4. Madecadryinus ranomafanensis Olmi, Copeland & van Noort, sp. nov.

(Figs 258F, H, 259)

Diagnosis. ♂ of Madecadryinus with POL about 3 × as long as OOL (Fig. 258F) and mesoscutum granulate.

Description. ♂ (Figs 259). Fully winged; body length 1.1–1.5 mm. Head brown, except mandible and clyp- eus testaceous; antenna brown-testaceous; mesosoma brown, occasionally black; metasoma brown-testaceous; legs yellow. Antenna filiform, setose, very slender, about as long as body; antennomeres in following proportions: 5:3:7:8:8:8:8:8:7:8. Head (Fig. 259C) convex, shiny, unsculpured; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; ocelli large (Figs 258F, 259C); POL = 3; OL = 1; OOL = 1; minimum distance from eyes to occipital carina as long as OL; TL = 4; greatest breadth of lateral ocelli longer than OL (2:1); lateral ocelli touching occipital carina; temple very long, shorter than eye (4:8); clypeus without short median longitudinal keel. Palpal formula 6/3. Mesoscutum (Fig. 259A) dull, completely slightly granulate. Notauli complete, posteriorly separated; minimum distance between notauli less than twice as long as breadth of lateral ocelli (3:2). Mesoscutellum and metanotum shiny, unsculptured. Epicnemium exposed. Mesopleuron and metapleuron shiny, unsculptured. Metapectal-propodeal disc and propodeal declivity reticulate rugose, with some areolae as large as tegulae. Forewing (Fig. 259D) hyaline without slight dark transverse bands; distal part of 2r-rs&Rs vein shorter than proximal part (8:12); 2R1 cell open. Hind wing hyaline, without dark transverse band. Paramere (Fig. 258H) without inner apical and pointed expansion; basivolsella with apex parallel to distivolsella (Fig. 258H). Tibial spurs 1/1/2.

♀. Unknown.

Material examined. Types: ♂ holotype (CASTYPE19469): MADAGASCAR: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21°15.99’S 47°25.21’E, 1020 m, 19–26.II.2002, MT, secondary tropical forest, R. Harin’Hala leg., MA-02-09C-17 (CAS) . Paratypes: MADAGASCAR: same locality label as holotype, 4–16.V.2003, MA-02-09C-60, 1♂ (CAS); same locality label as holotype, 12–23.IV.2003, MA-02-09C-58, 1♂ (MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21°13.57’S 47°22.19’E, 1110 m, 26.II–4.IV.2002, MT, high altitude rainforest, R. Harin’Hala leg., MA-02-09A-18 , 1♂ (MOLC); same locality label, 26–31.III.2002, MA-02-09A-22, 3♂♂ (2 in CAS, 1 in MOLC); Fianarantsoa, Manombo Special Reserve camp site, 32 km SSE of Farafangana, 23°01.31’S 47°43.20’E, 36 m, 27.VII–10.VIII.2005, MT, lowland rainforest, M. Irwin, R. Harin’Hala leg., MA-28-24 , 2♂♂ (CAS, MOLC); same locality label, 24.IV–10.V.2005, MA-28-18, 1♂ (CAS); same locality label, 6–- 24.VI.2005, MA-28-21, 1♂ (CAS); same locality label, 30.X–12.XI.2004, MA-28-03, 1♂ (MOLC) .

Hosts. Unknown.

Distribution. Madagascar.

Etymology. The species is named ranomafanensis after the collection site in the Ranomafana National Park.