen with CaBOL ID 1022671 (BGE-ID 00673 _ E 10) (BOLD: AGU 9535). The divergence of 6.53 % (p - distance) exhibited between the COI sequences of Georgian and Turkish populations indicates the presence of possible cryptic species. Further material sampling is needed.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF92FFD5FF0115E6FCA6CF52.taxon	discussion	Remarks: In neighboring territories, O. lendlii was previously recorded from Georgia, Turkey, Azerbaijan, and SW Russia (Staręga 1966, 1978; Snegovaya 2013). In Georgia, the species was previously recorded from the regions of Shida Kartli (Gori) and Samtskhe-Javakheti (Borjomi, Tsaghveri) (Modebadze et al. 2023); here, it is recorded in the Kvemo Kartli region (Dmanisi) for the first time.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF92FFD5FF011385FD3FCD8F.taxon	materials_examined	Material examined: GEORGIA • 3 ♂, (ISUIE); Paravani Lake vicinity; N 41.29411 °, E 43.73097 °; 2010 m a. s. l.; 19 May. 2024; leg. G. Iankoshvili; hand collecting; alpine meadow; CaBOL IDs 1037895, 1037896, and 1037897 (BGE-IDs 00654 _ G 04, 00654 _ G 05, and 00654 _ G 06). Barcoding: We obtained a single sequence from the specimen CaBOL-ID 1037895 (BGE-IDs 00654 _ G 04) (BOLD: AGQ 075). There were no sequences for P. mcheidzeae in BOLD Systems, as we submitted the first ones.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF92FFD5FF011385FD3FCD8F.taxon	discussion	Remarks: Phalangium mcheidzeae was previously recorded from a single location in Georgia (Snegovaya 2014); here we report it for the second time from the country and for the first time from the vicinity of Paravani Lake, suggesting a broader distribution of the species.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF92FFD5FF011188FD63CC42.taxon	type_taxon	Type species. Rilaena balcanica Silhavý, 1965	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	description	zoobank. org: act: 0 ACC 260 D- 27 A 3 - 4 A 4 F-B 2 BA-F 6 A 04 AE 9 D 0 ED Figs 3 – 6	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	materials_examined	Holotype. 1 ♂ (CaBOL ID 1038667): Georgia: Mtskheta-Mtianeti, Abudelauri Lake vicinity; N 42.5481 °, E 44.8803 °; 2240 m a. s. l.; leg. K. Dzuliashvili; hand collecting; subalpine meadow; 9 Oct. 2024. Paratypes. 6 ♂ (CaBOL-ID 1038668, 1038669, 1038670, 1038671, 1038672, and 1038673): same data as holotype. Barcoding: We submit the first five identical sequences of R. kopala sp. nov. to the BOLD Systems, obtained from the specimens CaBOL IDs 1038667, 1038668, 1038669, 1038670, and 1038671 (BGE-IDs 00665 _ H 04, 00665 _ H 05, 00665 _ H 06, 00665 _ H 07, and 00665 _ H 08) (BOLD: AGS 3235).	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	diagnosis	Diagnosis. Males of Rilaena kopala sp. nov. are most similar in general body and penis shapes to those of R. kasatkini Snegovaya, Cokendolpher & Mozaffarian, 2018 and R. kaschkataschica from Iran and Russia, respectively. From R. kasatkini they differ in lack of longitudinal ridge on eye tubercle (Fig. 1) (vs. present; Snegovaya et al. 2018: fig. 2), cephalothorax posteriorly with spines (Figs 1 – 2) (vs. posteriorly with denticles; Snegovaya et al. 2018: fig. 2), palpal Fe with ventral spines as prominent as dorsals (Fig. 6) (vs. dorsal denticles larger than ventrals; Snegovaya et al. 2018: fig. 2), BLI 2.18 (vs. 0.75), and dorsally convex glans with left-looking stylus (Figs 3 – 4) (vs. banana-form; Snegovaya et al. 2018: fig. 2). From R. kaschkataschica they differ in having more prominent denticles on legs (vs. less prominent; Snegovaya et al. 2025: fig. 2), cephalothorax posteriorly with spines (Figs 1 – 2) (vs. less prominent; Snegovaya et al. 2025: fig. 2 A), BLI 2.18 (vs. 1.04), palpal Fe with ventral spines as prominent as dorsals (Fig. 6) (vs. less abundant and spine-shaped; Snegovaya et al. 2025: figs 2 D – E), prominent denticles on basal and distal segments of chelicera (Fig. 5) (vs. less prominent; Snegovaya et al. 2025: figs 2 F – G), and dorsally convex glans with left-looking stylus (Figs 3 – 4) (vs. triangular-shaped glans and straightforward stylus; Snegovaya et al. 2025: figs 2 H – K).	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	description	Description. Male (Holotype). Total length 4.34 mm long, 3.04 mm wide. Eye tubercule 0.62 mm long, 0.83 mm wide, with 12 - 15 pointy denticles. Body dark-brown, with small light spots. Chelicerae simply shaped, with numerous spine-like denticles (Fig. 5). Cheliceral basal segment 1.31 mm long, distal segment 2.19 mm long. Penis medium size, 2.54 mm long; base wide, tapering towards neck, slightly expanded towards glans; glans 0.31 mm long, stylus 0.16 mm long. Measurements of leg and pedipalp segments as in Table 1. Variation (n = 7 ♂): Body 5.41 – 3.93 long, 3.88 – 2.73 wide. Penis 3.25 – 2.43 long, glans 0.32 – 0.27, stylus 0.18 – 0.17. Eye tubercle with 9 – 16 spines. Female. Unknown.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	etymology	Etymology. The new species is named after Kopala, a Khevsuretian mythological hero, honoring the cultural heritage of the region where the species was discovered.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF95FFD3FF0117FFFD3AC8EC.taxon	distribution	Distribution. Known from the type locality only.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF97FFD0FF011554FC79CF24.taxon	materials_examined	Material examined: GEORGIA • 1 ♂, (ISUIE); Gurjaani; N 41.8325 °, E 45.8532 °; 250 m a. s. l.; 13 Jul. 2018; leg. L. Ninua; hand collecting; CaBOL-ID 1035654 • 1 ♂, (ISUIE); Erisimedi; N 41.5855 °, E 46.3325 °; 190 m a. s. l.; 17 Jul. 2018; leg. G. Iankoshvili; hand collecting; CaBOL-ID 1035655. • 1 ♂, (ISUIE); Lagodekhi NP; N 41.874 °, E 46.357 °; 1940 m a. s. l.; 17 Jul. 2022; leg. B. Chitadze; hand collecting; BGE-ID 00665 _ G 10. Barcoding: Two sequences were obtained from the specimens with CaBOL-IDs 1035654 and 1035655, BGE-ID 00665 _ G 10 (BOLD: AGS 2943, maximum p - distance 0.46 %). There were no sequences for Z. shachdag in BOLD Systems, as we submitted the first ones.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
DC5287DAFF97FFD0FF011554FC79CF24.taxon	discussion	Remarks: Zachaeus shachdag, described from Azerbaijan (Snegovaya & Starega 2008), was previously regarded as endemic to that country and is currently reassessed as endemic to the Caucasus. Here, we record it for the first time in Georgia, extending its known distribution further west.	en	Chitadze, Beka, Dzuliashvili, Kristina, Seropian, Armen (2025): New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa 5725 (4): 591-599, DOI: 10.11646/zootaxa.5725.4.9, URL: https://doi.org/10.11646/zootaxa.5725.4.9
