Myotis adversus (Horsfield 1824)

Large-footed Myotis (Kelawar Kaki Panjang)

Specimens examined (3).—Bako NP: TTU 108277 (GenBank EU521628), TTU 108278, TTU 108307.

Type locality.— Indonesia, Java .

Malaysian distribution.—Records from Peninsular Malaysia and other parts of the Asian mainland remain uncertain (Francis 2008). In Sabah, this species is known only from Sandakan Bay (Payne et al. 1985). A new geographic record was added for Bako NP and for the state of Sarawak.

Remarks.—This species was captured in a harp trap and a mist net across a small stream between swamp forest and the beach. As the common name implies, M. adversus has a large foot with a wing membrane inserted at the ankle (Hill 1983; Kitchener et al. 1995; Fig. 3d). Myotis adversus also has a unique translucent wing membrane. A high genetic distance of 15.2% was documented when the Sarawak specimens were compared with a GenBank cyt- b gene sequence (AB106587; see Kawai et al. 2003) from Taiwan. Based on geographic distribution, our specimen would represent M. adversus carimatae and that from Taiwan should correspond to M. adversus taiwanensis (see Kitchener et al. 1995). However, the level of genetic variation suggests that these should be elevated to species status.