Nematheriinae

PP = 100, age = 18.60 Mya (16.85–20.28).

This clade contains the genera Nematherium and Analcitherium, and was recovered in all analyses performed in this study.

Nematheriinae was supported by seven synapomorphies, all recovered with both methods: anteroposteriorly ovate Mf4 cross-section; snout depressed anteriorly; external nares not greatly enlarged; absence of pterygoid inflation; presence of lacrimal eminence; presence of prominent lateral walls in lacrimal foramen; and jugal and lacrimal anteriorly overlapping facial portion of maxilla.