Cybaeus pan Bennett spec. nov.

Figs 33–39, 58, 86

Type material. U.S.A.: California: Holotype male, Sierra County, 2 mi. N of Calpine, 6.ix.1959, W.J. Gertsch & V. D. Roth (AMNH) . Paratypes: Butte. 1♀, Lumpkin Rd. near Feather Falls, Oroville, 18.iv.1994, J. Boutin (CAS) ; Plumas . 1♀, NE of Bucks Lake, 4.ix.1988, D. Ubick (CAS) ; Nevada. 1♀, 1 mi. N of Washington, 3050’, 27.xii.1986, D. Ubick (CAS) ; Sierra . 9♂ 10♀, 2 mi N of Calpine, 6.ix.1959, V. D. Roth & W.J Gertsch (AMNH) ; 1♀, 5 mi. E of Camptonville, 14.vii.2002, D. Ubick (CAS) ; 1♂ 1♀, Sierra City, The Cups, 6.ix.1959, V. D. Roth & W.J Gertsch (AMNH) ; 1♀, 1.5 mi. E of Sierra City, Wild Plum Campground, 4200’, 12–16.viii.1996, D. Ubick (CAS) ; 1♀, 1.5 mi. E of Sierra City, Wild Plum Campground, 4200’, 9.viii.2001, D. Ubick (CAS) ; 1♂, 1.5 mi. E of Sierra City, Wild Plum Campground, 4200’, 18.vii.2002, D. Ubick (CAS) ; 2♂, 3.5 mi. NE of Sierra City, 5100’, 17.viii.1993 ’, D. Ubick (CAS) ; 2♀, 6 mi. NE of Sierra City, SFSU Field Station, 15–18.viii.1991, D. Ubick (CAS) ; 1♂ 5♀, 6 mi. NE of Sierra City, SFSU Field Station, 5500’, 10–18.viii.1992, D. Ubick (CAS) ; 1♂ 1♀, 6 mi. NE of Sierra City, SFSU Field Station, 5500’, 16–20.viii.1993, D. Ubick (CAS) ; 1♂ 2♀, 6 mi. NE of Sierra City, SFSU Field Station, 5500’, 7–11.viii.2000, D. Ubick (CAS) ; 1♀, Yuba Pass, 8.viii.2001, D. Ubick (CAS) ; 1♀, Yuba Pass, 13.viii.1997, G. deNevers & H. Wilcox (CAS) ; 3♂ 3♀, Yuba Pass, 9.viii.2000, D. Ubick (CAS) ; 1♀, Yuba Pass, 8.viii.2001, R. Love (CAS) ; 2♀, Yuba Pass, 18.vi.2003, D. Ubick (CAS) .

Etymology. The specific epithet is a noun in apposition from Pan, the ancient pipe-playing Greek god of the countryside. Cybaeus pan spec. nov. is named in honor of the late John “Pan” Logan, dear friend and native of the northern Sierra Nevada region of California, who did much in his lifetime to uphold the traditions associated with mythical Pan; name in apposition.

Diagnosis. Among the males of the consocius group species, C. pan spec. nov. is only likely to be confused with C. vulpinus . The males of the two species are distinguished by, in C. pan spec. nov., a relatively simple, smoothly curved and untwisted proximal arm of the tegular apophysis in ventral view (Figs 35–36) with, in prolateral view (Fig. 58), a small triangular dorsal keel (versus smoothly curved but twisted proximal arm lacking a dorsal process [Figs 57, 82] in C. vulpinus). As well the patellar apophysis (Figs 33–34) is longer (about 3/4 the width of the patella), anterolaterally directed, and has more peg setae (about 35) in C. pan spec. nov. (versus about 1/3 the width of the patella, anteriorly directed, and bears only 13 peg setae in the single known male of C. vulpinus [Fig. 81]). Among the females of the consocius group, the female of C. pan is most likely to be confused with the other species which lack U-shaped copulatory ducts: C. ubicki spec. nov., C. penedentatus, C. vulpinus, C. simplex, and C. opulentus spec. nov. Among these species, the female of C. pan spec. nov. is most difficult to separate from C. vulpinus; the epigyna and vulvae of these two species are very similar and, in some cases, distribution may be the best or only way to separate them (Fig. 86). The only apparent morphological character separating the two species is the conformation of the spermathecal stalks: in C. pan spec. nov. they are relatively long, distinctly sinuous and describing an acute angle (approximately 55 ⁰ –65 ⁰) near the Bennett’s glands (Figs 38–39) versus in C. vulpinus (Figs 40–41) only slightly sinuous and obtusely angled (circa 120 ⁰ –130 ⁰). Separating the females of C. pan spec. nov. and C. opulentus spec. nov. is discussed under the diagnosis of the latter. From C. ubicki spec. nov. and C. penedentatus, the female of C. pan spec. nov. is distinguished by a combination of a large, conspicuous atrium occupying about 1/2 the width of the vulva (Figs 37–38) (versus atrium small, inconspicuous, occupying about 1/3 the width of the vulva in C. penedentatus [Figs 46–47, 49, 52]) and copulatory ducts relatively small, narrow, and inconspicuous (Figs 38–39) (versus large, broad, and prominent in C. ubicki spec. nov. [Figs 75–76, 78–79]). Finally, specimens of both sexes of C pan spec. nov. (as well as of C. vulpinus) usually have banded femora; specimens of all the other consocius group species usually have unbanded femora.

Description. Femora usually banded. Ventral tibia I macrosetae usually 2-1p(or 0)-2-1p-1p(or 0).

Male: (n=20). As per Diagnosis. Measurements (n=9). CL 2.33–2.8 (2.6±0.2), CW 1.70–2.05 (1.90±0.13), SL 1.20–1.38 (1.30±0.06), SW 1.16–1.33 (1.25±0.05). Holotype CL 2.43, CW 1.80, SL 1.22, SW 1.20.

Female: (n=35). Epigynum usually with small notch posteriorly (Fig. 38) (inconspicuous and may be difficult to discern, most easily observed in cleared specimens), absent in some specimens (Fig. 37); atrium (Figs 37–38) medially located on epigynum, usually strongly arched (some specimens have a weakly arched atrium approximating the characteristic atrial morphology seen in specimens of C. vulpinus [e.g., Fig. 40]). Copulatory ducts (Figs 38–39) separated except at atrium; anteriorly directed and diverging from atrium, anterior margin often indistinct. Stalks of spermathecae simple (Figs 38–39). Measurements (n=13). CL 2.20–3.0 (2.5±0.2), CW 1.50–2.05 (1.74±0.16), SL 1.13–1.40 (1.25±0.08), SW 1.05–1.33 (1.19±0.07).

Distribution and natural history. Northern Sierra Nevada region of eastern California northwest of Lake Tahoe in Butte, Plumas, Nevada, and Sierra Counties (Fig. 86). Mature males have been collected from July through September.