Urumacotheriinae

PP = 60, age = 17.82 Mya (14.03–21.51).

This clade was originally proposed with fewer taxa (Negri & Ferigolo, 2004), but was expanded here to include both Urumacotherium and Pseudoprepotherium . It did not receive high node support, despite being present in trees from all analyses.

Urumacotheriinae was supported by seven synapomorphies (five for both methods and two exclusively for BI): molariform morphology of Cf1/ cf1 (BI); posterior edge of mandibular condyle inclined posterodorsally; eversion of lateral edge of symphyseal spout (BI); premolariform portion of palate roughly equal or longer than the length of molariform toothrow; presence of a distinct neck at the base of occipital condyles; occipital condyles elongated anteroposteriorly in ventral view; and lesser tubercle of humerus proximally projected as much as greater tubercle.