taxonID	type	description	language	source
122556AB11275C5F9A5A7D663D1F982D.taxon	description	Fig. 5	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
122556AB11275C5F9A5A7D663D1F982D.taxon	etymology	Etymology. Epithet composed of the name of the host genus and the Latin-derived suffix “ - cola ” (dweller).	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
122556AB11275C5F9A5A7D663D1F982D.taxon	diagnosis	Diagnosis. Morphologically distinguished from Erysiphe glycines by having chasmothecia with shorter appendages, up to twice as long as the chasmothecial diameter (versus up to seven times as long as the diameter in E. glycines), and phylogenetically by forming a distant highly supported clade.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
122556AB11275C5F9A5A7D663D1F982D.taxon	description	Description. Mycelium amphigenous, forming effuse, arachnoid, whitish patches; hyphae branched, often at right angles, septate, hyaline, thin-walled, smooth, 3 – 5 µm wide; hyphal appressoria solitary, nipple-shaped to lobate, 4 – 10 µm wide; conidiophores 65 – 80 µm long, septate at base, foot cells 24 – 42 µm long and 6 – 8 µm wide, straight to usually somewhat flexuous, sinuous, followed by 1 – 2 shorter cells, about 15 – 20 × 7 – 8 µm conidia formed singly, cylindrical-doliiform, 23 – 35 × 10 – 12 µm, germination not seen. Chasmothecia scattered, subglobose to globose, dark brown, 90 – 154 × 104 – 157 µm in diameter; peridium cells irregularly polygonal, 6 – 13 × 10 – 19 µm; appendages in the lower half of the chasmothecium, number variable, few to numerous, mycelioid, often interwoven with the mycelium and with each other, sometimes poorly developed and hard to distinguish from the mycelial hyphae, length variable (up to two times the diameter of the chasmothecium), thin-walled, smooth, narrow (up to 4 µm wide), septate, hyaline; asci 4 – 6 per chasmothecium, 63 – 70 × 38 – 45 µm, oblong-ellipsoid, short-stalked, 6 - spored; ascospores ellipsoid-ovoid, 20 – 22 × 11 – 12 µm, colorless.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
122556AB11275C5F9A5A7D663D1F982D.taxon	distribution	Distribution. North America (Canada, USA), probably widespread.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
4DCFE85755435F61830405D421F5FF72.taxon	description	Fig. 6	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
4DCFE85755435F61830405D421F5FF72.taxon	etymology	Etymology. Epithet derived from the name of the host plant, Quercus virginiana.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
4DCFE85755435F61830405D421F5FF72.taxon	diagnosis	Diagnosis. Morphologically close to Erysiphe abbreviata s. lat., but differing by forming much larger chasmothecia, 116 – 159 µm diam., with up to 20 appendages, and 4 – 8 - spored asci. Phylogenetically well-distinguished from E. abbreviata and all other North American Erysiphe spp. on oaks by forming a highly supported clade.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
4DCFE85755435F61830405D421F5FF72.taxon	description	Description. Mycelium and anamorph not seen. Chasmothecia scattered to gregarious among trichomes on abaxial leaf surfaces, subglobose to globose, 116 – 159 × 123 – 144 µm; peridium cells conspicuous, brown, irregularly polygonal, 12 – 21 × 7 – 14 µm; appendages 8 – 20, equatorial, stiff, straight to somewhat curved, aseptate, hyaline, 60 – 125 µm long, relative length usually about 0.5 – 1 times the chasmothecial diameter or somewhat shorter, 4 – 7 µm wide [widest at base], apices 4 – 5 × regularly dichotomously branched not strictly in one dimension, tips of the ultimate branchlets recurved; asci 5 – 8 per chasmothecium, obovoid, saccate, short-stalked, 55 – 75 × 40 – 60 µm, walls up to 3 µm thick, 4 – 8 - spored; ascospores ellipsoid-ovoid, hyaline, 15 – 25 × 8 – 13 µm.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
4DCFE85755435F61830405D421F5FF72.taxon	distribution	Distribution. North America (USA, Florida, North Carolina).	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
55F4B9AFEC9259FB8CD6D5C14EEF9473.taxon	description	Fig. 7	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
55F4B9AFEC9259FB8CD6D5C14EEF9473.taxon	etymology	Etymology. Epithet referring to the name of the type host, Ulmus alata.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
55F4B9AFEC9259FB8CD6D5C14EEF9473.taxon	diagnosis	Diagnosis. Erysiphe ulmi-alatae is morphologically barely distinguishable from E. macrospora, but can be distinguished by the host and the fact that it phylogenetically forms a separate, highly supported clade.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
55F4B9AFEC9259FB8CD6D5C14EEF9473.taxon	description	Description. Mycelium in persistent, creamy-white patches, almost entirely on adaxial leaf surfaces; hyphae branched, often at right angles, septate, hyaline, 3 – 5 µm wide; hyphal appressoria solitary or in opposite pairs, lobed; conidia formed singly, cylindrical-doliiform, 34 – 40 × 11 – 18 µm. Chasmothecia scattered to gregarious, dark brown, subglobose to globose, 130 – 178 µm in diameter; peridium cells irregularly polygonal, 7 – 13 × 10 – 21 µm; appendages numerous, number variable (40 +), hyaline, aseptate, ± equatorial, 80 – 145 × 5 – 9 µm, mostly shorter than the chasmothecial diameter, width ± equal throughout, walls smooth, uniformly thickened from base to tip, apices uncinate to circinate when mature, uncinate-circinate apex not enlarged, circinate apices 10 – 15 µm across (appendages shorter, stiffer, and with pointed ends when immature); asci 10 – 25 per chasmothecium, obovoid, saccate, short-stalked, 48 – 70 × 23 – 24 µm, walls up to 3 µm thick, 2 - spored; ascospores ellipsoid-ovoid to slightly teardrop-shaped, 22 – 30 × 11 – 17 µm, hyaline.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
55F4B9AFEC9259FB8CD6D5C14EEF9473.taxon	distribution	Distribution. (based on specimens deposited in North American herbaria as ‘ Erysiphe macrospora ’ or ‘ Uncinula macrospora ’ on Ulmus alata): North America (USA: Alabama, Florida, Georgia, Illinois, Indiana, Mississippi, North Carolina, Oklahoma, South Carolina, Tennessee, Texas).	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
393024AF043A503CA5A8AAE92FB9805E.taxon	description	Fig. 9	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
393024AF043A503CA5A8AAE92FB9805E.taxon	etymology	Etymology. Epithet in honor of NCSU mycologist Larry F. Grand.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
393024AF043A503CA5A8AAE92FB9805E.taxon	diagnosis	Diagnosis. Takamatsuella grandii differs morphologically from T. circinata in having appendages with walls uniformly 3 µm thick, and genetically by forming a highly supported clade.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
393024AF043A503CA5A8AAE92FB9805E.taxon	description	Description. Mycelium on abaxial surfaces of leaves, effuse, thin, arachnoid, grayish white; hyphae dichotomously branched, hyaline, thin-walled, smooth, septate; hyphal appressoria nipple-shaped; anamorph not seen. Chasmothecia scattered to + / - gregarious, depressed globose, 120 – 170 µm in diameter; peridium cells irregularly polygonal, light brown, 7 – 15 × 14 – 19 µm; appendages very numerous, up to 150 per chasmothecium, arising below the equator, stiff to flexuous, simple, apices tightly uncinate to circinate, not enlarged, about 0.3 – 1 times as long as the chasmothecial diameter, uniformly 2 – 6 µm wide and walls uniformly 3 µm thick, hyaline, aseptate, smooth, thin-walled; asci up to 8 or more, clavate-saccate, 70 – 90 × 25 – 40 µm, usually stalked, wall uniformly 3 µm thick, 8 - spored; ascospores ellipsoid-obovoid, 10 – 25 × 9 – 20 µm, hyaline.	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
393024AF043A503CA5A8AAE92FB9805E.taxon	distribution	Distribution (based on specimens on Acer saccharum deposited in North American herbaria): North America (USA: Indiana, New York, New Hampshire, North Carolina, Pennsylvania).	en	LaGreca, Scott, Crouch, Uma, Paul, Andrew, Thomas, Jacklyn, Thompson, Jake, Shaw, Christian, Cubeta, Marc A., Braun, Uwe, Bradshaw, Michael (2025): Hidden treasures of herbaria - even small collections contain a wealth of diversity: the powdery mildews of the North Carolina State Larry F. Grand Mycological Herbarium. IMA Fungus 16: e 156231, DOI: 10.3897/imafungus.16.156231
