Key to the species based on male genitalia (males of S. selectoides sp. nov. and S. ethiopica sp. nov. are unknown)

1. Vincular process longer than sacculus (Figs 149, 150)........................................................ 2

- Vincular process equal to or shorter than sacculus........................................................... 3

2. Valva strongly curved at base; vincular process broader than sacculus at base; saccus extending slightly beyond the top of pedunculus; apical lobe of phallus short (Fig. 150).................................................... S. ergasima

- Valva not curved at base; vincular process as broad as sacculus at base; saccus extending well beyond the top of pedunculus; apical lobe of phallus large (Fig. 149)............................................................. S. aptatella

3. Vincular process and sacculus about 1/7 length of valva (Figs 131, 132, 151, 152)................................. 4

- Vincular process and sacculus longer than 1/7 length of sacculus............................................... 6

4. Valva distinctly bent; saccus very slender (Figs 151, 152)....................................... S. agassizi sp. nov.

- Valva weakly bent; saccus broad at base (Figs 131, 132)...................................................... 5

5. Uncus parallel-sided, posterior margin straight; vincular process strongly curved outwards (Fig. 132)........... S. nomias

- Uncus constricted in middle, posterior margin triangular; vincular process weakly curved outwards (Fig. 131).............................................................................................. S. nigristriana sp. nov.

6. Posterior margin of uncus emarginated to 1/4–1/3 length (Figs 147, 148).................................. S. swakopi

- Posterior margin of uncus straight or only slightly emarginated................................................. 7

7. Valva sigmoidal (Figs 121, 122, 146)..................................................................... 8

- Valva weakly curved.................................................................................. 9

8. Valva exceeding posterior margin of uncus; sacculus nearly 1/2 length of valva (Fig. 146)............... S. portosanctana

- Valva not exceeding posterior margin of uncus; sacculus 1/3 length of valva (Figs 121, 122)........... S. staudei sp. nov.

9. Vincular process very short, hump-shaped (Fig. 139)............................................ S. brandbergensis

- Vincular process long, from digitate to triangular........................................................... 10

10. Vincular process broader than sacculus at base (Figs 91–98).................................................. 11

- Vincular process narrower than sacculus at base........................................................... 12

11. Saccus short, not extending beyond the top of pedunculus, parallel-sided (Fig. 98).................. S. turiensis sp. nov.

- Saccus longer, extending beyond the top of pedunculus, tapered (Figs 91–97).................................... 12

12. Valva broader than vincular process at base (Figs 96, 97)....................................... S. wieseri sp. nov.

- Valva narrower than vincular process at base (Figs 91–95)................................................... 13

13. Phallus as long as valva; saccus apically inflated (Fig. 95).................................. S. ochroxantha sp. nov.

- Phallus longer than valva; saccus apically pointed or truncate (Figs 91–94)................................. S. incola

14. Vincular process distinctly narrowed apically, apex pointed (Figs 106–110)...................................... 15

- Vincular process weakly narrowed apically, apex rounded.................................................... 16

15. Saccus tapered apically (Figs 106–108).......................................................... S. subroseata

- Saccus truncate apically (Figs 109, 110)........................................................... S. biljurshi

16. Valva distinctly curved at 1/3 (Figs 113, 114)................................................ S. wolframi sp. nov.

- Valva gradually curved throughout...................................................................... 17

17. Culcitula covering uncus ventrally (Figs 133, 134)............................................. S. munita sp. nov.

- Culcitula not covering uncus ventrally................................................................... 18

18. Valva extending well beyond top of uncus (Figs 115, 116).................................................... 19

- Valva not or only slightly extending beyond top of uncus.................................................... 20

19. Vincular process 1/2 width of adjacent portion of valva (Fig. 116)............................. S. varivansoni sp. nov.

- Vincular process as broad as adjacent portion of valva (Fig. 115)............................... S. natalensis sp. nov.

20. Base of uncus 1/2 width of base of tegumen (Figs 140–142).................................................. 21

- Base of uncus 1/3 or less width of base of tegumen......................................................... 22

21. Saccus weakly constricted before apex, extending slightly beyond top of pedunculus (Figs 141, 142)......... S. traganella

- Saccus parallel-sided, not extending beyond top of pedunculus (Fig. 140)........................ S. etoshensis sp. nov.

22. Saccus parallel-sided or weakly constricted in middle, broad (Figs 99–105, 145).................................. 23

- Saccus broad at base, narrowing apically................................................................. 24

23. Outer margin of sacculus distinctly bent; vincular process 2/3–3/4 length of sacculus (Fig. 145)............... S. geomicta

- Outer margin of sacculus straight; vincular process as long as sacculus (Figs 99–105)....................... S. concreta

24. Vincular process digitate, tapered, extending to top of sacculus (Figs 111, 112)............................. S. diversa

- Vincular process a nearly equilateral triangle or an apically narrowed, outwardly curved process, not reaching top of sacculus .................................................................................................. 25

25. Posterolateral emargination of vinculum nearly an equilateral triangle (Figs 118–120)................. S. typica sp. nov.

- Posterolateral emargination of vinculum V-shaped or rounded (Figs 123–126).................................... 26

26. Posterolateral emargination of vinculum rounded; saccus extending well beyond top of pedunculus (Figs 123–125)................................................................................................. S. obsoletella

- Posterolateral emargination of vinculum V-shaped (Fig. 126)................................ S. afromontana sp. nov.

27. Vincular process a nearly equilateral triangle, extending 2/3 length of sacculus (Fig. 128)........... S. admirabilis sp. nov.

- Vincular process digitate, curved outwardly (Figs 117, 127).................................................. 28

28. Vincular process parallel-sided, extending to about apex, with outwardly curved tip (Fig. 117)................ S. phelotris

- Vincular process gradually curved outwardly (Fig. 127)......................................... S. erexita sp. nov.

29. Sacculus extending nearly 1/2 length of valva (Figs 143, 144)................................... S. meridioafricana

- Sacculus extending 1/3 or less length of valva (Figs 129, 130, 135–138)........................................ 30

30. Sacculus extending 1/3 length of valva; uncus narrowed distally; valva extending to top of uncus (Figs 137, 138)............................................................................................ S. asantesana sp. nov.

- Sacculus extending to 1/5–1/4 length of valva; valva not extending to top of uncus (Figs 129, 130, 135, 136)........... 31

31. Sacculus strongly curved inwards; valva weakly broadened posteriorly, not extending to top of uncus (Figs 135, 136).......................................................................................... S. ochracea sp. nov.

- Sacculus weakly curved inwards; valva distinctly broadened apically, extending to top of uncus (Figs 129, 130)................................................................................................ S. griseata sp. nov.

Key to the species based on female genitalia

(females of S. natalensis sp. nov., S. varivansoni sp. nov., S. phelotris, S. staudei sp. nov., S. afromontana sp. nov., S. admirabilis sp. nov., S. griseata sp. nov., S. nigristriana sp. nov. and S. ochracea sp. nov. are unknown)

1. Subgenital plates wrinkled medially, foam sculpturing indistinct (Figs 153–157)............................. S. incola

- Subgenital plate without wrinkles, unmodified or with foam sculpturing......................................... 2

2. Anterior margin of sternum VIII distinctly projected anteriorly (Figs 193, 194, 196)................................ 3

- Anterior margin of sternum VIII straight or weakly concave................................................... 5

3. Apophyses anteriores thick, slightly longer than anteromedial projection of sternum VIII (Fig. 194)........... S. aptatella

- Apophyses anteriores slender, distinctly longer than anteromedial projection of sternum VIII (Figs 193, 196)............ 4

4. Subgenital plates with pointed posterolateral corners; distal sclerite of signum short, straight; basal plate slender (Fig. 196)....................................................................................... S. agassizi sp. nov.

- Subgenital plates with rounded posterolateral corners; distal sclerite of signum long, curved at base; basal plate broad (Fig. 193)................................................................................ S. ethiopica sp. nov.

5. Foam-sculptured patches of subgenital plates displaced posteriorly (Fig. 195)............................. S. ergasima

- Foam-sculptured patches of subgenital plates, if present, medially or anteriorly.................................... 6

6. Papillae anales with microtrichia and inner sclerites (Fig. 181).................................... S. munita sp. nov.

- Papillae anales unmodified............................................................................. 7

7. Subostial sclerites triangular, distinct; subgenital plate folded anteromedially (Figs 182, 183)........ S. asantesana sp. nov.

- Subostial sclerites indistinct or absent; subgenital plate without anteromedial folds................................. 8

8. Apophyses anteriores distinctly inflated at base (Fig. 184)....................................... S. brandbergensis

- Apophyses anteriores not or only weakly inflated at base..................................................... 9

9. Apophyses anteriores 1/3 length of segment VIII (Fig. 188)..................................... S. meridioafricana

- Apophyses anteriores more than 1/3 length of segment VIII.................................................. 10

10. Ventromedial depression teardrop-shaped anteriorly (Figs 173, 174, 189–191).................................... 11

- Ventromedial depression not teardrop-shaped anteriorly..................................................... 13

11. Ventromedial depression exceeding anterior margin of segment VIII; distal sclerite of signum inflated at middle (Figs 173, 174)................................................................................ S. wolframi sp. nov.

- Ventromedial depression not reaching anterior margin of segment VIII; distal sclerite of signum not inflated at middle (Figs 189–191).......................................................................................... 12

12. Ventromedial depression covered with foam sculpturing; signum comparatively thick (Fig. 191).......... S. portosanctana

- Ventromedial depression unmodified; signum comparatively slender (Figs 189, 190)....................... S. geomicta

13. Ventromedial depression digitate, exceeding anterior margin of segment VIII (Figs 185–187, 192).................... 14

- Ventromedial depression rounded, not exceeding anterior margin of segment VIII................................. 17

14. Distal sclerite of signum 1/2 width of corpus bursae (Fig. 187)........................................ S. traganella

- Distal sclerite of signum less than 1/2 width of corpus bursae (Figs 185, 186, 192)................................ 15

15. Ventromedial depression covered with distinct foam sculpturing (Figs 185, 192).................................. 16

- Ventromedial depression wrinkled, foam sculpturing indistinct (Fig. 186)........................ S. etoshensis sp. nov.

16. Cells of foam sculpturing small; subgenital plates nearly connected posteromedially; base of signum stout (Fig. 185)......................................................................................... S. selectoides sp. nov.

- Cells of foam sculpturing large; subgenital plates broadly separated posteromedially; base of signum slender (Fig. 192)................................................................................................. S. swakopi

17. Apophyses anteriores twice length of segment VIII (Fig. 158)............................... S. ochroxantha sp. nov.

- Apophyses anteriores less than twice length of segment VIII.................................................. 18

18. Segment VIII twice as broad as long (Figs 160–172)........................................................ 19

- Segment VIII less than twice as broader as long (Figs 159, 175–180)........................................... 23

19. Papillae anales pointed posteriorly (Fig. 160)................................................ S. turiensis sp. nov.

- Papillae anales not pointed posteriorly (Figs 161–172)...................................................... 20

20. Ventromedial depression elongated, length 3x width in the broadest part (Figs 161–167).................... S. concreta

- Ventromedial depression subovate or rounded, length less than 3x width in the broadest part (Figs 168–172)........... 21

21. Distal sclerite of signum with hump on posterior margin (Figs 170–172)........................................ 22

- Distal sclerite of signum without hump on posterior margin (Figs 168, 169)............................. S. subroseata

22. Ventromedial depression rounded (Fig. 170)....................................................... S. biljurshi

- Ventromedial depression more elongated, subovate (Figs 171, 172)...................................... S. diversa

23. Subgenital plates 1/4 width of sternum VIII (Fig. 180)................................................ S. nomias

- Subgenital plates 1/3–1/2 width of sternum VIII (Figs 159, 175–179)........................................... 24

24. Signum weakly curved (Figs 175–177, 179)............................................................... 25

- Signum distinctly curved (Figs 159, 178)................................................................. 26

25. Signum slender, long; ventromedial depression narrow (Fig. 179).................................. S. erexita sp. nov.

- Signum thicker; ventromedial depression broad (Figs 175–177)................................... S. typica sp. nov.

26. Apophyses anteriores weakly broadened at base; margin of signum smooth (Fig. 159)................ S. wieseri sp. nov.

- Apophyses anteriores not broadened at base; margin of signum serrate (Fig. 178)........................ S. obsoletella