Megatheriini

PP = 100, age = 9.85 Mya (9.07–10.67).

This clade was consistently recovered in all analyses performed in this study with an invariable arrangement:( Anisodontherium,( Pyramiodontherium, ( Megatheriops, ( Megatherium, ( Proeremotherium, Eremotherium))))).

Megatheriini was supported by eight synapomorphies, all but one, recovered with both methods: thickness of cementum much larger than orthodentine; mf3 smaller than next smallest molariform; reflexed basicranial/basifacial axis; profile of nasal region and braincase relatively horizontal, but nasal region depressed relative to braincase; maximum length of nasal bones greater than or equal to three times the width of both nasals (BI); presence of ventral extension in maxilla for dental alveoli; concave mediolateral contour of palate; and occipital condyles situated well dorsal to the dentition.