Maxomys whiteheadi (Thomas, 1894) . Ann. Mag. Nat. Hist., ser. 6, 14:452.

TYPE LOCALITY: Malaysia, Sabah (N Borneo), Gunung Kinabalu .

DISTRIBUTION: Peninsular Thailand south of Isthmus of Kra, Malay Peninsula, Sumatra, Borneo, and various adjacent islands; absent from Java and Bali.

SYNONYMS: asper, batamanus, batus, coritzae, klossi, mandus, melanurus, melinogaster, perlutus, piratae, subitus.

COMMENTS: A Sundaic endemic. Ellerman and Morrison-Scott (1951) included whiteheadi in the Sulawesi musschenbroekii, but the two are separate species (Chasen, 1940; Medway, 1977; Musser, 1991; Musser et al., 1979; Tate, 1936). Spermatozoal morphology (Breed and Yong, 1986), and data from biochemical, morphological, and cytological studies (Chan et al., 1978, 1979; Yong, 1969) pointed to a close relationship between M. whiteheadi and the Malayan M. inas .