Metalycaeus heudei (Bavay & Dautzenberg, 1900)

Fig. 41

Alycaeus (Charax) heudei Bavay & Dautzenberg, 1900a: 121–122 .

Alycaeus (Charax) heudei — Bavay & Dautzenberg 1900b: 458–459, plate 11, figs 15–18.

Alycaeus paviei Bavay & Dautzenberg, 1912: 50–51, pl. 4, figs 5–8.

Alycaeus paviei var. minor Bavay & Dautzenberg, 1912: 51, pl. 4, fig. 9.

Chamalycaeus (Dicharax) compressicosta Zilch, 1957: 145–146, fig. 33.

Chamalycaeus (Dicharax) fractus Varga, 1974: 165–167, figs 1–5.

Alycaeus zhuangiyucuii Yang, Fan, Qiao & He, 2012: 32, fig. 2.

Metalycaeus heudei — Páll-Gergely et al. 2017: 74–84, figs 49C–D, 50–52, 53C–D. ( paviei Bavay & Dautzenberg, 1912, paviei var. minor Bavay & Dautzenberg, 1912, compressicosta Zilch, 1957, fractus Varga, 1974 and zhuangiyucuii Yang, Fan, Qiao & He, 2012 are synonyms; Inkhavilay et al. 2019: 16, fig. 6C.

Alycaeus kengtungensis Godwin-Austen, 1914: 409, plate 139, figs 6, 6a. new synonym

Material examined. Holotype: Myanmar, Shan State, “Kengtung, Siam Frontier, leg. Woodthorpe ”, NHMUK 1903.7.1.3037 (holotype of A. kengtungensis). For the examined type specimens of other taxa see Páll-Gergely et al. (2017) . Other material: 4 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Bo Pha Gu Cave sink, 19°21.121’N, 97°3.759’E (locality code: JG10), 10 Feb. 2019, J. Grego leg., JG ; 2 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village, under bridge over Phruno River, 19°22.966’N, 97°02.153’E (locality code: JG100), 12 Dec. 2019, J. Grego leg., JG ; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village, rocks above bridge over Phruno River, 19°23.011’N, 97°02.108’E (locality code: JG101), 12 Dec. 2019, J. Grego leg., JG ; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Entrance of Phruno River Cave, 19°22.744’N, 97°02.570’E (locality code: JG102), 12 Dec. 2019, J. Grego leg., HNHM 104867 (photographed shell: 4.77 mm, H: 3.66 mm) ; 17 shells, same data as preceding, JG .

Diagnosis of specimens from Myanmar. A medium-sized to large species with very long R2 having typical Metalycaeus - type elevated, thin, widely spaced ribs, a narrow swelling on R3, and a rounded aperture.

Measurements. D: 4.65–5.05 mm, H: 3.66–3.71 mm (newly collected specimens).

Remarks. Metalycaeus heudei is a widely distributed, variable species reported from China, Vietnam and Laos. Alycaeus kengtungensis (Fig. 41B) from the eastern border of Shan State, Myanmar, and the newly collected shells from Kayah State (Fig. 5) fall within the known morphological variation of Metalycaeus heudei . Therefore, we consider A. kengtungensis a new synonym of M. heudei . With this, M. heudei is the most widely distributed alycaeid species known.