Key to the identification of the climbing plants of Igarassu based on fruiting material

1. Tendrils present, but not associated with the inflorescence.......................................................................................... 2

1. Tendrils absent, or where present just a simple branch .............................................................................................. 34

2. Leaves opposite............................................................................................................................................................. 3

2. Leaves alternate ............................................................................................................................................................ 6

3. Leaves simple ................................................................................................................................ Strychnos parviflora

3. Leaves compound ......................................................................................................................................................... 4

4. Leaflets bearing two dark glands at their base, branches of the inflorescence flattened in cross-section ...................... ............................................................................................................................................. Phryganocydia corymbosa

4. Leaflets without dark glands, branches of the inflorescence cylindrical in cross-section ............................................ 5

5. Fruits oblongoid, seeds not winged ................................................................................... Adenocalymma hypostictum

5. Frutis elongated, seeds winged .............................................................................................................. Lundia cordata

6. Leaves simple or 1-foliolate compound........................................................................................................................ 7

6. Leaves compound and with more than one foliole..................................................................................................... 26

7. Tendrils hook-like ......................................................................................................................................................... 8

7. Tendrils twining ............................................................................................................................................................ 9

8. Basal third of the leaflet connate; fruit indehiscent ..................................................................... Phanera trichosepala

8. Basal two-thirds of the leaflets connate; fruit dehiscent .................................................................. Phanera outimouta

9. Tendrils axillary to the leaves ..................................................................................................................................... 10

9. Tendril not axillary to the leaves................................................................................................................................. 19

10. Fruits winged, grouped in short (<20cm) racemes ............................................................................... Gouania virgata

10. Fruits berries, solitary, in pairs or long (> 20cm) racemes .......................................................................................... 11

11. Leaves entire ............................................................................................................................................................... 12

11. Leaves 2- or more lobbed ........................................................................................................................................... 14

12. Fruits grouped in racemes up to 1.5 m in length ................................................................................. Passiflora ovalis

12. Fruits solitary or in pairs............................................................................................................................................. 13

13. Stem 4-angulate in cross-section; petiole with 2-6 sessile glands ........................................................ Passiflora alata

13. Stem cylindrical in cross-section; petiole with 2 stalked glands ..................................................... Passiflora galbana

14. Leaves 5-lobbed ............................................................................................................................ Passiflora cincinnata

14. Leaves 2- or 3-lobbed ................................................................................................................................................. 15

15. Mature fruits never more than 15mm long ................................................................................................................. 16

15. Mature fruits always more than 15mm long ............................................................................................................... 17

16. Stem hairy, flattened in cross-section ................................................................................................. Passiflora misera

16. Stem glabrous, cylindrical in cross-section .................................................................................... Passiflora suberosa

17. Petiole with two sessile glands ............................................................................................................ Passiflora edulis

17. Petiole with 2-6 stalked glands ................................................................................................................................... 18

18. Apex of the leaves and stipules mucronate ................................................................................. Passiflora watsoniana

18. Apex of the leaves and stipules truncate ........................................................................................ Passiflora edmundoi

19. Tendrils opposite to the leaves (Fig. 3A) ........................................................................................... Cissus verticillata

19. Tendrils in the leaf sheath, at the basis of the inflorescence, or forming an acute angle with petiole (Fig. 3B, C, D, F, G) ................................................................................................................................................................................ 20

20. Tendrils two, in the apex of the leaf sheath (Fig. 3G) ....................................................................... Smilax syphilitica

20. Tendril solitary, in the leaf’s axils, forming an acute angle with the petiole or at the basis of the inflorescence (Fig. 3D) .............................................................................................................................................................................. 21 21. Leaves glabrous .......................................................................................................................................................... 22

21. Leaves hairy................................................................................................................................................................ 23

22. Latex watery; seeds arilate................................................................................................................. Psiguria umbrosa

22. Latex absent; seeds without an aril .............................................................................................. Melothria fluminensis

23. Mature fruits green or yellow striped ......................................................................................................................... 24

23. Mature entirely yellow to orange................................................................................................................................ 25

24. Petioles to 3 cm long; leaves scabrous on the adaxial surface.......................................................... Gurania spinulosa

24. Petioles to 15 cm long; leaves pubescent on the adaxial surface..................................................... Guranis acuminata

25. Mature fruit orange, dehiscent; seeds red .................................................................................... Momordica charantia

25. Mature fruit yellow (rarely red), indehiscent; seeds white ................................................................ Cayaponia tayuya

26. Tendrils borne opposite to the leaves (Fig. 3A).......................................................................................... Cissus erosa

26. Tendrils borne in the axils of the leaves or at the basis of the inflorescence (Fig. 3B, C, D, F, G)............................ 27

27. Fruits fleshy ................................................................................................................................................................ 28

27. Fruits dry..................................................................................................................................................................... 32

28. Leaves with 2 or 3 foliolules....................................................................................................................................... 29

28. Leaves with more than 3-foliolules ............................................................................................................................ 30

29. Plants glabrous; fruits solitary ........................................................................................................... Psiguria triphylla

29. Plants pubscent to scrabous; fruit borne in racemes ..................................................................... Gurania bignoniacea

30. Fruits without wings ........................................................................................................................... Paullinia pinnata

30. Fruit 3-winged............................................................................................................................................................. 31

31. Leaf margin entire; wings of the fruit obovate, apiculate................................................................ Paullinia racemosa

31. Leaf margin dentate; wings of the fruit elliptical, not apiculate ........................................................ Paullinia trigonia

32. Stem flattened in cross-section; leaves pinnate ........................................................................ Machaerium aculeatum

32. Stem cylindrical in cross-section; leaves 3-foliolate or ternate .................................................................................. 33

33. Fruit divided into three oblong-winged samaras ....................................................................... Serjania subinpunctata

33. Fruit divided into three ovate-winged samaras............................................................................ Serjania salzmaniana

34. Leaves opposite or verticillate .................................................................................................................................... 35

34. Leaves alternate .......................................................................................................................................................... 48

35. Stipules borne between the petioles........................................................................................................ Sabicea grisea

35. Stipules not borne between the petioles...................................................................................................................... 36

36. Plants hairy ................................................................................................................................................................. 37

36. Plants glabrous............................................................................................................................................................ 42

37. Latex present............................................................................................................................................................... 38

37. Latex absent ................................................................................................................................................................ 40

38. Glands absent from the leaf limb ................................................................................................. Temnadenia odorifera

38. Glands present on the leaf limb .................................................................................................................................. 39

39. Leaves scabrous, glands 2-6, distributed along the midvein ............................................................ Mandevilla scabra

39. Leaves not scabrous, glands 2-6 grouped at the base of the midvein ................................................ Matelea maritima

40. Leaf covered with long fine hairs; fruit divided into 3 samaras ........................................... Stigymaphyllon blanchetii

40. Leaf covered with short thick hairs............................................................................................................................. 50

41. Fruit a 3-valved capsule; seeds winged .................................................................................... Prionostemma asperum

41. Fruits grouped into a capitate compound infrutescence ...................................................................... Mikania obovata

42. Fruit winged ................................................................................................................................................................ 43

42. Fruit not winged.......................................................................................................................................................... 45

43. Wings less than three times the length of the fruit.................................................................... Niendenzuela acutifolia

43. Wings as long as or longer than three times the length of the fruit ............................................................................ 44

44. Wings oblong-falcate (Fig. 2J) ................................................................................................ Heteropterys nordestina

44. Wings asymmetrically obovate (Fig. 2I) ................................................................................. Stigmaphyllon salzmanii

45. Stigma feather-like................................................................................................................................. Clematis dioica

45. Stigma not feather-like................................................................................................................................................ 46

46. Fruit a 3-carpelar capsule; seeds winged .................................................................................... Hippocratea vollubilis

46. Fruit a berry or a 2-carpelar capsule; seeds not winged ............................................................................................. 47

47. Fruit a capsule, 2-carpelar................................................................................................................... Thunbergia alata

47. Fruit a berry ............................................................................................................................ Mendoncia blanchetiana

48. Ochrea present (Fig. 2N) ............................................................................................................. Coccoloba ochreolata

48. Ohcrea absent.............................................................................................................................................................. 49

49. Leaves compound ....................................................................................................................................................... 50

49. Leaves simple or compound unifoliolate.................................................................................................................... 69

50. Palms. Terminal leaflets reduced, stiff and hook-like (Fig. 4A)................................................................................. 51 50. Not palms. Terminal leaflets not hook-like, foliaceus ................................................................................................ 52

51. Raphe unarmed, leaflets unarmed or sometimes armed with thorns at the basis of the leaflets; apex of bracts covered in narrow thorns (Fig. 4B) ..................................................................................................... Desmoncus orthacanthos

51. Raphe and abaxial surface of the leaflets armed with stiff and twisted thorns; apex of bracts covered in pustular thorns (Fig. 4C) ...................................................................................................................... Desmoncus polyacanthos

52. Inflorescences opposite to the leaves .......................................................................................................... Cissus erosa

52. Inflorescences never opposite to the leaves................................................................................................................ 53

53. Leaves digitate ............................................................................................................................. Merremia macrocalyx

53. Leaves with 3 or more leaflets, never digitate ............................................................................................................ 54

54. Prickles or thorns present ................................................................................................................ Senegalia tenuifolia

54. Prickles or thorns absent ............................................................................................................................................. 55

55. Calyx persistent, 2-lobed ............................................................................................................................................ 56

55. Calyx persistent, 4 or 5 lobed ..................................................................................................................................... 58

56. Abaxial surface of the leave glabrous ........................................................................................... Canavalia parviflora

56. Abaxial surface of the leave glabrescent to pubescent ............................................................................................... 57

57. Calyx lobes of unequal lengths ................................................................................................... Canavalia braziliensis

57. Calyx lobes of equal lengths ............................................................................................................ Canavalia dictyota

58. Calyx 5 lobed .............................................................................................................................................................. 58

58. Calyx 4-lobed.............................................................................................................................................................. 66

59. Calyx including a spur ................................................................................................................................................ 60

59. Calyx lacking a spur ................................................................................................................................................... 61

60. Stem cylindrical; calyx lobes unequal in size, the larger lobe twice the length of the calyx tube.................................. ................................................................................................................................................... Centrosema pubescens

60. Stem angulate; calyx lobes of equal size ...................................................................................... Centrosema plumieri

61. Bracteole caducous; calyx campanulate ..................................................................................................................... 62

61. Bracteole persistent; calyx tubular.............................................................................................................................. 63

62. Stems bearing lenticels, glabrous; leaflets lanceolate; calyx persistent in fruit ............................. Connarus blanchetii

62. Stem lacking lenticels, hairy; leaflets ovate; calyx not persistent ............................................. Rynchosia pyramidalis

63. Seeds more than ten per fruit ................................................................................................... Macroptilum prostratum

63. Seeds 4-6 per fruit....................................................................................................................................................... 64

64. Legume obtusely quadrangular; seeds spherical ................................................................................... Clitoria falcata

64. Legume oblong to linear ............................................................................................................................................. 65

65. Plants glabrous; bracteoles stiff, adpressed to the rachis ........................................................................... Vigna luteola

65. Plants hirsute; bracteoles membranaceous, free ..................................................................................... Vigna vexillata

66. Lobes of the calyx of unequal length (Fig. 4D); fruit with urticating hairs… .................................... Mucuna pruriens

66. Lobes of the calyx of equal length; fruit without urticating hairs .............................................................................. 67

67. Plants herbaceous ..................................................................................................................... Centrosema brazilianum

67. Plants woody............................................................................................................................................................... 68

68. Stem hairy; calyx glabrous .................................................................................................................... Dioclea virgata

68. Stem glabrous; calyx hairy .................................................................................................................. Dioclea violacea

69. Fruits grouped in a flower-like inflorescence ............................................................................................................. 70

69. Fruits borne in racemes, panicles, glomerules or solitary........................................................................................... 73

70. Fruits arranged into a capitate compound inflorescence ....................................................... Cyrtocymura scorpioides

70. Fruits protected by two persistent, bright involucral bracts, not arranged into a capitulum ...................................... 71

71. Involucral bracts entire (Fig 4E) ....................................................................................... Dalechampia convolvuloides

71. Involucral bracts 3-lobed ............................................................................................................................................ 72

72. Stipules linear-subulate; seed rough, dark-brown (Fig 4F) ................................................... Dalechampia brasiliensis

72. Stipules reniform; seeds smooth, light-brown (Fig 4F) .................................................. Dalechampia pernambucensis

73. Plant bearing urticating hairs; fruit a 3-lobed capsule; a single seed per locule......................................................... 74

73. Plant not bearing urticating hairs; fruit 3-lobed or not, where 3-lobed more than one seed per locule ..................... 75

74. Leaves 5-veined ..................................................................................................................................... Tragia volubilis

74. Leaves 3-veined ................................................................................................................................. Tragia lessertiana

75. Fruit solitary................................................................................................................................................................ 76

75. Fruit borne in racemes, panicles or glomerule............................................................................................................ 77

76. Pseudo-stipules reniform ......................................................................................................... Aristolochia brasiliensis

76. Pesudo-stipules absent .............................................................................................................. Aristolochia pappillaris

77. Fruit a pod................................................................................................................................................................... 78

77. Fruit not a pod............................................................................................................................................................. 79

78. Apical third of the leaflets connate; bracts linear-lanceolate ....................................................... Phanera trichosepala

78. Basal third of the leaflets connate; bracts reniform ......................................................................... Phanera outimouta

79. Fruits winged .............................................................................................................................................................. 80

79. Fruits not winged ........................................................................................................................................................ 81

80. Fruits 3-winged; seeds numerous, flattened ................................................................................. Dioscorea marginata

80. Fruits 1-winged; one seed per fruit, spherical ............................................................................ Securidaca diversifolia

81. Fruits fleshy ................................................................................................................................................................ 82

81. Fruits dry..................................................................................................................................................................... 84

82. Fruits born in a cime, opposite to the leaves ....................................................................................... Cissus verticilata

82. Fruits born in caulinar or terminal racemes ................................................................................................................ 83

83. Leaves peltate; bracts cordate .................................................................................................. Cissampelos glaberrima

83. Leaves no peltate; bracts filiform ........................................................................................ Cissampelos andromorpha

84. Stems not releasing latex when cut............................................................................................................................. 85

84. Stems releasing white or colourless latex when cut ................................................................................................... 86

85. Sepals persistent, unequal, internal 2 bigger than the external 3, surrounding the fruit (Fig. 4H); seeds with a white aril ............................................................................................................................................................ Davila aspera

85. Sepals persistent, equal in size, not surrounding the fruit; seeds without a white aril ............................................... 87

86. Fruits solitary ................................................................................................................................. Operculina altissima

86. Fruits grouped in infrutescences ................................................................................................................................. 88

87. Fruits born in terminal racemes ...................................................................................................... Tetracera breyniana

87. Fruits born in axillary or cauliflorous glomerules ....................................................................... Doliocarpus dentatus

88. Leaves pinatifid ................................................................................................................................ Ipomoea quamoclit

88. Leaves entire to 5-lobed.............................................................................................................................................. 89

89. Leaves sagitate; inner surface of sepals with liguliform appendix ................................................... Ipomoea bahiensis

89. Leaves never sagitated; inner surface of sepals without appendix ............................................................................. 90

90. Leaves 3–5-lobed; sepals ............................................................................................................... Ipomoea hederifolia

90. Leaves entire; sepals not corniculate .......................................................................................................................... 91

91. Leaves and sepals glabrous......................................................................................................................................... 92

91. Leaves and sepals hairy .............................................................................................................................................. 93

92. Fruits irregulary dehiscent; seeds triangular ................................................................................... Merremia umbelata

92. Fruits regulary dehiscent; seeds with two surfaces flat, one convex .................................................. Ipomoea obscura

93. Fruits 4-valved, each valve dividing into two ................................................................ Jacquemontia menispermoides

93. Fruits 4-valved, valves not subdividing upon opening............................................................................................... 94

94. Leaves elliptic; hairs golden .......................................................................................................... Bonamia maripoides

94. Leaves cordate; hairs white .................................................................................................................... Ipomoea martii