Ahytheriini plus Choloepodini

PP = 69, age = 8.66 Mya (6.84–10.85).

This clade was recovered in most of the analyses performed in this study but was not well supported. The exceptions for the presence of the clade were two H models—IW10 and IW5, par_IW10 and par_IW5—in which the intertropical megalonychids were recovered more closely related to a clade uniting Megalonyx and Pliometanastes .

The clade was supported by 11 synapomorphies (seven for both methods and four exclusively for BI): mandibular symphysis extends posterior or to the level to mf1; lateral edge of symphyseal spout not everted; temporal lines approximate midline, but do not meet to form a sagittal crest (BI); presence of a contact between maxilla and lacrimal, within orbit; absence of a crest at median suture of palatine; interpterygoid region narrower than interpalatine region (BI); presence of pterygoid inflation; presence of a median ridge of basisphenoid (BI); jugular foramen well separated from hypoglossal foramen; presence of glenoid posterior shelf; and convex posterior margin of ulnar diaphysis in lateral view (BI).