Nothrotheriinae plus Megatheriinae

PP = 54, age = 17.96 Mya (16.39–19.87).

This clade, which unites the clades Nothrotheriinae and Megatheriinae, was recovered only for H models and par_IW10 and par IW5, as stated above, and was poorly supported.

The clade was supported by four synapomorphies, recovered with both methods: presence of lingual and labial grooves in mf3; temporal lines approximate midline but do not meet to form a sagittal crest on skull roof; well-developed buccinator fossa of maxilla, with a deep depression; and Glaserian fissure opening into a distinct groove in squamosal lying medial to entoglenoid process.